This research considers human impacts on three components of biodiversity (composition, spatial structure and function). Given the relict character and unusual biogeochemical balance of tropical montane cloud forests in Mexico, logging poses a pressing threat to their survival. Specifically, this thesis explores the effect of selective logging and above-ground secondary succession on the biogeochemical cycling and soil macroinvertebrate community in a cloud forest in Oaxaca, Mexico. The research investigates: (1) whether the above-ground chain of successional changes in tree dominance, litterfall, litter diversity and soil microenvironmental conditions are coupled with a below-ground succession of soil nutrient availability and macroinvertebrate communities, (2) the role of spatial structuring of environmental conditions and litter resources as determinates of the nutrient availability and macroinvertebrate taxa abundance, (3) the implications of successional changes for decomposition and (4) whether the local influence of single trees explains the spatial structure of macroinvertebrate communities in late successional forests. The work was carried out in three chronosequences (c.15, 45, 75 and 100 year-old stages) of high altitude (1500-2000m) tropical montane cloud forest, two recently logged sites and two pristine sites. the macroinvertebrates in the litter and mineral soil were hand sorted from monoliths. Parametric statistics and canonical correspondance analysis were used to determine mean successional trends, and Spatial Analysis by Distance Indices and geostatistical methods were used in combination to determine spatial patterns. Two decompositional experiments were performed to explore the relationship between decompositional rate, litter quality and macroinvertebrate community higher taxa composition in different successional stages and under the canopy of different tree species. The research showed that: 1. The macroinvertebrate community composition in both recently logged sites and pristine forests were distinct compared to secondary successional stages. A decrease in soil temperature and nutrient availablity but increase in litter diversity and soil organic matter recorded through succession were accompanied by an increase in the number of macroinvertebrate taxa in the soil. For exampl, Collembola were most abundant in recently logged sites and earthworms (Megascolecidae) were almost excluvely found in pristine forests. 2. The oldest secondary forest (100 year-old) showed the highest frequency of aggregation in the abundance of individual macroinvertebrate taxa, and the highest and most uniform value of Shannon's diversity. This suggests that high levels of diversity in litter resources and soil chemistry in late succession are associated with complex spatial structuring of highly diverse macroinvertebrate communities. 3. The leaves of a late successional species (Persea americana)decomposed at a slightly slower rate than an early successional species (Pinus chiapensis) in all successional stages, yet the number and Shannon's diversity of macroinvertebrate taxa that invaded decomposing P.Americana leaves was consistently higher. The preference of macroinvertebrate taxa for the late-successional leaves was ultimately explained by differences in leaf quality during decomposition. P.americana leaves had higher concentrations of cations throughout decomposition and their concentration of lignin and nitrogen became higher. 4. In the 100-year-old forest, the effect of seasonal variation on soil microenvironmental conditions and litter availability was different under the canopy of different tree species. Furthermore, the chemical evolution of the same leaf type (e.g. Oreopanax xalapensis) was differentwhen decomposing under different canopies. The highly diverse and spatially complex macroinvertebrate community found in late succession (and experimental litter) was largely explained by the interactive effects of seasonal variation, tree species, litter quality and availability of the decomposing leaf type. The results provide the first analysis of the relationship between soil biodiversity and the tight biogeochemical cycling in this relict ecosystem type. Overall the results indicate that mature cloud forests sustain a diverse and spatially heterogenous macroinvertebrate community. The compositional and spatial components of soil biodiversity are compromised by logging and full recovery may take mopre than 100 years.

The effects of sewage sludge application and the additional effects of clearfelling on nutrient cycling have been assessed in a mature Scots pine stand, north-east Scotland. The forest site was situated in a low rainfall area with high evapotranspiration and low leaching losses. The ground vegetation formed an important sink for N and P following sludge addition and clearfelling. Sludge application caused a foliar N response, increased foliar needle weight, and litterfall showed a significant and rapid potential to immobilize NH₄-N and PO₄-P added in sludge. The soil availability of NH₄-N and PO₄-P together with rates of mineralization of N and P and nitrification in sludge-treated areas were significantly increased compared with that of the control. Concentrations of NH₄-N, NO₃-N, PO₄-P and organic P were increased significantly following sludge addition and fluxes were an order of magnitude greater than that of the control. Peaks in concentrations of NH₃-N in sludge-treated areas exceeded the EC limit of 11.3 mg 1^-1 on several occasions. Of the total N and P applied to the sludge, throughfall and litterfall over a 17 month period, 2.4 % and 0.72 % were measured in the B_s soil horizon flux. After clearfelling, initial increases in soil availability of NH₄-N, NO₃-N andPO₄-P were measured. Felling increased nitrification rates, although increases were significant only in areas previously treated with sludge. Fluxes of NO₃-N, NH₄-N and PO₄-P and organic P were increased at least an order of magnitude greater in felled areas than those of the control. Immobilization of N and P was greater in green and abscised litters placed in previously sludge-treated areas than compared with those placed in control areas.

Inherent in the structure of timber are features affecting dimensional stability, stiffness and strength. These include knots, compression wood and spiral grain. Physical characteristics such as log shape, density, rate of growth, presence of juvenile wood and microfibril angle also affect these properties and, in turn, utilisation. In this thesis, the relationships between tree, log and board variables to the properties of stiffness and distortion of British-grown Sitka spruce (Picea sitchensis) are examined. The research‟s main aim was to identify variables which could potentially be used to sort timber. Different behaviour in terms of the relationship between stiffness and variables such as density, knot content, log taper, tree height and axial position were noted between groups of upper log and butt log material, and between the four stands and two sites studied. Relationships between stiffness and these variables were generally weak, notably for density. Overall, variables based on radial position within the stem were not found to be useful sorting parameters. No relationship between batten stiffness and compression wood content or slope of grain was observed. The relatively low stiffness of butt wood material was associated with high microfibril angle. Batten twist was found to be a function of spiral grain angle and distance from pith. However, practical difficulties with the use of slope of grain measurements to sort timber were noted. Although severe forms of compression wood were seen to be associated with stem-form correction, relatively straight logs and those which were round, or without pith eccentricity, were also observed to contain compression wood and hence yield timber which distorted on drying. In particular, this work demonstrates the effect of sorting timber using combined variables (e.g. log shape and knot content). These findings may be of use to foresters, sawmillers and developers of timber scanning technologies.

Sitka spruce (Picea sitchensis (Bong.) Carr.) has been the most common forestry species in UK plantations for many decades. It is generally fast grown in short rotation, which has implications for quality. The harvested timber has a number of end-use applications. Of these, sawn construction grade timber is at the high end of market value. Currently, under one third of home grown sawn timber processed by sawmills achieves the required C16 or higher grading for construction class timber. Presently there is significant standing stock of 40 years old or under. Improvements in timber quality would create significant benefits for growers and processors. Distortion in kiln dried sawn timber is a considerable barrier to improving grading classification and marketability. Twist in particular accounts for around six percent of the rejections after kiln drying. Additional twist when timber dries further; in storage, at construction sites or in-service, is perceived as a marketing issue. The focus of this work was to better understand the evolution of twist during the drying process and seek to improve quality by examining the key properties in sawn timber that contribute to the formation of twist. Samples of regularly processed full size (3 m in length) sawn timber were kiln dried with no restraint to promote maximum twist. This was done in three stages to assess the impact of intermittent drying. Measurements of twist, moisture content, acoustic velocity and basic dimensions were taken before and after each drying cycle. The results showed highly significant differences between battens containing the pith and those that did not. Drying in stages did not seem to affect the linearity of the moisture content against twist relationship. Twist was seen to develop at a consistent rate throughout the process regardless of the final amount of twist accrued or the position from which the batten was sawn. Where timber is likely to dry to lower moisture content after sale it is therefore possible to predict the additional twist from the moisture content expected and allow for it by stricter grading than the twist standard demands. Twist was seen to decrease slightly during a lengthy period of storage in controlled conditions between drying cycles. This may allude to a relaxation effect which could have implications for timber distortion in-service but this would require a more extensive study before any such conclusions could be drawn. A second batch of unrelated samples, were kiln dried to 12% MC under restraint. All of these samples contained the pith along their length and all developed significant amounts of twist. This suggests that battens containing the pith will twist regardless of restraint. A method was developed to measure longitudinal shrinkage. This required the batten size to be reduced to fit the method and battens were sawn into 3 x 1 m sections. Battens were re-saturated and measured daily throughout air drying until an equilibrium moisture content was achieved. Measurements of transverse shrinkage were also taken concurrently. No relationship could be found between the longitudinal and transverse results. When plotted against the data for twist a positive relationship was found with longitudinal shrinkage. Acoustic tools were used to calculate dynamic stiffness. As the values for acoustic velocity squared correlated well with dynamic stiffness, other possible relationships were explored. Good agreement was found with twist and longitudinal shrinkage. This suggests it may be possible to predict twist in sawn timber directly or as a function of microfibril angle, using acoustic tools. A simple method for measuring grain angle was devised. Discs taken from the trees used in the drying experiment were measured for grain angle to compare with other measured properties. The measurement method worked well but the sample size was small and the discs too thin to provide good quality data quality. The results were not as instructive as expected. A vague relationship with twist was found. A repeat of the experiment with an improved method could yield more conclusive results. A method was developed to measure radial and tangential shrinkage in discs. Discs of Sitka spruce from a different source were used. Discs were saturated and allowed to dry with the expectation of cracks developing. The intention was to explore the nature of internal stresses present in intact discs and why tangential and radial shrinkage differs. As only one disc cracked the scope of comparison was limited but it was apparent that the greater shrinkage tendency in the tangential direction was responsible for radial cracking.

Trees and forests are under increasing threat from pathogens which cause huge economic and ecological damage. The unprecedented global movement of pathogens into new areas creates novel pathosystems, while the changing climate affects the dynamics of endemic pathosystems. Co-evolution within endemic pathosystems affects the genetic composition of hosts and pathogens. Spatial heterogeneity in pathogen pressure leads to genetic variation in disease-related traits among host populations. In contrast, novel hosts or populations are expected to be highly susceptible to exotic pathogens as there has been no evolution of defence responses. Host response to disease can therefore be an indicator of a novel or endemic pathosystem. The long term resilience of forests to pathogens depends on the adaptive capacity of both the host and pathogen species. Establishing the extent of genetic and phenotypic variation within both the host and pathogen is therefore fundamental in understanding past, current and future pathosystem dynamics. The most significant current threat to Scots pine (Pinus sylvestris) is Dothistroma needle blight (DNB) caused by the foliar pathogen Dothistroma septosporum which is assumed to be exotic to Great Britain. This study aimed to increase understanding of the genetic and phenotypic variation in this pathosystem. Results from this study show that there are high levels of variation in the Dothistroma – Pinus pathosystem. Genetic variation, elucidated using neutral genetic markers, mating type specific markers and in vitro analysis of phenotypic variation in D. septosporum collected from Scottish pinewoods, was found to be high: there was high allelic diversity, particularly within plantation forests outside the native pinewood range, and high phenotypic plasticity in response to different temperature treatments. Both mating type idiomorphs were found in one forest which demonstrates their potential for sexual as well as asexual reproduction. There is also tentative evidence from this study that the pathogen is either introduced to Great Britain or that endemic pathogen populations have been augmented with introduced pathogens. Artificial and natural inoculations of native Scots pine provenances with D. septosporum indicate that there is considerable variation in susceptibility to DNB across the native range in Scotland and that variation in this trait is both highly heritable and evolvable. Furthermore, provenance mean susceptibility to DNB is negatively and significantly associated with water-related variables at site of origin, a finding that is potentially indicative of a co-evolutionary history between host and pathogen. Genetic differences among individuals which are ‘resistant’ or ‘susceptible’ to DNB were identified in Pinus radiata for which there has been extensive research in this pathosystem, by comparing the transcriptome sequences of the two phenotypic groups. Nearly half of the genetic differences identified among phenotypes were found in genes with a putative defence function. In conclusion, native Scots pine provenances contain the necessary heritable genetic diversity to evolve a decrease in their susceptibility to D. septosporum through natural selection in response to elevated prevalence of this pathogen. However, implementation of key native pinewood management strategies, including encouraging regeneration in particular, are necessary in order to facilitate the adaptive evolution of native forests to increased levels of DNB. The effectiveness of this response will depend on the rapidity of adaptation of the pathogen. Measures to limit adaptation where possible, including the use of pathogen monitoring and control in nurseries and the limitation of pathogen movement into native pinewoods, should be continued.

The anatomy of bifurcations in trees requires further scientific investigation as the current anatomical model for them is logically flawed. The provision of a better model will assist in scientific studies of woody plants, the risk assessment of junctions in mature trees and provide bio-inspiration for Y-shaped joints in composite materials. In this study, the xylem formed in the central axis of a hazel (Corylus avellana L.) bifurcation is shown to provide a disproportionately greater amount of its tensile strength. CT scanning identified that this centrally-placed xylem was 28.1% denser, with 63% less vessels formed in this tissue, such vessels being 50.5% of the diameter and 32.5% of the length of those formed in adjacent stem tissues. The wood grain pattern at the bifurcation apices were 22 times more tortuous, forming interlocking patterns that acted to resist tensile forces by requiring the extraction or breaking of wood fibres along their length (the axial tensile strength of wood). Subsequent tests confirmed that this conferred more than 100% additional tensile strength to these specialised xylem tissues. These findings provided the basis of a novel anatomical model for bifurcations in woody plants. Further to this, the effects of several factors upon junction strength and biomechanical behaviour were assessed in bifurcations of hazel, identifying the weakening effect of bark inclusions and three types of artificial modification as well as differences in wind-induced movement between bifurcation types. This study concludes that further investigations of bifurcations in a wider range of woody plants and observations of the developmental stages of the interlocking wood grain patterns found at bifurcations would usefully add to existing knowledge.

Succession following shifting cultivation was studied in secondary forests (5 to 40 years old) and a primary forest in Bragantina region, Para, Brazil. Secondary forests covered 62% of the study area (89 km² ) and crops and pasture 18%. Soil analyses showed an increase in pH, exchangeable calcium, magnesium and potassium and a decrease of organic matter and exchangeable aluminium, immediately after burning. Organic matter and extractable phosphorus increased with forest age, exchangeable aluminium and potassium were the same in both secondary and primary forests and exchangeable calcium and magnesium remained higher in the secondary forests. The highest densities of individuals and the least basal area and height were found in the younger secondary forests. The number of woody species (≥5 cm dbh) 250-m2 plot ranged from 8 to 17 in the secondary forests and 17 to 27 in the primary forest. Ordination and classification of floristic data suggested four successional communities. The forests regenerated readily and studies showed: the largest seed bank (0-5 cm) in the 5-years-old forest (1190 ± 284 seeds m-2) and a decrease with age to 137 ±19 seeds m-2 in the primary forest; the highest seed rain in the 5-years-old forest (883 ± 230 seeds m-2) and least in the primary forest (220 ± 80); 46% of the smaller plants (≥1 m tall, < 5 cm dbh) were seedlings and 54% were sprouts in the 5-years-old forest but in the 10-years and 20-years-old forests, seedlings (81%) predominated; adult plants (≥5 cm db h) in 5 to 20-years-old plots regenerated mainly from sprouts. Even after about 90 years of shifting cultivation the region has the potential for forest regeneration and the soil nutrients are able to recover to values similar to those in the primary forest. This raises the hope that, if a land use plan for Bragantina could be implemented, then it would be successful.

This policy brief aims to propose options to narrow the wood supply and demand gap through a critical scenario analysis at the national level, complemented with wood value chain analysis that reflects the on-the-ground realities. Limited access to credit services, complicated bureaucratic processes, insecure land tenure, and the dominant influence of government forestry enterprises in the wood value chain are among the key constraints to augmenting the national wood supply. The government needs to remove some of the complicated bureaucratic land acquisition processes to ensure tenure security for private investors, small- and medium-sized wood producers. Wood import substitution with locally produced wood products can reduce government spending, while promoting local industrialization, innovation, and job creation. Cross-sectoral collaboration among government departments and the need to extend the participation of small- and medium-scale wood producers beyond wood production to wood processing and marketing activities is crucial to upscale the wood supply. This policy brief is intended for forest and natural-resource policymakers, wood industrial sector, non-governmental organizations, and academia dealing with the forestry and wood sector in Ethiopia.:Key Messages: Page 1 Introduction: Page 2 Project methods: Page 2 National wood product supply scenarios: Page 3 Wood value chain: Page 4 Policy implications and recommendations: Page 5 Contact: Page 6

Based on recent estimates, the average demand for wood at 1.39 m3/year/capita surpasses the average annual allowable cut at 0.95 m3/year/capita. The annual wood supply deficit in Tanzania is estimated to be 19.5 million m3. Underlying causes for the wood supply gap include: weak value chains that undermine market access, unsustainable wood production, and harvesting practices. Policies targeted at strengthening the institutional capacity of the forestry sector at local and national levels are suggested to increase the share contribution of the sector to the national economy. To foster small-scale wood production, government needs to expedite efforts in tenure security, and make financial services, extension services, and market information accessible to farmers. Continuous collaboration between the government and actors in the wood value chain is crucial. This policy brief is intended for forest and natural-resource policymakers, wood industrial sector, non-governmental organizations, and academia dealing with the forestry and wood sector in Tanzania.:Key Messages: Page 1 Prevailing challenges: Page 2 Project methods: Page 2 National wood scenarios: Page 3 Wood value chain: Page 4 Policy actions and recommendations to achieve a sustainably wood supply: Page 5 Contact: Page 6

Die Softwareanwendung »Sächsisches Wildmonitoring« löst Papierdokumente zwischen Jagdausübungsberechtigten und Jagdbehörden ab und gestattet eine einheitliche Erfassung jagdrelevanter Daten. Das Internetportal optimiert somit den Verwaltungsaufwand für die Jagdbehörden in Bezug auf Abschussplanung und Streckenerfassung. Redaktionsschluss: 31.10.2016