Forests are a valuable resource for humans providing a range of products and services such as construction timber, paper and fuel wood, recreation, as well as living quarters for indigenous populations and habitats for many animal and bird species. Most recent international political agreements such as the Kyoto Protocol emphasise the role of forests as a major sink for atmospheric carbon dioxide mitigation. However, forest areas are rapidly decreasing world wide. Thus, it is vital that efficient strategies and tools are developed to encourage sustainable ecosystem management. These tools must be based on known ecological principles (such as tree physiological and soil nutrient cycle processes), capable of supplying fast and accurate temporal and spatial predictions of the effects of management on both timber production and carbon sequestration. This thesis had two main objectives. The first was to investigate the environmental factors affecting growth and carbon sequestration of Scots pine (Pinus sylvestris L.) across Scotland, by developing a knowledge base through a statistical analysis of old and novel field datasets. Furthermore, the process-based ecosystem model 3-PGN was developed, by coupling the existing models 3-PG and ICBM. 3-PGN calibrated using a Bayesian approach based on Monte Carlo Markov Chain simulations and it was validated for plantation stands. Sensitivity and uncertainty analyses provided an understanding of the internal feedbacks of the model. Further simulations gave a detailed eco-physiological interpretation of the environmental factors affecting Scots pine growth and it provided an assessment of carbon sequestration under the scenario of sustainable, normal production and its effects from the environment. Finally, the study investigated the spatial and temporal patterns of timber production and carbon sequestration by using the spatial version of the model and applying advanced spatial analyses techniques. The second objective was to help close the gap between environmental research and forest management, by setting a strategic framework for a process-based tool for sustainable ecosystem management. The thesis demonstrated the procedures for a site classification scheme based on modelling results and a yield table validation procedure, which can provide a way forward in supporting policies for forest management and ensuring their continued existence in the face of the present and future challenges.

The accurate assessment of forest persistence under environmental change is dependent on the fundamental understanding of the genetic consequences of human intervention and its comparison to that of natural processes, as declines in genetic diversity and changes in its structuring can compromise the adaptive ability of a population. The European beech, Fagus sylvatica, has experienced prolonged human impact over its 14 million ha range with contemporary forests harbouring high ecological, economic, and cultural value. Historical traditional management practices, such as coppicing and pollarding, have impacted a large portion of Europe’s forests. This form of management encouraged vegetative regeneration, prolonging the longevity of individual trees. In several cases, the structure and function of managed trees and their associated ecosystems were significantly altered. Specifically, coppiced beech forests in Europe displayed significantly larger extents of spatial genetic structuring compared to their natural counterparts, revealing a change in the genetic composition of the population due to decades of management. Humans have also aided in the dispersal of beech within and outside of its natural range. In Great Britain, the putative native range retained signals of past colonisation dynamics. However, these signals were obscured by the wide-spread translocation of the species throughout the country. Evidence of post-glacial colonisation dynamics can be found in Sweden as well. In contrast to Britain, the structure of this natural leading range edge displays a gradual reduction in population size where isolation was found to have acted as an effective barrier to gene flow reducing the genetic diversity of populations.

This research investigates soil respiration (Rs) in a boreal jack pine (Pinus banksiana Lamb.) fire scar chronosequence at Sharpsand Creek, Ontario, Canada. During two field campaigns in 2006 and 2007, Rs was measured in a chronosequence of fire scars in the range 0 to 59 years since fire. Mean Rs adjusted for soil temperature (Ts) and soil moisture (Ms) (Rs T,M) ranged from 0.56 μmol CO2/m2/s (32 years post fire) to 8.18 μmol CO2/m2/s (58 years post fire). Coefficient of variation (CV) of Rs adjusted for Ts and Ms ranged from 20% (16 years post fire) to 56% (58 years post fire). Across the field site, there was a significant exponential relationship between Rs adjusted for soil organic carbon (Cs) and Ts (P = 1.24*10-06; Q10 = 2.21) but no effect of Ms on Rs adjusted for Cs and Ts for the range 0.21 to 0.77 volumetric Ms (P = 0.702). Rs T,M significantly (P = 0.030) decreased after burning mature forest, though no significant (P > 0.1) difference could be detected between recently burned and unburned young forest. Rs was measured in recently burned boreal jack pine fire scar age categories that differed in their burn history and there was a significant difference in Rs T,M between previously 32 v 16 year old (P = 0.000) and previously 32 v 59 year old (P = 0.044) scars. There was a strong significant exponential increase in S R T,M with time since fire (r2 = 0.999; P = 0.006) for the chronosequence 0, 16 and 59 years post fire, and for all these age categories, Rs T,M was significantly different from one another (P < 0.05). The Joint UK Land Environment Simulator (JULES) was used to model vegetation re-growth over successional time at Sharpsand Creek, though it appeared to perform poorly in simulating leaf area index and canopy height. JULES probably over estimated heterotrophic Rs at Sharpsand Creek when Ts corrected simulated values were compared with measured Rs T,M. The results of this study contribute to a better quantitative understanding of Rs in boreal jack pine fire scars and will facilitate improvements in C cycle modelling. Further work is needed in quantifying autotrophic and heterotrophic contributions to soil respiration in jack pine systems, monitoring soil respiration for extended time periods after fire and improving the ability of JULES to simulate successional vegetation re-growth.

The objective of this thesis is to quantitatively understand the economic performance of protected area management strategies for forest and biodiversity conservation. Examples such as integrated conservation and development and eco tourism are assessed in terms of their ability to deliver on welfare benefits to local communities, and an assessment of the opportunity costs of forest conservation as a land use strategy. In addition the contribution of forest conservation in protected areas can make to poverty alleviation and economic development is also examined. The geographical focus of this study is the Albertine Rift region of East and Central Africa, stretching north from the southern end of Lake Tanganyika through the spine of Africa to the northern end of Lake Albert. The Albertine Rift is one of Africa’s most important landscapes for the conservation of forests and biodiversity. The overarching objective is addressed using a series of case studies empirically valuing the opportunity costs of conserving forests in a selection of sites in the central part of the Albertine Rift. The success of conservation is most often measured against progress in reducing habitat or species loss and not often in terms of the contribution of the protected area to poverty alleviation and local economic development. Achieving improvements of conservation strategies in the social dimension requires objective evidence on their effects. Economic valuation of protected area resources provides a quantitative means of assessing the promise and performance of conservation policies in achieving welfare benefits to local communities. This thesis provides three case studies each addressing current valuation and social issues in conservation and sets them in a context of managing protected areas in the broad dynamic setting of poverty alleviation and economic growth from a developing economy perspective. In addition two of the empirical studies are as concerned with methodological enquiry and the performance of novel environmental economic valuation techniques, such as the contingent valuation and choice modelling approaches, as the application of results to conservation questions. The empirical studies show that the benefits to local households and communities from their local forests may be greater than at first perceived. Across all protected area categories, biomes and income groups, households derived significant amounts of their overall income from their local protected area with large proportions of the value of goods harvested from forests being consumed in the home. Amongst income groups high income households often appropriated a greater share of the value of forest goods. There was no significant difference found between the household consumption and the sale of protected area products between income groups. The findings indicate that imposing reductions in forest use may increase poverty amongst local people whilst increasing household income will not necessarily reduce forest exploitation. This indicates that community conservation and integrated conservation and development programmes must target the poor forest adjacent households more actively to ensure poverty alleviation, whilst providing improved protection and law enforcement for effective conservation. It is also shown that biodiversity conservation can have an economic return through mountain gorilla eco-tourism. Findings show a disparity between what constitutes eco-tourism and the real values of tourists towards biodiversity conservation and local social benefits from protected areas. Despite showing a high marginal utility for biodiversity conservation, consumers are unwilling to pay for local community benefits from tourism as part of the permit price to view gorillas. Clearly the link between successful conservation and the welfare status of local communities is not sufficiently established in the minds of consumers to influence their spending decisions. The challenges of effectively mobilising communities to protect biodiversity are discussed in the context of the variable impacts of integrated conservation and development programs over the last three decades. Direct payment payments for conservation services schemes are discussed as an innovative tool to add to the gamut of community approaches currently on offer. Payments for conservation schemes are viewed with cautious optimism in terms of their possibility for success. Despite their allure of being more economically and socially efficient at achieving welfare and conservation objectives, given the complex nature of any society, no less research in to social and economic dynamics of protected area use by local communities would be needed to ensure success of such schemes. However, the overwhelming majority of benefits form protected areas are tied up in ecosystem services values. Mechanisms to generate funding and distribute payments for these benefits in terms of offsetting the local opportunity costs are essential to change local behavior and reduce forest degradation and destruction.

This study reports forest structure, regeneration and the soil properties from unlogged and logged forest in the Bulungan Research Forest, Malinau District, East Kalimantan, Indonesia. Four sites were compared by using four 1-ha replicate plots in each of primary forest (PF), 5, 10 and 30-yr old logged forest (LF-5, LF-10, LF- 30). The tree species composition differ among forest types, as it was shown that the mean value of similarity indices for all pairs were 0.215 (for the Jaccard index) and 0.353 (for the Sorensen index). The low values for similarities among forest types were most probably caused by low numbers of species shared between each forest type. Both correlation values, r = 0.023 for Jaccard index and r = 0.031 for Sorensen index, showed no strong correlation between the similarity index (C) and the distance between forest types. This supports the use of a chronosequence approach. A total of 914 tree species with ³ 10 cm dbh were recorded from 223 genera and 65 families. There were no significant differences in mean species numbers (166 – 180/ha) among treatments. Mean density of species was lower in LF-5 and LF-10 (501/ha) than in PF or LF-30 (605/ha and 577/ha); similarly to mean basal area (LF-5, 28.5 m2/ha; LF-10, 32.6 m2/ha) vs. PF (45.8 m2/ha) and LF-30 (46.9 m2/ha). Dead wood on the forest floor was significantly higher in LF-10 (75 m3/ha) than in the other treatments. Seedlings (< 2 cm dbh) of 1,022 species were recorded from 408 genera and 111 families. The mean number of tree seedling species ranged between 170-206; the mean density of seedlings was about two-fold lower in LF-10 (2790/ha) than in the other treatments. Saplings (>2 – 9.9 cm dbh) of 802 species belonged to 241 genera and 65 families. There was a high variability in species richness across treatments (89 – 191/ha), but not in stem numbers. The Dipterocarpaceae family was dominant in all treatments, followed by the Euphorbiaceae. The soils were acidic, low in nutrients and had low to very low fertility. Both primary and logged forest areas are marginal or not suitable for sustained production of plantation crops. Logging caused soil compaction in LF-30. Although in terms of number of species and trees, amount of BA, number of saplings and seedlings LF-30 appeared to have satisfied prescriptions for a second harvest, ecologically the forest is far from mature. The Indonesian Selective Cutting and Replanting (TPTI) system may need to be revised to a 35 – 45 year cycle to ensure long-term forest productivity in terms of not only timber but other goods and ecosystem services, the value of which are never quantified in monetary terms, but can be higher than the timber revenue.

Les changements environnementaux à l’échelle globale peuvent se traduire par une modification des conditions abiotiques et de la diversité biologique. La fonction de production des écosystèmes forestiers a la particularité d'être à la fois soumise à, et régulatrice de ces changements environnementaux. Il apparaît alors primordial de comprendre la manière dont la diversité en essences et les facteurs abiotiques exercent le contrôle de cette fonction. Au cours de mes travaux j’ai étudié la variation de l'effet du mélange d’essences sur leur productivité dans un contexte climatique et édaphique changeant. Grâce à une double approche, basée sur des données locales et des données de l'Inventaire forestier national, je me suis concentrée sur cinq espèces forestières européennes majeures en peuplements purs et bispécifiques : Quercus petraea, Pinus sylvestris, Fagus sylvatica, Abies alba et Picea abies. En plaine je montre que l'effet du mélange d’essences varie peu avec les facteurs abiotiques. De plus cet effet est déterminé par la tolérance à l'ombrage de l'espèce accompagnatrice. En montagne l’effet du mélange d’essences sur leur productivité est plus fort et positif lorsque les facteurs abiotiques sont les plus limitants pour la croissance. Ces résultats illustrent l’intérêt d'intégrer les facteurs abiotiques et les caractéristiques fonctionnelles des espèces comme déterminants de l'effet de la diversité biologique sur les fonctions de l'écosystème. / Global environmental changes may lead to a modification of abiotic factors and biological diversity. The production function in forest ecosystems has the particularity to be both subjected to, and a regulator of, these environmental changes. Understanding how forest productivity is driven by species diversity and environmental factors is therefore a critical issue. This PhD thesis studies how tree species mixture affects their productivity along edaphic and climatic gradients. Based on an approach using both an original dataset and the national forest inventory dataset, I focused on five major species of European forests in pure and two-species forests: Quercus petraea, Pinus sylvestris, Fagus sylvatica, Abies alba and Picea abies. In lowlands, abiotic factors had little impacts on the outcome of tree species mixture on productivity. In addition, the effect of tree species mixture was determined by the shade tolerance of companion species. In highlands, the positive effects of tree species mixture were strongest when the abiotic factors were the most limiting for growth. These results highlight the importance of considering abiotic factors and the functional characteristics of species as drivers of the effect of biological diversity on ecosystem functions.

Ce travail étudie l’influence des questions environnementales sur le commerce international à partir des échanges de bois tropicaux Malaisie – Europe, la Malaisie étant un important exportateur de bois. Les avantages comparatifs des produits forestiers Malaisiens sont évalués, ainsi que la propension à payer le bois issu de gestion forestière durable par les consommateurs français (en tant qu’Européens). La première partie envisage les différences de perception entre pays développés et pays en développement pour le lien entre commerce et environnement. Il apparaît que les normes environnementales agissent comme des barrières non-tarifaires. Ces barrières sont accentuées par les critères de marquage, d’étiquetage, et de technologie imposés par les pays importateurs. La seconde partie analyse le rôle clé de la Malaisie dans le commerce des bois tropicaux. Les principaux marchés d’exportation des bois Malaisiens sont évalués. Le marché Européen est étudié plus en détail. Il apparaît que les ventes de meuble ont dépassé en 2004 celles des autres principaux produits forestiers. Sur le marché Européen la Malaisie fait face à la concurrence de produits tropicaux à bas prix venant de Chine, et à celle de produits forestiers éco-certifiés venant du Brésil. Concomitamment, l’engagement de la Malaisie dans une dynamique de gestion plus durable y crée une pénurie relative de matériau brut. La troisième partie calcule l’index de Balassa d’avantage comparatif, pour 21 produits forestiers Malaisiens sur le marché Européen. Seuls 5 produits industriels intermédiaires ou à transformation fortement mécanisée, ont un avantage comparatif marqué. Il s’agit des sciages, moulures, contreplaqués, placages, charpente et menuiserie industrielle. Les autres produits présentent des avantages comparatifs faibles ou même négatifs. La quatrième partie estime la propension à payer pour différents attributs environnementaux, ainsi que d’autres tels le commerce équitable et l’origine géographique. Un questionnaire reprenant ces attributs pour du parquet bois hypothétique a été utilisé. Il semble que les consommateurs soient prêts à rémunérer le plus les critères de commerce équitable et d’origine Française, la gestion durable étant recherchée dans une moindre mesure. La propension à payer les tous les attributs varie en fonction des notions et attitudes qu’ont les consommateurs sur l’éco-certification, l’environnement, ainsi qu’en fonction de leur lieu d’habitation, niveau d’éducation et de revenu, et type de profession. Enfin les résultats des 4 parties sont synthétisés en reliant les échelles micro et macroéconomiques, avec les dimensions de demande et d’approvisionnement. D’une façon générale, les résultats suggèrent que les opportunités et contraintes propres à la l’industrie du bois de Malaisie façonnent les exports de produits. La Malaisie s’adapte en se tournant vers des produits à plus haute valeur ajoutée et à moindre impact environnemental, pour pallier aux barrières commerciales et à la pénurie relative de matériau. La Malaisie s’est dotée d’une certification nationale (Malaysian Timber Certification) propre à remplir les critères de durabilité et de légalité de l’Europe, et s’est engagée la gestion durable des forêts. / This study analyses the international timber trade between Malaysia and Europe with respect to the importance of environmental issues on trade and the role of Malaysia as a major timber exporter to Europe. It also evaluates the comparative advantage of Malaysian wood products and the willingness of French consumers (to represent European communities) to pay for sustainable forest management. The first part gives an overview the clashes of perception between developed and developing countries on the environmental concerns over trade. It was observed that environmental standards may act as non-tariff barriers to exporting countries. In addition, the stringent requirements posed by importing countries on technical, marking and labelling to some extent provide unnecessary barriers to trade. The second part deals with the role of Malaysia as a key player in the tropical timber trade. This part evaluates the main export market for Malaysian wood products to the world. For the purpose of this thesis, the analysis focuses on the European market. From the observations, it was found that the export of wooden furniture surpassed major timber exports in 2004. However, to penetrate the European market, Malaysia has to compete with the Chinese with their lower cost tropical wood products, and Brazil with their advantage in certification and labelling of tropical wood products. In tandem with that, the commitment towards sustainable forest management at national level causes shortage of raw materials in Malaysia. To a certain extent, the internal and external factors create necessary challenges to enter the European market. In the third part, the Balassa approach was used to classify the comparative advantage of Malaysia’s twenty one types of wood products in Europe. It was estimated that Malaysia had high comparative advantage only in five products which were mechanized and intermediary industrial products. The products identified were sawn wood, wooden mouldings, plywood, veneer and builders’ joinery and carpentry. The remaining products had lower comparative advantage and disadvantage to export to the European market based on the Balassa index. In the last part, the estimation on the willingness to pay for sustainable forest management attributes was conducted. Besides that, additional attributes such as fair trade and wood origin were included. A questionnaire was set up using all the attributes reflected in the hypothetical wood flooring product in the market. Based on the result, consumers were willing to pay the highest for the presence of fair trade and wood origin (in this study referring to French origin); nevertheless they were still willing to pay for sustainable aspects of forest. However, the willingness to pay for all the attributes was altered depending on the respondents’ knowledge of forest labelling, their attitudes towards environmental preservation, living area, education level, type of job and income level. In the overall finding of the thesis, all the results from each part were synthesized in a systemic approach simultaneously deliberating on the macro and microeconomic perspectives as well as the dimensions on demand and supply. Overall, the findings suggest that the challenges and constraints facing the Malaysian timber industry indirectly shaped the export of Malaysian wooden products. Malaysia has adapted by going into value-added products to lessen the impact of environment-related trade barriers and to circumvent the shortage of raw materials supply. Malaysia has successfully customized the wooden products to the sustainability and legality requirements of the European market by pursuing the national certification (Malaysian Timber Certification) and being committed to sustainable forest management objectives.