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How and why species’ ranges shift has long been a focus of ecology but is now becoming increasingly important given the current rate of climatic and environmental change. In response to global warming, species will need to migrate northward or upward to stay within their climatic tolerances. The ability of species to migrate will determine their fate and affect the community compositions of the future. However, to more accurately predict the future extent of species, we must identify and understand their responses to past and current climatic and environmental changes. The first place change is expected to occur is within ecotones where the ranges of many species converge and individuals exist at the limits of their environmental tolerances. In montane regions, these boundaries are compressed, creating a situation in which even relatively small changes in conditions can lead to shifts in the elevational ranges of species. In this dissertation, I examine the responses of forests in the Green Mountains of Vermont to recent climatic and environmental change in an attempt to understand how future climate change will affect their location and composition. I focus on the Boreal-Deciduous Ecotone (BDE), where the high elevation spruce-fir forests converge with the lower elevation northern hardwoods. In addition to investigating adult trees within the BDE, I also examine the responses of understory herbs and tree seedlings to changes in environmental and climatic factors. Factors considered in these investigations include temperature, soil environment, light environment, invasive species, competition, disturbance and many others. I will examine the complex range of responses in forest species that results from prolonged exposure to these forces alone and in combination. I have attempted to identify the responses of forest species to environmental changes by resurveying historic vegetation plots (Chapter 2), experimentally manipulating the growing environment of tree seedlings (Chapter 3) and performing dendrochronological analyses on tree rings (Chapter 4). Through my resurvey of historic vegetation plots, I determined the degree to which understory species have shifted as individuals or as groups. I also identified a set of novel understory communities that have developed since the 1960's in response to recent climate change, acid deposition and invasive species (Chapter 2). By transplanting and artificially warming tree seedlings, I identified factors responsible for limiting the growth and survival of northern hardwood species above the BDE. Temperature was the primary factor limiting sugar maple (Acer saccharum) at high elevations, while yellow birch (Betula alleghaniensis) was limited almost exclusively by light (Chapter 3). Dendrochronological studies of sugar maples indicated that prolonged exposure to acidified soils has only recently caused growth declines and has altered their relationship to climate (Chapter 4). Together, these studies have produced a cohesive picture of how northeastern montane forests have responded to recent climate change and other anthropogenic impacts. These findings can be used to help predict future species' ranges and identify species that may not be capable of migrating fast enough on their own to keep pace with changes in climatic conditions.

Forest management practices emphasizing stand structural complexity are of interest across the northern forest region of the United States because of their potential to enhance carbon storage. Our research is nested within a long-term study evaluating how silvicultural treatments promoting late-successional forest characteristics affect aboveground biomass development in northern hardwood forests. We are testing the hypothesis that biomass development (carbon storage) will be greater in structural complexity enhancement (SCE) treatments when compared to conventional uneven-aged treatments. SCE treatments were compared against selection systems (single-tree and group) modified to retain elevated structure. Manipulations and controls were replicated across 2-hectare treatment units at two study areas in Vermont, USA. Data on aboveground biomass pools (live trees and coarse woody material, standing dead and downed wood) were collected pre- and post-harvest then again a decade later in 2013. Species group-specific allometric equations were used to estimate live and standing dead biomass and downed log biomass was estimated volumetrically. We used Forest Vegetation Simulator to project "no-treatment" baselines specific to treatment units, allowing measured carbon responses to be normalized relative to differences in site-specific characteristics and pre-treatment conditions. Results indicate that 10 years post-harvest biomass development and carbon storage were greatest in SCE treatments compared to conventional treatments, with the greatest increases in coarse woody material (CWM) pools. Structural complexity enhancement treatments contained 12.67 Mg ha-1 carbon in CWM compared to 6.62 Mg ha-1 in conventional treatments and 8.84 Mg ha-1 in areas with no treatment. Percentage differences between post-harvest carbon and baseline values indicate that carbon pool values in SCE treatments returned closest to pre-harvest or untreated levels over conventional treatments. Total carbon storage in SCE aboveground pools was 15.90% below baseline conditions compared to 44.94% less in conventionally treated areas (P = 0.006). Results from CART models indicated treatment as the strongest predictor of aboveground C storage followed by site-specific variables, suggesting a strong influence of both on carbon pools. Structural enhancement treatments have potential to increase carbon storage in managed northern hardwoods based on these results. They offer an alternative for sustainable management integrating carbon, associated climate change mitigation benefits, and late-successional forest structure.

The health of the northern hardwood forest in the Southern Appalachian Mountains of Tennessee, North Carolina, and Virginia has recently gained attention from the media and environmental stakeholders. This project was designed to examine concentrations of metals, including lead, aluminum, and calcium in growth rings of an important northern hardwood species, American beech (Fagus grandifolia Ehrh.) at Mount Rogers and Whitetop Mountain, Virginia. Dominant and codominant trees were sampled from sixteen research plots located at two different elevations. Samples were crossdated, divided into sections of ten-year periods, and analyzed using atomic absorption spectroscopy. Concentrations of metals were negatively correlated with ring width. Elevation and aspect were significantly associated with the concentration of the cations analyzed. Both lead and calcium concentrations increased during the 1860s and again during the mid-1900s. This increase in lead and calcium may suggest the mobilization of these elements in soil.