Marine reserves have increasingly been recognised for their potential to address the pervasive problem of unsustainable harvest of fisheries worldwide. Biologists advocate the benefits of increased spawning biomass, larger modal sizes and greater densities of fish within marine reserves, and the possibility of spillover to adjacent fishable areas. Bioeconomic studies, however, find that pay-offs from stand-alone marine reserves rarely compete with sustainable yield management schemes, but that they can be beneficial when stocks are heavily exploited. Most of these bioeconomic models are analytical and deterministic in nature, and therefore ignore the redistribution of effort in response to closure and the inherent uncertainty of the marine environment. We present a bioeconomic analysis of a network of no-take areas around Stewart Island in New Zealand applied to the shellfish species paua (abalone) that incorporates both predicted redistribution and reduction in effort, as well as stochastic recruitment. A nested logit model is applied to spatially recorded catch and effort data by the Ministry of Fisheries between 1998 and 2003 to capture the two level decision-making process of divers. On any given day, divers decide whether to go diving at all, and if so, which of the 16 statistical areas around Stewart Island to visit. Weather conditions, spatially varying levels of catch per unit of effort and distance are used as explanatory variables to select areas for closure according to the �least economic impact� in terms of loss of diving trips. An age-structured biological model is developed with parameters specifically applied to paua stocks around Stewart Island. Virgin paua biomass as of 1974 is estimated on the basis of growth, survival, post-larval recruitment and egg production in the absence of fishing. Historic catch rates are then applied to find overall and area-specific levels of exploitation rates, spawning biomass, egg production, legal biomass and numbers of paua. In a final step, the economic model is linked to the biological model to simulate the imposition of no-take areas when taking account of the initial disproportional shift of harvest to fished areas in the first year, and the increase in overall pressure on legal biomass in the years thereafter. We contribute to the marine reserve debate by showing that in the very long run, the overall yield under closure of a relatively small area approaches and even slightly surpasses the yield under no closure for an assumed spillover gradient of 40% despite the redistribution of effort. The most important benefits of marine reserves emerge when stochastic recruitment is included in the recruitment function. In practice, predictions about the stock status and the impact of different harvest levels become much more difficult when acknowledging the inherent variability of the marine environment. The likelihood of stock collapse depends on the assumed value of two recruitment parameters, which highlights the effects of parameter uncertainty and emphasizes the role of marine reserves for population persistence. We also show that under uncertainty average yields under a management regime of a network of no-take areas in addition to the quota system can equal yields under no closure for an assumed spillover gradient of 40%, despite the increased pressure on areas adjacent to the closed areas. Our findings have significant implications for the management of the paua fishery at Stewart Island. For a heterogeneously abundant species, such as paua, spatial management in addition to quota limits could be vital in ensuring the long-term sustainability of the fishery given the inherent variability of the marine environment.

Marine reserves have increasingly been recognised for their potential to address the pervasive problem of unsustainable harvest of fisheries worldwide. Biologists advocate the benefits of increased spawning biomass, larger modal sizes and greater densities of fish within marine reserves, and the possibility of spillover to adjacent fishable areas. Bioeconomic studies, however, find that pay-offs from stand-alone marine reserves rarely compete with sustainable yield management schemes, but that they can be beneficial when stocks are heavily exploited. Most of these bioeconomic models are analytical and deterministic in nature, and therefore ignore the redistribution of effort in response to closure and the inherent uncertainty of the marine environment. We present a bioeconomic analysis of a network of no-take areas around Stewart Island in New Zealand applied to the shellfish species paua (abalone) that incorporates both predicted redistribution and reduction in effort, as well as stochastic recruitment. A nested logit model is applied to spatially recorded catch and effort data by the Ministry of Fisheries between 1998 and 2003 to capture the two level decision-making process of divers. On any given day, divers decide whether to go diving at all, and if so, which of the 16 statistical areas around Stewart Island to visit. Weather conditions, spatially varying levels of catch per unit of effort and distance are used as explanatory variables to select areas for closure according to the �least economic impact� in terms of loss of diving trips. An age-structured biological model is developed with parameters specifically applied to paua stocks around Stewart Island. Virgin paua biomass as of 1974 is estimated on the basis of growth, survival, post-larval recruitment and egg production in the absence of fishing. Historic catch rates are then applied to find overall and area-specific levels of exploitation rates, spawning biomass, egg production, legal biomass and numbers of paua. In a final step, the economic model is linked to the biological model to simulate the imposition of no-take areas when taking account of the initial disproportional shift of harvest to fished areas in the first year, and the increase in overall pressure on legal biomass in the years thereafter. We contribute to the marine reserve debate by showing that in the very long run, the overall yield under closure of a relatively small area approaches and even slightly surpasses the yield under no closure for an assumed spillover gradient of 40% despite the redistribution of effort. The most important benefits of marine reserves emerge when stochastic recruitment is included in the recruitment function. In practice, predictions about the stock status and the impact of different harvest levels become much more difficult when acknowledging the inherent variability of the marine environment. The likelihood of stock collapse depends on the assumed value of two recruitment parameters, which highlights the effects of parameter uncertainty and emphasizes the role of marine reserves for population persistence. We also show that under uncertainty average yields under a management regime of a network of no-take areas in addition to the quota system can equal yields under no closure for an assumed spillover gradient of 40%, despite the increased pressure on areas adjacent to the closed areas. Our findings have significant implications for the management of the paua fishery at Stewart Island. For a heterogeneously abundant species, such as paua, spatial management in addition to quota limits could be vital in ensuring the long-term sustainability of the fishery given the inherent variability of the marine environment.

This thesis examines the implementation of tourism strategies in a rural community context. Applying public policy implementation theory to tourism research, this study investigates questions related to connections and collaborations between stakeholders, decision-making and inducing action and the relation between objectives in a strategy document and actual outcomes of a policy process. Two tourism planning strategies in rural peripheral communities of New Zealand�s South Island are investigated adopting a case study approach. While these strategies have in common their geographical and content foci, they differ in terms of commissioning agencies, stakeholder and community involvement, age, resource allocation and planning and implementation approaches. The strategies are four and eleven years old. One is a top-down strategy by a government agency, the other is the result of a community bottom-up process. Using a pragmatic methodical approach and applying a framework that incorporates the implementation environment including stakeholders and decisive events during strategy making, this research assesses the two cases individually and comparatively. Thus applying a non-linear framework and examining strategies that allow for an investigation of longer implementation timeframes, this study overcomes long-standing issues in this type of research. It fills a gap in the literature as it is the first comprehensive analysis of tourism plan implementation to employ public policy implementation theory in one coherent case study approach. The methods applied in this study include semi-structured interviews as well as content analysis of strategy and policy documents, internal documents such as minutes of meetings and newspaper articles. Starting with strategy makers and �key implementers�, a snowballing process was used to identify further interviewees and to also follow the policy process as perceived by the main actors in it. This combination of methods allowed for a discovery of the policy story; it assisted a process-oriented investigation of tourism plan implementation while at the same time providing factual information and verifying stakeholder statements through triangulating interview results with the content analysis of documents. Overall, for both the top-down and the bottom-up planning case, implementation was found to be dynamic and highly dependent on the actors at the grassroots level. Many critical stakeholders are volunteer community members who have little or no experience in tourism planning. Most actors are tourism operators or are otherwise involved with the industry. However, as some actors are not actively involved in the tourism industry but represent interests related to the protection of the social and natural environment, their views and actions have the potential to be anti-tourism development and therefore they evoke conflict. The two planning approaches do not differ significantly in terms of bureaucratic control and political guidance. Institutions such as, for example, district councils or economic development agencies, mainly impact on the availability of resources to plan and implement strategy goals. Whether or not goals are achieved therefore ultimately depends on the commitment and interest of all stakeholders involved. Other important factors for policy success are the stability of stakeholder relationships and the level of information and knowledge held by those stakeholders, particularly in situations where there is little support from agencies. This is evident where a stakeholder�s previous experience in community work and relevant know-how is considered an asset among the volunteers who are responsible for many of the projects. The potential for conflict and the necessity for implementers to �make do� with the funding, knowledge and information they have, accounts for high volatility of priorities and goals during implementation. Key findings that relate to both the tourism planning and the public policy literature refer to the similarities of implementation processes for top-down and bottom-up strategies at a grassroots level, the significant role and decisive influence that inexperienced volunteer implementers have in such settings and the high degree to which policy change takes place during such processes.

Field evidence from exposed plutonic rocks indicates that mafic-felsic magma interaction is an important process during the construction and evolution of magma chambers. The exhumed, ~140 Ma, Halfmoon Pluton of Stewart Island, New Zealand is characterized by a sequence of mingled mafic sheets and enclaves preserved within an intermediate-felsic host, and provides a unique opportunity to directly investigate the physico-chemical processes that operate within an arc setting. Interpretation of mingling structures and textures in the field, in combination with extensive petrographic, geochemical and geochronological data, allow for conclusions to be reached regarding the nature of the mafic-felsic magma interactions, and the physical, chemical and thermal processes responsible for the generation and evolution of the calc-alkaline magmas. Detailed documentation and interpretation of mafic-felsic magma mingling structures and textures reveal that the Halfmoon Pluton formed incrementally as the result of episodic replenishments of mafic magma emplaced onto the floor of an aggrading intermediate-felsic magma chamber. Physico-chemical processes identified include fractional crystallization and accumulation of a plagioclase – hornblende – apatite – zircon mineral assemblage, episodic replenishment by hot, wet basaltic magmas, magmatic flow and compaction. Early amphibole and apatite crystallization played an important role in the compositional diversity within the Halfmoon Pluton. Variations in the style of magma mingling preserved within the magmatic “stratigraphy” indicate that processes operating within the chamber varied in space and time. Variations in mineral zoning and composition within hornblende indicate that the Halfmoon Pluton crystallized within a magma in which melt composition fluctuated in response to repeated mafic magma replenishments, fractionation, crystal settling and convection. Mineral assemblages, chemical characteristics, isotopic data and geochronological evidence indicate that the amphibole-rich calc-alkaline Halfmoon Pluton was emplaced into a juvenile arc setting, most probably an island-arc. Data are consistent with a model whereby ‘wet’ amphibole-rich basaltic magmas pond at the crust-mantle interface and episodically rise, inject and mingle with an overlying intermediate-felsic magma chamber that itself represents the fractionated product of the mantle melts.

This thesis examines the implementation of tourism strategies in a rural community context. Applying public policy implementation theory to tourism research, this study investigates questions related to connections and collaborations between stakeholders, decision-making and inducing action and the relation between objectives in a strategy document and actual outcomes of a policy process. Two tourism planning strategies in rural peripheral communities of New Zealand�s South Island are investigated adopting a case study approach. While these strategies have in common their geographical and content foci, they differ in terms of commissioning agencies, stakeholder and community involvement, age, resource allocation and planning and implementation approaches. The strategies are four and eleven years old. One is a top-down strategy by a government agency, the other is the result of a community bottom-up process. Using a pragmatic methodical approach and applying a framework that incorporates the implementation environment including stakeholders and decisive events during strategy making, this research assesses the two cases individually and comparatively. Thus applying a non-linear framework and examining strategies that allow for an investigation of longer implementation timeframes, this study overcomes long-standing issues in this type of research. It fills a gap in the literature as it is the first comprehensive analysis of tourism plan implementation to employ public policy implementation theory in one coherent case study approach. The methods applied in this study include semi-structured interviews as well as content analysis of strategy and policy documents, internal documents such as minutes of meetings and newspaper articles. Starting with strategy makers and �key implementers�, a snowballing process was used to identify further interviewees and to also follow the policy process as perceived by the main actors in it. This combination of methods allowed for a discovery of the policy story; it assisted a process-oriented investigation of tourism plan implementation while at the same time providing factual information and verifying stakeholder statements through triangulating interview results with the content analysis of documents. Overall, for both the top-down and the bottom-up planning case, implementation was found to be dynamic and highly dependent on the actors at the grassroots level. Many critical stakeholders are volunteer community members who have little or no experience in tourism planning. Most actors are tourism operators or are otherwise involved with the industry. However, as some actors are not actively involved in the tourism industry but represent interests related to the protection of the social and natural environment, their views and actions have the potential to be anti-tourism development and therefore they evoke conflict. The two planning approaches do not differ significantly in terms of bureaucratic control and political guidance. Institutions such as, for example, district councils or economic development agencies, mainly impact on the availability of resources to plan and implement strategy goals. Whether or not goals are achieved therefore ultimately depends on the commitment and interest of all stakeholders involved. Other important factors for policy success are the stability of stakeholder relationships and the level of information and knowledge held by those stakeholders, particularly in situations where there is little support from agencies. This is evident where a stakeholder�s previous experience in community work and relevant know-how is considered an asset among the volunteers who are responsible for many of the projects. The potential for conflict and the necessity for implementers to �make do� with the funding, knowledge and information they have, accounts for high volatility of priorities and goals during implementation. Key findings that relate to both the tourism planning and the public policy literature refer to the similarities of implementation processes for top-down and bottom-up strategies at a grassroots level, the significant role and decisive influence that inexperienced volunteer implementers have in such settings and the high degree to which policy change takes place during such processes.

Field evidence from exposed plutonic rocks indicates that mafic-felsic magma interaction is an important process during the construction and evolution of magma chambers. The exhumed, ~140 Ma, Halfmoon Pluton of Stewart Island, New Zealand is characterized by a sequence of mingled mafic sheets and enclaves preserved within an intermediate-felsic host, and provides a unique opportunity to directly investigate the physico-chemical processes that operate within an arc setting. Interpretation of mingling structures and textures in the field, in combination with extensive petrographic, geochemical and geochronological data, allow for conclusions to be reached regarding the nature of the mafic-felsic magma interactions, and the physical, chemical and thermal processes responsible for the generation and evolution of the calc-alkaline magmas. Detailed documentation and interpretation of mafic-felsic magma mingling structures and textures reveal that the Halfmoon Pluton formed incrementally as the result of episodic replenishments of mafic magma emplaced onto the floor of an aggrading intermediate-felsic magma chamber. Physico-chemical processes identified include fractional crystallization and accumulation of a plagioclase – hornblende – apatite – zircon mineral assemblage, episodic replenishment by hot, wet basaltic magmas, magmatic flow and compaction. Early amphibole and apatite crystallization played an important role in the compositional diversity within the Halfmoon Pluton. Variations in the style of magma mingling preserved within the magmatic “stratigraphy” indicate that processes operating within the chamber varied in space and time. Variations in mineral zoning and composition within hornblende indicate that the Halfmoon Pluton crystallized within a magma in which melt composition fluctuated in response to repeated mafic magma replenishments, fractionation, crystal settling and convection. Mineral assemblages, chemical characteristics, isotopic data and geochronological evidence indicate that the amphibole-rich calc-alkaline Halfmoon Pluton was emplaced into a juvenile arc setting, most probably an island-arc. Data are consistent with a model whereby ‘wet’ amphibole-rich basaltic magmas pond at the crust-mantle interface and episodically rise, inject and mingle with an overlying intermediate-felsic magma chamber that itself represents the fractionated product of the mantle melts.

The choice of a place to live and reproduce is crucial for species� survival in providing them with adequate resources and shelter from predators or climatic conditions. Determining habitat suitability in endangered species is important for the success of translocation as a conservation tool. In addition, understanding mechanisms (source/sink system versus ecological traps) that drive habitat selection in translocated animals may be critical to population viability. In New Zealand, where ecosystems are highly vulnerable to extinction, habitat restoration on predator-free off-shore islands is an important recovery tool. Therefore, there is a need to understand the relationship between the establishment of the translocated populations and the characteristics of their new environment. Previous research indicated that re-introduced populations of Stewart Island robin (Petroica australis rakiura - Toutouwai) and South Island saddleback (Philesternus carunculatus carunculatus - Tieke) on Ulva Island (Stewart Island), New Zealand, showed preferences for coastal habitats that were characterized by low-lying dense vegetation and open ground cover. In this study, we further investigated territorial establishment in these two populations since re-introduction and looked at how birds utilised the landscape. I hypothesised that sites colonised soon after re-introduction were of high quality and later on, birds moved into unsuitable habitats. I defined habitat quality at a micro-scale in terms of vegetation structure, nest characteristics and food availability. I modeled bird presence and nesting success in relation to habitat components to determine factors in the environment that influenced breeding site selection and contributed to successful nesting in these two species. I discussed results in comparison to similar bird-habitat models developed for the South Island saddleback population on Motuara Island (Marlborough Sounds) and examined explanatory variables in each model. Translocated birds in the three studied populations first established territories in coastal scrub, and in the following years moved into larger coastal forest stands. Although vegetation structure was the primary variable explaining site selection in these populations, vegetation composition should still be considered important as it dictated the suitability of nesting substrate and the availability of food items. There was no evidence that first-colonised areas were more suitable habitats, and I concluded that these cases could not be used as examples of ecological traps. Instead, results suggested that with increased density robins and saddlebacks on Ulva have more recently settled in sites less suitable to nesting and foraging, thus underlying a source/sink structure. However, the sparse distribution of food items on Motuara contributed to a lack of territorial behavior and environmental effect on breeding success; therefore a source/sink system could not be confirmed in this population. I recommended that future translocation sites give preference to mixed-size stands with broadleaved species that are characterised by dense canopy below 4 m height and with suitable cavities in live trees. Lastly, due to robins� and saddlebacks� attraction to conspecifics and their territorial behavior, resources evenly distributed across the landscape could also increase their survival and reproductive success.

The choice of a place to live and reproduce is crucial for species� survival in providing them with adequate resources and shelter from predators or climatic conditions. Determining habitat suitability in endangered species is important for the success of translocation as a conservation tool. In addition, understanding mechanisms (source/sink system versus ecological traps) that drive habitat selection in translocated animals may be critical to population viability. In New Zealand, where ecosystems are highly vulnerable to extinction, habitat restoration on predator-free off-shore islands is an important recovery tool. Therefore, there is a need to understand the relationship between the establishment of the translocated populations and the characteristics of their new environment. Previous research indicated that re-introduced populations of Stewart Island robin (Petroica australis rakiura - Toutouwai) and South Island saddleback (Philesternus carunculatus carunculatus - Tieke) on Ulva Island (Stewart Island), New Zealand, showed preferences for coastal habitats that were characterized by low-lying dense vegetation and open ground cover. In this study, we further investigated territorial establishment in these two populations since re-introduction and looked at how birds utilised the landscape. I hypothesised that sites colonised soon after re-introduction were of high quality and later on, birds moved into unsuitable habitats. I defined habitat quality at a micro-scale in terms of vegetation structure, nest characteristics and food availability. I modeled bird presence and nesting success in relation to habitat components to determine factors in the environment that influenced breeding site selection and contributed to successful nesting in these two species. I discussed results in comparison to similar bird-habitat models developed for the South Island saddleback population on Motuara Island (Marlborough Sounds) and examined explanatory variables in each model. Translocated birds in the three studied populations first established territories in coastal scrub, and in the following years moved into larger coastal forest stands. Although vegetation structure was the primary variable explaining site selection in these populations, vegetation composition should still be considered important as it dictated the suitability of nesting substrate and the availability of food items. There was no evidence that first-colonised areas were more suitable habitats, and I concluded that these cases could not be used as examples of ecological traps. Instead, results suggested that with increased density robins and saddlebacks on Ulva have more recently settled in sites less suitable to nesting and foraging, thus underlying a source/sink structure. However, the sparse distribution of food items on Motuara contributed to a lack of territorial behavior and environmental effect on breeding success; therefore a source/sink system could not be confirmed in this population. I recommended that future translocation sites give preference to mixed-size stands with broadleaved species that are characterised by dense canopy below 4 m height and with suitable cavities in live trees. Lastly, due to robins� and saddlebacks� attraction to conspecifics and their territorial behavior, resources evenly distributed across the landscape could also increase their survival and reproductive success.

Yellow-eyed penguins have suffered major population declines and periodic mass mortality without an established cause. On Stewart Island a high incidence of regional chick mortality was associated with infection by a novel Leucocytozoon sp. The prevalence, structure and molecular characteristics of this leucocytozoon sp. were examined in the 2006-07 breeding season. In 2006-07, 100% of chicks (n=32) on the Anglem coast of Stewart Island died prior to fledging. Neonates showed poor growth and died acutely at approximately 10 days old. Clinical signs in older chicks up to 108 days included anaemia, loss of body condition, subcutaneous ecchymotic haemorrhages and sudden death. Infected adults on Stewart Island showed no clinical signs and were in good body condition, suggesting adequate food availability and a potential reservoir source of ongoing infections. A polymerase chain reaction (PCR) survey of blood samples from the South Island, Stewart and Codfish Island found Leucocytozoon infection exclusively on Stewart Island. The prevalence of Leucocytozoon infection in yellow-eyed penguin populations from each island ranged from 0-2.8% (South Island), to 0-21.25% (Codfish Island) and 51.6-97.9% (Stewart Island). The high prevalence on Stewart Island represented the infection of 100% of chicks and 83% of adult yellow-eyed penguins when tested by PCR. Sequencing of Leucocytozoon sp. DNA found similarities between infections in yellow-eyed penguin adults and chicks, but differences to Leucocytozoon sp. DNA obtained from Fiordland crested penguins. These findings support the suggestion of cross infection between adults and chicks, and indicate that endemic infection in yellow-eyed penguins is unrelated to that in Fiordland crested penguins. Examination by histology and electron microscopy showed tissue megaloschizonts and circulating round gametocytes. Megaloschizonts up to 440µm diameter showed an affinity for hepatic and splenic tissue and were observed releasing occasional intact cytomeres. Round gametocytes were observed within leucocytes in visceral sections, but not peripheral blood smears. The morphology of Leucocytozoon sp. in yellow-eyed penguins showed similarities to the pathogenic species L. simondi and L. sakharoffi but not L. tawaki. A successful treatment protocol for leucocytozoonosis has not been established, although treatment in a Fiordland crested penguin was able to suppress parasitaemia. The role of Leucocytozoon in yellow-eyed penguins as a cause of morbidity and mortality remains unclear. Further investigation into direct pathogenicity, and the interaction of concurrent disease and environmental influences is required. The findings of this thesis provide potential management recommendations and highlight areas requiring further investigation.

Inbreeding depression is one of the factors that can increase the risk of extinction of small populations, and therefore understanding its effects is currently an important issue in conservation biology. Until recently, few studies on inbreeding depression were carried out in wild populations. These recent studies have highlighted the variability in detecting inbreeding depression among natural populations and the multitude of factors that can influence its expression. Many of the factors affecting inbreeding depression in wild populations remain largely unexplored and most of the recent studies in this area have tended to focus on incidents of inbreeding in populations with a history of large population size. The aim of this study is to investigate the relative importance inbreeding depression has had on individual fitness parameters in a population of New Zealand's Stewart Island robins Petroica australis rakiura introduced to Ulva Island. This island population has historically gone through several population bottlenecks. Four main factors that potentially influence the rate of inbreeding and the extent of inbreeding depression, were investigated: environmental variability, life history stage, genetic load and dispersal. Generalized Linear Mixed Modelling was first used to determine how weather affected nest survival. Weather effects were then incorporated into models containing demographic factors to control for environmental variability, and finally parental, maternal and paternal inbreeding co-efficients (=f) were added to models to determine the relative importance of inbreeding depression. Interactions between inbreeding depression and environmental factors were explored. Three different life history stages were compared to determine the differences in inbreeding depression at each stage as well as cumulative effects over time. The genetic load of the population was estimated using lethal equivalents allowing for standardised comparison of inbreeding depression with other species. The likelihood of inbreeding in the population was also explored by investigating the factors affecting dispersal patterns and evaluating evidence for inbreeding avoidance. Inbreeding depression was found to be mild in the robin population. Weather did not have strong effects on nest survival or interactions with inbreeding. Female age was the only factor interacting with inbreeding, with younger inbred females experiencing significantly reduced offspring juvenile survival. Parental and paternal f did not significantly affect brood survival at any life history stage, however, maternal f showed significant effects on nest juvenile survival with the strongest effect occurring when survival was examined cumulatively over all life history stages. The Stewart Island robin had a relatively low lethal equivalent value compared to the closely related North Island robin and other avian species. This difference was associated with the Stewart Island robin having a low genetic load, most likely due to historical genetic purging during periods of population bottleneck. The Ulva Island robin population did not appear to be avoiding inbreeding through dispersal. Dispersal distance was most strongly influenced by the location of the natal nest of the dispersing offspring. In conclusion, the genetic history of the population was likely to have had the strongest impact on the severity of inbreeding depression in the Ulva Island robin population. The results of the thesis highlight the need to examine a number of factors to be able to explain variability in inbreeding depression among populations.