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Stability of coastal systems are threatened by oceanic and atmospheric drivers of climate change. Sea-level rise compounded with increased frequency and intensity of storms emphasizes need for protection of inner island systems by dune formations. Dune building processes are affected by interactions between growth of ecosystem engineering dune grasses and environmental factors associated with disturbance such as sand burial and salt spray. Climate change may also cause latitudinal expansion of some species, resulting in emergence of competitive interactions that were previously absent. Topographic structure of coastlines, traditionally influenced by sand burial, could change as a result of competition emergence. My goal was to determine if species functional trait responses to common abiotic factors are altered by novel and current biotic interactions. I performed a multi-factorial greenhouse experiment by planting three common dune grasses (Ammophila breviligulata, Uniola paniculata, and Spartina patens) in different biotic combinations, using sand burial and salt spray as abiotic stressors. I hypothesized that biotic interactions will cause these dune grasses to shift functional trait responses to abiotic factors that are associated with dune building. I found that plants consistently decreased in biomass when buried. I also found that competition between A. breviligulata and U. paniculata negatively affected dune building function traits of A. breviligulata. This indicates that competition with U. paniculata could alter dune structure. In comparison A. breviligulata had a positive interaction with S. patens, which increased functional trait responses to abiotic stress. Last, we found that competitive intransitivity could occur between these species. My results can be used to make predictions on cross-scale consequences of novel competitive events. This experiment also provides evidence that consideration of local biotic interactions is important in understanding connections between plant level dynamics and large-scale landscape patterns in high stress environment.

Phenomenological models are presented which predicts transpiration rates (E) of individual leaves of Scaevola plumieri, an indigenous dune pioneer, and Ammophila arenaria, an exotic grass species introduced to stabilise mobile sand. In both cases E is predictably related to atmospheric vapour pressure deficit (VPD). VPD is calculated from measurements of ambient temperature and humidity, hence, where these two environmental variables are known, E can be calculated. Possible physiological reasons for the relationships of E to VPD in both species are discussed. Scaling from measurements of E at the leaf level to the canopy level is achieved by summing the leaf area of the canopy in question. E is predicted for the entire canopy leaf area by extrapolation to this larger leaf area. Predicted transpiration rates of individual shoot within the canopy were tested gravimetrically and shown to be accurate in the case of S. plumieri, but less so in the case of A. arenaria. Using this model, the amount of water used by a known area of sand dune is shown to be less than the rainfall input in the case of S. plumieri in wet and dry years. The water use of A. arenaria exceeds rainfall in the low-rainfall year of 1995, while in 1998 rainfall input is slightly higher than water extraction by the plants. Using a geographic information system (GIS), regional maps (surfaces) of transpiration were calculated from surfaces of mean monthly temperature and mean monthly relative humidity. Monthly surfaces of transpiration were subtracted from the monthly median rainfall to produce a surface of mean monthly water deficit. Areas of water surpluses along the coast correspond with the recorded distribution of both species in the seasons that the plants are most actively growing and reproducing. This suggests that unfavourable water availability during these two species growth periods limit their distributions along the coast. In addition to unfavourable water deficits, additional climatic variables that may be important in limiting the distribution of these two species were investigated using a discriminant function analysis.

Arbuscular mycorrhizal (AM) fungi are root and soil inhabiting symbionts with higher plants. The fungi are especially nutrient-function efficient in nutrient deficient soils. There have been innumerable studies of AM fungal facilitation of plant nutrient uptake in controlled environments. Comparatively little similar investigation has been undertaken in natural soils, including investigation of taxon specific nutrient-function efficiency in the phylum. Plant diversity and frequency, soil chemistry statuses, and AM spore diversity and abundance were sampled in an interrupted-belt transect in an aggrading dune sytem on west-coast India, followed by foredunes and transect nutrient amendment experiments in selected plant species. The transect extends 175 m inland from mean high-water mark (MH-WM). Examination showed nutrients were consistently deficient. A plant zonation pattern and increasing frequency over the transect were indicated, as well as decreasing pH and increasing organic matter (OM)-amendment AM species diversity gradients. Plant zonation does not correlate with soil chemistry. There was a distinct soil transition at the 175 m point and evidence of further system partition between foredune and behind-foredune regions. Plant and AM demographies bore no resemblance suggesting neither is driven by the other. Four AM genera were recovered, Acaulospora, Gigaspora and Scutellospora in high abundance, Glomus in comparatively low abundance. The two co-dominant species, A. spinosa and Gi. margarita, displayed divergent strategies in OM amendment. Certain AM taxa may be functionally associated with particular soil nutrients. There was no evidence of taxon-specific nutrient-function efficiency.

Thesis (MScEng (Civil Engineering))--University of Stellenbosch, 2011. / ENGLISH ABSTRACT: The hypothesis postulated in this research, namely that the effectiveness of natural and constructed buffer dune systems can be assessed by a set of indicators that defines the integrity of the dune system and triggers informed management decisions, was evaluated and proved to be essentially true. Two key objectives, namely (1) the identification of key indicators that define the buffer dune integrity; and (2) the development of a scientifically defendable and practical checklist-based method of gathering qualitative information on the identified key indicators so as to guide decision-making at municipal level formed the core of the study. The six dune integrity indicators that collectively define the risk profile of a particular site along the Southern Cape coastline are (1) the degree of protection from prevailing wave energy, (2) the characteristics of the dominant winds and sand supply during the dry season, (3) the relative height of the foredune, (4) the degree of pressure on the buffer dune due to humans, (5) the vulnerability of the type of coastline to erosion, and (6) the coastline stability considering the prevailing coastal processes. The first two indicators relate to the natural (permanent) characteristics of the site and can be defined by experts and presented in the form of a risk and vulnerability atlas layer for direct use by non-experts. The third and fourth indicators relate directly to the implementation of proactive assessment and appropriate management actions to ensure a high level of buffer dune integrity. The last two indicators allow for management intervention to reduce the vulnerability but may entail costly engineering solutions and require expert input. A conceptual risk profile assessment procedure and a decision support guideline incorporating these indicators were developed and evaluated for relevance and practicality through a series of workshops with municipal officials along the south coast of South Africa. It was seen that although some initial basic training may be required, carrying out rapid assessments of the environmental status of key components of an identified human–nature system, such as a buffer dune, is practical and achievable by non-experts. / AFRIKAANSE OPSOMMING: Die hipotese wat in hierdie navorsing gepostuleer is, naamlik dat die doeltreffendheid van natuurlike en geboude bufferduinstelsels geassesseer kan word deur ’n stel aanwysers wat die integriteit van die duinstelsel bepaal en ingeligte bestuursbesluite tot gevolg het, is getoets en bewys hoofsaaklik waar te wees. Twee sleuteldoelwitte, naamlik (1) die identifisering van sleutelaanwysers wat die bufferduinintegriteit bepaal; en (2) die ontwikkeling van ’n praktiese kontrolelys-gebaseerde metode wat wetenskaplik verdedigbaar is om kwalitatiewe inligting oor die geïdentifiseerde sleutelaanwysers in te samel ten einde besluitneming op munisipale vlak te bevorder, vorm die kern van die studie. Die ses duin-integriteitsaanwysers wat gesamentlik die risikoprofiel van ’n bepaalde terrein langs die kuslyn bepaal, is (1) die graad van beskerming teen die heersende golfenergie, (2) die kenmerke van die dominante winde en sandbron gedurende die droë seisoen, (3) die relatiewe hoogte van die voorduin, (4) die graad van druk op die bufferduin as gevolg van mense, (5) die eroderingskwesbaarheid van die soort kuslyn, en (6) die kuslynstabiliteit met inagname van die kusprosesse. Die eerste twee aanwysers het betrekking op die natuurlike (permanente) eienskappe van die terrein en kan deur kundiges bepaal word en in die vorm van ’n kaart in ’n risiko-enkwesbaarheidsatlas aangebied word vir direkte gebruik deur niedeskundiges. Aanwysers 3 en 4 hou direk verband met die implementering van tydige en deurlopende proaktiewe assessering en gepaste bestuursaksies om ’n hoë vlak van bufferduinintegriteit te verseker. Aanwysers 5 en 6 bevorder bestuursaksies om kwesbaarheid te verminder, maar kan moontlik duur ingenieursoplossings inhou en kundige insette benodig. ’n Konseptuele risikoprofielassesseringsprosedure en ’n besluitondersteuningsriglyn wat die aanwysers insluit, is ontwikkel en geëvalueer vir toepaslikheid en uitvoerbaarheid deur ’n reeks werkswinkels met munisipale amptenare aan die suidkus van Suid-Afrika. Hoewel aanvanklike basiese opleiding nodig kan wees, bly dit dat vinnige assessering van die omgewingstatus van sleutelkomponente van ’n geïdentifiseerde mens–natuurstelsel, soos ‘n bufferduin, prakties en haalbaar deur niedeskundiges is.

Une zone d’élargissement/rétrécissement (ZER) du chenal principal de la Loire moyenne (France), incluant une bifurcation asymétrique, est étudiée par courantométrie, bathymétrie multifaisceaux et jaugeages solides. Elle offre une morphologie dont l’évolution est influencée par la morphodynamique des barres sédimentaires. Les forçages induits par les variations de largeur du chenal et la configuration des barres structurent les flux liquides et solides qui, par rétroaction, conditionnent l’évolution des barres. En crue, les relations entre l’hydrodynamique, le charriage et la dynamique des dunes s’ajustent aux variations de débit et engendrent l’érosion du lit en montée des eaux puis son aggradation en décrue. En période de basses eaux, la forte mobilité des sédiments favorise une active morphogenèse. Le charriage total est estimé à 480 000 t pour 2010. La bifurcation module la structure des écoulements seulement après submersion totale du seuil d’entrée du chenal secondaire. L’exportation de la charge de fond vers l’annexe hydraulique semble facilitée par l’érosion du seuil d’entrée ainsi que par la présence d’une barre au pied de cette même marche. / A widening/narrowing area (ZER) of the main channel of the middle reaches of the Loire River (France), including an asymmetrical bifurcation, are studied from hydraulic measurements, multibeam bathymetries and sediment sampling. The evolution of the ZER’s morphology is influenced by the morphodynamics of bars. The forcing effects induced by channel width variations and by bars configuration determine the structure of water and sediment fluxes which affect, in turn, the bars evolution. During floods, the links between hydrodynamics, bedload transport and dunes dynamics adjust to discharges variations and cause bed erosion before the flood’s peak and bed aggradation after. Due to the high sediment mobility, the low flow periods highly affect the morphological evolution of the ZER. In 2010, the total bedload transport reached 480 000 t. The bifurcation influences the flow structure only after the total submersion of the inlet step of the secondary channel. The exportation of the bed-material load toward the secondary channel becomes probably significant after the erosion of the inlet step as well as by the presence of a bar in front of this step.

Un point crucial pour l'avenir de la Physique de Particules est représenté par la mesure de la masse et des paramètres qui gouvernent l'oscillation du neutrino, qui représentent la preuve de Physique au-delà du Model Standard. Depuis 2011, la valeur élevée de l'angle de mélange θ13 a ouvert la voie à l'étude sur la violation de symétrie CP dans le secteur de neutrinos. La nouvelle d'expérience DUNE (Deep Underground Neutrino Experiment) a un potentiel sans précédent pour effectuer une mesure très précise des paramètres d'oscillation de neutrinos. Les grands détecteurs souterrains nécessaires pour ces mesures sont également un milieu idéal pour la recherche sur la désintégration du proton et sur la détection de neutrinos provenant des supernovæ.Les détecteurs utilisés pour ces expériences sont des Chambres à Projection Temporelle (TPC) dont la cible, très massive, est constituée d'Argon liquide. Ce type de détecteur fournit une très bonne résolution de l'image reconstruite, une excellente résolution en énergie et la possibilité d'identifier les particules. Les interactions de neutrino produisent des particules secondaires qui ionisent l'Argon liquide. Les électrons produits par ionisation dérivent sur de longues distances, sous l'effet d'un champ électrique uniforme, jusqu'à rejoindre l‘anode, équipé pour la détection de la charge. Un signal électrique est alors produit et est utilisé pour la reconstruction 3D de l'interaction primaire. La TPC en double phase liquide-gaz permet l'amplification du signal d'ionisation par des avalanches qui se produisent dans la phase gazeuse, au-dessus du niveau de l'Argon liquide. Cette technique améliore les performances de TPC en augmentant son rapport signal-bruit.Le sujet de cette thèse est la reconstruction et l'analyse de la charge de ionisation dans une TPC à Argon liquide : la mesure de la charge déposée par ionisation fournit des informations sur l'énergie cinétique des particules chargées secondaires produites lors de l'interaction de neutrino. De cette manière, il est possible de reconstruire l'énergie du neutrino entrant, de reconnaître et d'exclure les gerbes électromagnétiques produites par la désintégration du π0, puis d'effectuer l'indentification des particules à partir de la mesure des pertes d'énergie par ionisation.La mesure de l'ionisation implique une connaissance approfondie de la réponse du détecteur et de l'algorithme de reconstruction. Afin d'atteindre cette connaissance nous avons effectué une analyse détaillée des pertes d'énergie simulée en étudiant les divergences entre les connaissances théoriques et la simulation. Ensuite, la simulation de la réponse du détecteur a été étudiée, en examinant les effets qui se produisent au cours de la dérive des charges et les effets liés à la réponse de l'instrumentation électronique. Ces effets systématiques qui affectent l'exactitude de l'algorithme de reconstruction sont alors caractérisés par rapport à la génération de Monte-Carlo.Par la suite, nous montrons comment il est possible d'effectuer la rejection du π0 en étudiant les pertes d'énergie. En mesurant les pertes par ionisation au début d'une gerbe électromagnétique, il est possible de comprendre si elle a été produite par un électron ou par un photon. Cela permet d'exclure le bruit dans la détection des produits d'interaction du neutrino électronique fondamental pour la recherche de la violation de CP.Par l'échantillonnage de la trajectoire d'une particule chargée et en mesurant ses pertes d'énergie, il est possible d'identifier sa nature. Une TPC à Argon liquide est également un milieu idéal pour la recherche de la désintégration du proton, en particulier en regardant certaines chaînes de désintégration exclusives, comme p  K + ν. Dans cette thèse, nous montrons ainsi comment il est possible d'identifier des particules à partir de la mesure des pertes d'énergie, et plus abordons plus précisément l'identification des kaons chargés / This is an important period for High Energy Physics: many recent results, including the Higgs discovery and its characterization, confirm the Standard Model. A crucial point for the future of Particle Physics is the study of neutrino masses and mixing representing the first established evidence of physics beyond the SM. Since 2011, the large value of the ?13 mixing angle opened the way to the investigation of CP violation in the neutrino sector. A next generation long baseline neutrino experiment (DUNE) has unprecedented potential to precisely measure the neutrino oscillation parameters, determine the neutrino mass hierarchy and has a very good chance to discover evidence for CP violation in the leptonic sector. The large underground neutrino detectors needed for this task will also address the search for proton decay and the observation of supernovae neutrinos. Giant Liquid Argon Time Projection Chambers (LAr TPCs) will be employed as neutrino targets and detectors. They provide bubble-chamber quality imaging coupled to excellent energy resolution and particles identification capabilities. Neutrino interactions produce secondary particles, which ionize the liquid argon. The ionization electrons drift for long distances along a uniform electric field until they reach finely segmented and instrumented anodes, producing electrical signals that are used for 3D imaging and analysis of the primary interactions. The dual-phase readout technique foresees the amplification of the ionization signal in avalanches occurring in the gas phase above the liquid argon level. This technique further enhances the performance of the LAr TPC by increasing its signal to noise ratio. The subject of thesis is the ionization charge reconstruction and analysis in the dual-phase LAr TPC: the ionization charges measurement provides information about the kinetic energy of secondary charged particles produced in neutrino interactions. In this way, it is possible to reconstruct the incoming neutrino energy, identify and reject electromagnetic shower generated by photons from pi0 decay and perform particles identification from the measurement of the specific ionization losses.The measurement of the ionization implies a detailed knowledge of the detector response and of the reconstruction algorithm. In order to achieve this knowledge a detailed analysis of the simulated energy losses has been performed by studying the differences between the theoretical knowledge and the simulation

The importance of understanding the ecological functioning of coastal dune systems is emphasized as being fundamental to the correct management of the dune landscape. Dune vegetation along the Eastern Cape coastline, from Cape St Francis in the west to Kei Mouth in the east was described in terms of the distribution and phytochorological affinities of the taxa. At a regional scale species distribution was strongly influenced by both the climate, particularly rainfall, and the phytochorological affinities of the taxa. Seven communities were defined using TWINSPAN, and the interrelationships between these communities in anyone area appeared to be linked to a successional gradient. Dune Slacks are thought to play a key role in this successional sequence, and a temporal study of this community led to a conceptual model of plant succession in these dunefields. Climate, particularly rainfall and wind, are major factors influencing plant succession. Wind-borne sand causes the slacks to migrate in an easterly direction under the influence of the predominantly westerly winds, although easterly winds, mainly in summer months may reverse these trends. Autogenic changes appeared to be important in this succession, and a comparative study of a good example of a primary succession at Mtunzini in Natal was undertaken to elucidate the main mechanism of change. Eight communities that were identified here were concluded to be distnbuted along a gradient of increasing age, with successional changes predictable, linear and directional. Species were grouped in distinct zones along the continuum and edaphic changes (decrease in soil pH, increase in organic matter and exchangeable bases) were related to the community based changes in species composition. The mechanism of change supported the facilitation model of plant succession which is a modification of the original Clementsian concept. Similar results were found in the Eastern Cape, but due to the harsh environment, multiple pathways of succession exist. Data from this study lent support to the model of plant succession developed earlier, and confinned that the dune slacks played an important role in this facilitation by acting as centres of diversity. The foredunes were found to have an indirect role in protecting these slacks from salt spray and sand movement. The central theme of the management studies was to investigate the ecological consequences of recreational pressure within the dune environment. Current levels of beach utilization at East London were lower than other beaches in South Africa, but a general trend of increasing utilization due to sociopolitical changes can be expected. The suitability of questionnaire surveys to assess aspects such as the adequacy of facilities, perceptual carrying capacity and the beach users opinion of natural vegetation and preference for particular beaches was demonstrated. The dune vegetation was found to be sensitive to human trampling, but at current levels, the ecological carrying capacity will not be exceeded since results of the aerial census counts and questionnaire survey revealed that few people entered sensitive zones such as the coastal forest. More detailed long term studies on the susceptibility of dune vegetation to both trampling and off-road vehicle impacts revealed a low resilience of dune plant communities to these effects. Although susceptibility differed between the three communities tested, generally the greatest amount of damage occurred after the first few passages, and vehicles caused a more significant decrease in height when compared to trampling. Recovery rates were slow and low levels of repeated damage were sufficient to retard or prevent the recovery of the plants. Stricter control of vehicle use on beaches is therefore required, and in high use zones the ecological carrying capacity should be increased by providing access tracks if possible, or if not possible, by restricting access. A historical account of the process of dune stabilization showed that although first initiated in 1845, indigenous species were only used in the past three decades. The use of alien species has resulted in problems such as a reduction in the ecological integrity and aesthetic appeal of coastal systems. The techniques applied in the stabilization of drift sands with indigenous vegetation have been successful, as revealed by a quantitative survey of 17 sites in the Eastern Cape. Sites were grouped by multivariate analysis on the basis of their species composition, and variability between sites was dependent on the types of species planted. Selection of suitable species is therefore important and is discussed with respect to their natural distribution along the coast. The long term objective of stabilization should be the creation of functional, diverse, aesthetic ecosystems, since the intrinsic and economic value of the dune landscape for tourism lies therein. However, detailed studies should be undertaken prior to implementing a manipulative process such as dune stabilization, since ecological processes may be disrupted. An understanding of such processes is therefore important if one wishes to effectively manage the dune landscape.

Ammophila arenaria (L.) Link is a European sand binding plant which was introduced to South Africa in the 1870's for the purpose of dune stabilisation. Because of its known invasiveness along the west coast of North America, and the problems South African ecosystems experience with alien invader plants, it was deemed necessary to study the biology and ecology of this species in South Africa. The aim of this thesis is to establish the potential invasiveness of A. arenaria on Cape coastal dunes and assess whether its use for dune stabilisation is still justifiable. A. arenaria occurs nowadays between the Langebaan area on the west coast and Gonubie in the Eastern Cape. Although widespread, the grass appears to occur only in areas where it has been planted. Its unaided spread may be prevented by adverse climatic conditions. Studies on the community biology of South African A. arenaria communities as compared to indigenous dune plant communities and natural A. arenaria communities in Europe cannot confirm the aggressive behaviour that A. arenaria shows in California and Oregon. In South Africa, A. arenaria does not exert strong floristic control over other species or outcompete and replace them, neither does it alter the topography of South African beaches and dunes. It forms weaker species associations and tends to develop communities of little species variability along the coast, thereby proving its alienness in South Africa, but this does not imply its invasiveness. Studies on succession of A. arenaria stabilisation areas show that monospecific A. arenaria plantings can be succeeded by a species-rich indigenous dune scrub or dune fynbos within a few decades. Plant-parasitic nematodes have been recorded, which may play an important role in the succession of A. arenaria stands in South Africa as was observed in Europe. Monitoring of A. arenaria communities and indigenous communities over nearly three years shows that A. arenaria is not spreading and replacing indigenous plants but in fact rather being replaced by the latter. A. arenaria profits from a superior sand burial tolerance but is affected by adverse climatic factors, mostly the lack of rainfall and strong radiation. In comparison to the indigenous dune grasses Thinopyrum distichum and Ehrharta villosa, it does not show any superior demographic traits such as an unusually high growth rate or large aboveground biomass production. Although A. arenaria produces viable seed in South Africa, the indigenous grasses show better germination and seedling establishment in the field. This study indicates that A. arenaria is not invasive in South Africa, nor likely to become an invader species in the near future. However, more research is required to confirm these results and more caution recommended regarding the further use of this alien grass for dune stabilisation.

No description available.