Thesis (Ph. D.)--University of Western Sydney, Hawkesbury, 1996. / Includes bibliographical references (p. 511-557).

Thesis (Ph. D.)--University of Hawaii at Manoa, 2003. / Includes bibliographical references.

Thesis (M.S.)--University of North Texas, Dec., 2009. / Title from title page display. Includes bibliographical references.

The prairie communities that once dominated the landscape of Illinois have been reduced to a fraction of their former extent. Subsequently, considerable effort has been invested in the restoration of these lost communities, yet the comprehensive assessment of restoration success has only recently garnered interest. The objectives of this study were 1) to gauge the success of a prairie restoration project by measuring the components of ecological fidelity (structure/composition, function, and durability), and 2) to determine the factors that influenced success. Nineteen prairie plantings (ranging from two to 19 growing seasons old) at The Nature Conservancy's Nachusa Grasslands preserve were chosen for the assessment. Floristic quality was calculated to assess the composition component of ecological fidelity. Aboveground net primary productivity, soil bulk density, total soil nitrogen and total soil carbon were measured to assess the function component (soil measurements were taken at 0-5 and 5-15 cm depths). Results were compared to benchmark values taken from the literature and from samples of remnant prairies. Durability was determined by comparing measurements across a restoration chronosequence. To further evaluate the prairie plantings and restoration success, non-metric multidimensional scaling ordination was used to compare plantings based on their vegetation composition and soil characteristics (prairie remnants were also included in the comparison based on soil characteristics). Values of Mean C and FQI indicated successful levels among younger plantings, but durability was less successful according to the chronosequence. Seed-mix quality had the greatest influence (positive) on composition success. Aboveground net primary productivity levels were successful and durable overall, however, younger plantings exhibited successful levels of production more consistently than older plantings. Aboveground net primary productivity was most influenced (negatively) by the abundance of the exotic C3 grass genera Poa and Bromus. Functional success based on soil characteristics was limited. Soil bulk density, total nitrogen, and total carbon levels all differed among plantings and remnants at both depths, and evidence of levels recovering toward levels of remnants was not detected. The results of this study indicated that some components of ecological fidelity have been successfully restored, while others have not, and using a high-quality seed mix that resembles the species pool of remnant prairie and limiting the abundance of the dominant native C4 and exotic C3 grasses can improve the restoration of plant composition and ecological function in Illinois prairie plantings. The mixed results underscore the importance of examining more than one component of ecological fidelity when measuring success. Long-term monitoring is also recommended for evaluating restoration durability, especially for detecting changes among soil properties over time.

No description available.

The grasslands of South Africa are the foundation of commercial and subsistence agriculture yet they are being degraded at an alanning rate. The coastal grasslands of the Eastern Cape are no exception and they suffer added pressure of coastal resort development because of their proximity to an attractive coastline. In order to determine the degree of conservation necessary for any area it is essential to know what species occur there in order to determine if protection from habitat destruction is required. Four aims were defined for this studv. The initial aim was a phytogeographical classification of the grasslands in the coastal region west of the Kei River. This was done by vegetation sampling followed by computer based analysis with TWINSPAN. This analysis defined ten grassland associations. five being located in the area west of the Keiskamma River and five occurring east of it. The associations in the eastern half are termed mesic while those in the western balf are xeric. The second aim was to determine the presence of any underlying ecological gradients affecting the distribution of tbe associations. Indirect gradient analysis was carried out where samples are analysed irrespective of environmental factors. Direct gradient analysis was then carried out using scores per sample of various environmental factors. Environmental factors whicb migbt produce such gradients are both naturally-ocurring and man-induced. Natural factors which were recorded in the field include depth of soil at sample site, soil family, aspect and distance from the shore. An important factor is the land / sea interface. Natural factors analysed in the laboratory include soil pH. conductivity. percent organic matter, calcium, magnesium, phosphate and potassium. Man-induced factors are land use history e.g. Ploughing and grazing. Both direct and indirect gradient analysis were carried out with the computer based programme CANOCO. The third aim of the study was to determine the presence of any successional trends between the ten defined associations. This was done using several characteristics of the associations. Alpha and beta diversity were the first factors compared between associations. The percentage contribution of the Cyperaceae, Fabaceae, Asteraceae and Poaceae to each association was assessed and compared. The presence of various life forms of the species occurring in each association was determined together with the cover abundance of different classes of grass species. These results were then combined and analysed in the light of the results from CANOCO analvsis. They show that the mesic Themeda Iriandra - Anthospermum herhoceum, association which occurs closest to the shore and with the least disturbance is a depauperate form of the mesic climax Themeda triandra association. The mesic Hyparrhenia hirta - Dtheteropogon amplectens association may be a secondary grassland on account of phosphate and potassiun poor soils and land-use. The mesic Hyparrhenia hirta - Themedo triandra association is a slightly degraded form of the climax association due to grazing. The severely disturbed Stenotaphrum secundatum - C'entella coriacea association, which is located close to the shore, is a secondary grassland. Analysis of the xeric associations indicates a clearly defined ordination of associations on the basis of land-use. The Cynodon daclylon - Helictolrichon hirtulum and Sporobolis africanus - Setaria sphacelata associations which occur where ploughed lands have been left to lie fallow are secondary in nature. The Themeda triandra - Ehrharta calycina association is thought to be the closest representati ve of a climax xeric association but the presence of E. calycina indicates that some disturbance has occurred. The vegetation is subject to moderate grazing. The Cynodon dactylon - Ehrharla calycina and Diheleropogon filifolius - Ehrharla calycina associations are subject to varying intensities of trampling and grazing and are degraded forms of the Themeda triandra - Ehrharla calycina association. Because of tbe overriding influence of the land-use gradient separation along gradients of the remaining eight factors was limited. The effect of fire and temporal change in the mesic Themeda triandra and Hyparrhenia hirta - Themeeia trianda associations was assessed via long-term studies at Potter's Pass outside East London. The results show a quick response to burning with a return to 100% cover within six months by both associations. During spring the two associations could not be separated by either TWINSPAN or DECORANA. indicating a temporal shift from the Hyparrhenia hirta - Themeda triandra association to the Themeda triandra association. The fourth and final aim was to identify plants requiring protection from habitat destruction and to make recommendations for further conservation areas and management of the coastal grasslands. Through the collection and identification of as many plants as possible a species checklist was prepared and the status of each was determined through reference to the Red Data Book of threatened and endangered species. Kniphofia rooperi is vulnerable in the Cape and Euphorbia hupleuroides is considered rare in kwazulu-Natal. Ten species are endemic to the Cape and / or South Africa. The richness of the vegetation lies in the presence of species representative of the four major floras which converge in the eastern Cape. i.e. The Cape, NamaKarroo, Tongoland-Pondolaod floras and the Kalahari Highveld Regional Transitional Zone. Suggestions are made for the conservation of the grassland associations at various sites within the study area based on the present area conserved. Less than 2% of the coastline in the study area is conserved and it is felt that the area under conservation should be increased. Management suggestions based on the available literature are given for both farmed and conserved areas. There is a great potential for further studies on grassland dmamics within the areas surveyed. In terms of management, the grasslands require careful examination to determine the most effective season in which to burn if at all. As with most agricultural systems in South Africa. grazing strategies will benefit from further research. Any additional research on the dynamics of these grasslands can only be of benefit to the sustained utilisation of this vital resource.

The aim of this study is to gain a predictive understanding of the patterns and determinants of plant biodiversity in temperate, mesic grasslands of South Africa with a primary focus on the geographical area of the Eastern Cape. From a review of the literature on hypotheses explaining diversity (Chapter 2) it was possible to formulate a number of hypotheses that could be tested to explain species richness patterns in Eastern Cape grasslands. This thesis is organised so that each main chapter deals with a specific body of theory concerning the explanation of diversity patterns. A detailed description of the study area is provided (Chapter 3), including environmental variation and a description of major vegetation patterns. A summary is provided of grassland plant community patterns, as determined by phytosociological studies in the study area. A multivariate analysis of environmental variables was undertaken to determine which variables contributed the most towards explaining environmental variation in the study area and to determine whether any variables co-vary, a possible problem for any multivariate analysis in later chapters. Altitude produced one of the strongest gradients in the study area. There were a number of variables that were correlated with altitude, most notably temperature. Rainfall co-varied partially with altitude, but there was also a strong rainfall gradient perpendicular to the altitude gradient. A description of species richness, diversity and evenness patterns at the plot scale within different grassland plant communities of the Eastern Cape is provided in Chapter 4. To determine whether the environment acts differently on different growth forms, the contribution to species richness by different major growth forms is analysed. Furthermore, since the majority of literature attempts to explain diversity in terms of environmental factors, it was necessary to analyse the relationship between species richness and various environmental variables. The results indicate that there is high variation in species richness both within and among grassland communities. Forbs make the most significant contribution to overall species richness per 100 m2, followed by grasses. Variance in richness of all species together is not significantly related to environmental variables in mesic grasslands, but is significantly related to environmental variables in semi-arid grasslands. The result of greatest interest from this chapter is the fact that richness amongst different life-forms in the same place is explained by different environmental factors, indicating that the environmental factors that affect coexistence of species have a different effect on different life-forms. A classification of all the species of the dataset into plant functional types using a multivariate approach based on functional traits was conducted (Chapter 5). The grass species were classified into 16 functional types and the forbs into 14 functional types. The functional type classification provided the opportunity for undertaking analyses to develop an understanding of 8 the contribution by niche differentiation towards promoting species richness (Chapter 6). The results provide evidence of niche differentiation in the grasslands of the study area and also that niche differentiation promotes species richness in the grasslands of the study area. It was found that higher rainfall grasslands are less structured by niche differentiation than semi-arid grasslands. A regional / historical analysis is undertaken (Chapter 7) to investigate the relationship between the regional species pool and local richness, and the relationship between local richness and phytochorological diversity. Regional richness appears to have little effect in promoting local richness in grassland plant communities of the study area except at sites where there is high local richness. This provides an indication that regional richness only promotes local richness in the absence of local limiting factors. Phytochorological diversity promotes local richness, but mostly through diversity amongst species with narrow distribution ranges. Some theories ascertain that seasonal uncertainty may provide opportunities to species that would otherwise be outcompeted and thereby promote local richness. The degree to which seasonal uncertainty and seasonality promote local richness in the Eastern Cape grasslands was therefore investigated (Chapter 8). A weak relationship exists between these variables and local species richness in grassland communities of the study area, indicating that they do not promote niche differentiation to a significant degree in the study area. It is clear that in the grasslands of the Eastern Cape, environmental limiting factors are more important in semi-arid grasslands and species interactions are more important in mesic grasslands for structuring plant communities (Chapter 9, Discussion). Regional processes do not appear to be important in structuring local communities, but the analysis undertaken in this study shows that they may be significant when factors operating at the other two levels are overcome (species interactions and environmental limiting factors.

In this study, I investigated the interaction between two megaherbivores, the African elephant and white rhinoceros, that has the potential to impact grazing lawns of which white rhino are the creators and maintainers and elephants are potentially the modifiers. I hypothesized that as elephants browse, they discard a variety of coarse woody debris onto the ground; should this woody debris (of varying amounts and sizes) fall onto grazing lawns, white rhino either move them, consume grass around the woody debris or abandon the lawn entirely. If high levels of woody debris are deposited here, grazing by white rhino is likely to be prevented, at which time I predicted that mesoherbivores would have a competitive advantage in accessing forage that white rhino cannot. I examined the mechanistic links between different levels of elephant-deposited woody debris and grass response at a point scale and feeding patch spatial scale of grazing lawns in an African savanna. In addition, I assessed the response of mesoherbivores in terms of vigilance behaviour with increasing levels of predation risk posed by increasing levels of woody debris. I present the first evidence of an indirect effect of elephant on white rhino foraging behaviour. I demonstrate how increasing levels of woody debris lead to a decreasing probability of foraging by white rhino. I also demonstrate how the probability of foraging by mesoherbivores increases as the amount of forage increases. However, since this study took place during a severe drought where resources are extremely limited, I was unable to properly separate the effects of elephant-deposited woody debris from the severe lack of rainfall on grass response and subsequently herbivore foraging behaviour. Due possibly to the drought, mesoherbivores responded less or not at all to risk factors such as woody debris therefore woody debris was not a predictor of vigilance behaviour in my study. This study contributes to our understanding of how the impacts of elephants, as ecosystem engineers, have cascading effects on savanna ecosystems. My study showed that elephant impact mediates the foraging behaviour of white rhino during a drought. However, under average rainfall periods, my original hypothesized effect of the indirect impacts of elephants on white rhino foraging and grazing lawn dynamics could still hold. This key hypothesis that I was unable to test under ‘normal’ conditions due to the drought is still valid and functionally important for understanding the ecosystem processes driving grazing lawn formation, persistence and composition in African savannas where elephants and white rhinos coexist.

A thesis submitted to the Faculty of Science, University of the Witwatersrand, Johannesburg, in fulfilment of the requirements for the degree of Doctor of Philosophy / The study aimed to differentiate the effect of rainfall and grazing on the compositional change of savanna grasslands, and to explain responses in terms of the life history patterns of the main species, An extinction prone perennial grass under heiry grazing was identified as a palatable, obligate seed reproducer with low seed production and poor dispersal (e.g. Themeda triandra and Reteropogon contortus), wnich could become extinct if the established population and the seed bank were eliminated. Aristrida species were expected to show the converse pattern. Grazing was manipulated for three or five years, and rainfall for one year, on a sandveld site, and two clay soil sites which supported the same species in different amounts, respectively. (Abbreviation abstract) / Andrew Chakane 2019

A long-standing question in ecology is understanding how the environment structures species occupancy in space and time. Specifically, identifying associations between environmental drivers - climate and land cover - and species occupancy is crucial to predicting species distributional dynamics in the future. Over the last century, research on the abiotic drivers of species occupancy has largely focused on temperate regions. Tropical mountain ecosystems harbor extraordinary levels of diversity and face some of the highest anthropogenic pressures of climate and land cover change. Yet, such regions have remained historically understudied. Bird species, due to their sheer diversity and occurrence across climatic zones and land cover types, are an ideal model for understanding how climate and land cover structure occupancy in space and time. The goal of this dissertation is to disentangle spatial and temporal associations between environmental drivers - climate and land cover - and bird species occupancy along a tropical montane gradient. I use an integrative approach that relies on citizen science, historical ecology, and bioacoustics to study bird communities in the Western Ghats biodiversity hotspot of southern India. In Chapter 1, we used observations from the world’s largest citizen science database, eBird, to ask how contemporary climate and land cover are associated with bird species occupancy across the Nilgiri and the Anamalai hills of the Western Ghats. We show that the occupancy of several forest specialist birds was negatively associated with temperature seasonality, highlighting narrow thermal niches for such species. We also show that a small number of generalist bird species are positively associated with human-modified land cover types. In Chapter 2, we combined colonial-era and modern datasets on bird species observations and land cover to ask how a century of landscape change across the Nilgiri hills has impacted bird communities. Between 1848 and 2017, 75% of grassland habitat across the Nilgiri hills was lost toward timber plantation and cash crop expansion. Such drastic declines in grassland habitat have resulted in declines in species persistence and relative abundance of grassland specialist birds over the last century. As a result, the functional trait space has undergone biotic homogenization. In Chapter 3, we ask if the reversal of landscape changes significantly affects bird communities. Using passive acoustic monitoring, we examined the impacts of ecological restoration on bird communities and other vocalizing fauna along a gradient of forest regeneration (consisting of actively restored, naturally regenerating, and mature benchmark sites) in the Anamalai hills. Encouragingly, we show that the bird community composition of actively restored sites is transitioning toward mature benchmark sites. However, when we moved beyond birds, we found that vocalizations at higher frequencies (> 12 kHz) were largely missing from actively restored and naturally regenerating sites, while the same frequency space was occupied in mature benchmark sites. Taken together, we find that climate and land cover are key determinants of bird species occupancy in the Western Ghats, and in a globally changing world, conservation interventions such as ecological restoration along with the preservation of naturally occurring land cover types are key to sustaining montane avifauna in the long run.