The permanent coresidence of males within a troop is unusual but occurs in vervet monkeys. Several hypotheses have been projected to explain the coexistence of male vervets (predation risk, breeding season length) but these hypotheses fall short in explaining the multimale nature of vervet monkeys. In order to determine the explanation for coresiding males, I collected male behavioural data from two troops over the course of nine months. My dataset was divided into two categories, male-male interactions and female-male interactions. The male-male data indicate that breeding season is the most active time for migration, aggressions and wounds. Coalitions were described for the first time, and affiliative interactions between males highlighted coping tactics of males in regards to their coexistence. The female-male data indicate there was little indication for distinct male or female choice for mating. Specifically, olfactory information appears to lower successful copulations of males, and female resistance also decreased successful copulations. Grooming was not a commodity traded for mating access. Male dominance was not correlated with mating success, and females seem to express their preference for sexual partners. The large cohort of males of my troops appears to alter behaviours observed at other sites. The maintenance of male-female associations after breeding season suggests that males may be preparing for next breeding season, and males may co-reside for breeding purposes. / xiii, 123 leaves : ill. maps ; 29 cm

This study explored the organizing principles of female sociality in free-ranging vervet monkeys (Chlorocebus aethiops) inhabiting the Klein Karoo, South Africa. Females groomed more than males, grooming peaked at the end of the day and less grooming occurred during the mating season. Although females competed over food, they did not compete over grooming partners, rarely formed coalitions and did not trade grooming against other activities. Instead, they maintained grooming whilst trading between feeding and resting and feeding and moving. Despite seasonal shifts in food competition, grooming was not traded for tolerance and there was an upper limit to cohort size before clique size declined. Inter-population comparisons revealed no troop size effects on clique size, aggression and competition over high-ranking grooming partners. The rarity of coalitions suggests coalitions are unlikely to be a central component of female relationships. / xiii, 162 leaves : ill., maps ; 29 cm

Vervet monkeys (Chlorocebus pygerythrus) are generally described as a territorial species, though variations in territoriality have been observed. This research examined the impact of high population density, large group sizes and extensively overlapping home ranges on the expression of territoriality in a population of vervet monkeys. Over a period of eighteen months, data were collected on three large troops of vervet monkeys (PTN = 31, RBMN = 39, RSTN = 57) on the Samara Game Reserve, South Africa. I examined the spatial dynamics demonstrated in this population, assed the extent and use of home range overlap and examined the occurrence of intertroop encounters in relation to mate defence, resource defence and home range defence. The home ranges of the troops in this study population overlap substantially, and overlap areas are exploited extensively. The frequency of occurrence of intertroop encounters does not increase in the mating season, nor is there evidence of the defence of particular food patches or home range defence. The findings from this thesis emphasize the importance of examining the expression of territoriality of a population within the context of its environment. / xvi, 125 leaves : ill. ; 29 cm

Narrow riparian woodlands along non-perennial streams have made it possible for vervet monkeys to penetrate the semi-arid karoo ecosystem of South Africa, while artificial water points have more recently allowed these populations to colonize much more marginal habitat away from natural water sources. In order to determine the sequelae of life in these narrow, linear woodlands for historically 'natural' populations, I determined the size of troops in relation to their reliance on natural and artificial water sources and collected detailed data from two river-centred troops on activity, diet and ranging behaviour over an annual cycle. These data indicate that river-centred troops were distinctive primarily for their large group sizes and, consequently, their large adult cohorts, and the extent of home range overlap in what is regarded as a territorial species. While large group size carried the corollary of increased day journey length and longer estimated interbirth intervals, there was little other indication of ecological stress. Specifically, the rate of predation appears to be lower than observed at other sites. Predation encounters here, encourage the use of predator vigilance rather than influencing the use of space within the habitat. The high density of Acacia karoo, which accounted for a third of annual foraging effort in what was a relatively depauperate floristic habitat, allows for an adequate energy intake for groups of this size. I ascribed the large group size and home range overlap to the inability of groups to undergo fission. / xiii, 79 leaves : ill., maps ; 29 cm

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