This study documented effects of fragmentation and disturbance on a coastal eucalypt dry open-forest plant community at Port Stephens-Myall Lakes, south-eastern Australia. The study evaluated the relative influence of fragment size (range &lt1-75ha), fragment or edge age (&lt 1-25y), time since fire (5-25y) and anthropogenic disturbance (minor, major) on microclimate, native plant species richness and weed invasion. Plots were sampled for native and exotic species richness per 25 m2 and edge transects for light, soil moisture, soil temperature, vegetation structure, native species richness and exotic species richness and cover. Depths of edge influence (DEI) were all &lt 20m inside the forest edge. Younger edge zones were lighter, had hotter and drier soils and more native species than forest interiors. Older edge zones were shadier, had warmer and moister soils and fewer native species than forest interiors, due to vegetation thickening in the edge zone. Light and soil moisture followed linear or monotonic edge-to-interior gradients in younger edges, but more complex patterns in older edges. Soil-temperature DEI decreased with increasing edge age. Fragment size had little influence on edge effects, but those for light developed more rapidly in smaller fragments, and recent fire was associated with reduced richness in edge zones of smaller fragments. Both anthropogenic disturbance and fire enlarged DEI for native species richness. Major anthropogenic disturbance coupled with fragmentation produced a stronger and more immediate loss of native species than fragmentation alone. Small fragments with minor disturbance had fewer native species per 25 m2 than larger fragments, but only after &gt10y since fragmentation. Analysis of forest interior portions of transects revealed that edge effects, and possibly disturbance, were largely responsible for this loss of native species with time, rather than effects of area. Despite a viable soil seed bank in fragments and the surrounding matrix, weed invasion in fragments was minimal. Exotic species were concentrated in edge zones, and were promoted by major anthropogenic disturbance within fragments. Results were highly dependent on fragment or edge age, and external influences of fragmentation (edge effects and disturbance), were more important than biogeographic (area-driven) factors. Impacts of fragmentation were compounded when combined with disturbance.

This thesis reports the results obtained, and conclusions made regarding research into the hydrochemistry of a small impounded karst area in southeastern Australia. Water samples were collected from a variety of sites over a period of approximately six years and include samples that are representative of baseflow (drought conditions) and flood (high flow conditions) the two extremes of the flow regime. Four distinct water types were found to be associated with the karst spring and cave water sites sampled. In terms of physical, chemical and computed parameter values these have been classed as TYPE 1 water (median parameter values similar to those noted for the surface stream sites sampled with catchments of non-carbonate material); TYPE 2 water (catchment predominantly of limestone); TYPE 3 water (catchment predominantly of dolomite); TYPE 4 water (high median chemical concentrations). Spatial variation within the study area is considerable and intrinsic factors such as catchment lithology, the residence time of recharge, aspect and vegetation cover are the major natural controls in determining physical and chemical characteristics.

Intertidal wetland habitats in southeastern Australia have changed significantly during the past sixty years. Mangrove habitats have expanded both seawards and landwards, the latter being at the expense of saltmarsh habitats. This relatively common phenomenon is generally suggested to be an outcome of sea-level rise. Several factors potentially responsible for this change are examined, including changes in mean sealevel during the past 50 to 100 years, changes in climate, population growth, catchment landuse, and estuary type. A protocol for mapping estuarine habitats was developed and implemented, incorporating the application of geographic information systems. Spatial and temporal coastal wetland habitat changes at nine sites along the New South Wales coast are illustrated. These habitat dynamics were shown to not correlate between sites. The results demonstrate that sea-level rise in this region cannot solely account for the extent of change during the past sixty years. With the exception of one site (Careel Bay), there have been no correlations between contemporary mean sea-level rise and mangrove incursion of the saltmarsh habitats at the study sites, or with rainfall patterns, at the scale of observation in this study, which was largely decadal. The only correlations determined during this study have been between population growth and coastal wetland habitat dynamics in some sites. In spite of saltmarsh habitat loss being a regional phenomenon, local factors appear to have a profound bearing on the rates of change. Neither contemporary mean sea-level rise, rainfall patterns, estuary type, catchment landuse, catchment natural cover nor population pressure can account solely for the patterns in the spatial and temporal dynamics of the coastal wetlands of New South Wales. It seems apparent that regional factors create preconditions favourable for mangrove incursion, but that localised conditions have been responsible for the extent of these incursions from site to site. That is, despite higher sea-level and greater rainfall, the extent of change has been determined by the unique characteristics of each site. The results have important implications for current estuary management practices in the state of New South Wales. The lack of spatial and temporal trends in coastal wetland habitat dynamics point to the need for management to be conducted on a localised, rather than regional scale. Additionally, anthropogenic influences must be carefully managed, since the extent of mangrove habitat expansion into saltmarsh areas is unlikely to be a natural occurrence.

Fauna conservation outside protected areas can make an important complementary contribution to conservation within reserves. This thesis aimed to contribute new information and analytical frameworks to the science of fauna conservation in human-modified landscapes. Two approaches were used: (1) empirical data collection and analysis, and (2) the discussion and development of conceptual landscape models. ¶ Empirical work focused on lizard distribution patterns in two production landscapes in southeastern Australia. Lizards were targeted because ectotherms are frequently neglected by conservation biologists. The “Nanangroe grazing landscape” was used for sheep and cattle grazing. In this landscape, approximately 85% of pre-European woodland cover had been cleared, and understorey vegetation was sparse. Lizards were surveyed at 16 landscape units, which were stratified by aspect, topographic position and amount of tree cover. Each landscape unit contained three sites, and each site contained three plots. Regression modelling showed that different species responded differently to their environment. For example, the four-fingered skink (Carlia tetradactyla) and Boulenger’s skink (Morethia boulengeri) were more likely to occur at woodland sites with northerly aspects, whereas the striped skink (Ctenotus robustus) and olive legless lizard (Delma inornata) were more likely to inhabit sites with a simple microhabitat structure. Statistical analysis further showed that the habitat attributes that lizards were related to varied continuously through space, and over different spatial scales. For example, invertebrate abundance (a proxy for food availability) varied most strongly over tens of metres, whereas the amount of grass cover varied most strongly over hundreds to thousands of metres. Thus, work at Nanangroe revealed spatially complex patterns of lizard occurrence and habitat variables. ¶ The “Tumut plantation landscape” was a spatial mosaic of native eucalypt (Eucalyptus) forest patches embedded within a plantation of the introduced radiata pine (Pinus radiata). In this landscape, thirty sites were surveyed for lizards. Sites were stratified by forest type and patch size, and included eucalypt patches, pine sites, and extensive areas of eucalypt forest adjacent to the plantation. Regression modelling showed that lizard species responded to various habitat attributes, including elevation, the amount of eucalypt forest within 1 km of a site, invertebrate abundance and ground cover. Variables related to habitat fragmentation often were significant predictors of lizard occurrence. However, work at Tumut suggested that important additional insights into lizard distribution patterns could be obtained by considering variables related to food and shelter resources, and climatic conditions. ¶ The Nanangroe and Tumut landscapes were in close proximity, but together spanned an altitudinal gradient of 900 m. An investigation of changes in lizard community composition with altitude showed that (1) only one species was common to Nanangroe and Tumut, (2) different species had different altitudinal preferences, and (3) ecologically similar species replaced one another with increasing altitude. These results highlighted that even in highly modified landscapes, natural gradients (such as climate) can play an important role in shaping animal assemblage composition and species distribution patterns. ¶ Empirical work suggested that, in some landscapes, the frequently used “fragmentation model” is a relatively weak conceptual basis for the study of animal distribution patterns. The fragmentation model implicitly assumes that “habitat patches” can be defined unequivocally across many species, and that patches are located within a relatively inhospitable matrix. Where these assumptions are breached, conservation guidelines arising from the fragmentation model may be too simplified. In spatially complex production landscapes, it may be more appropriate to maintain habitat heterogeneity at multiple spatial scales than to focus solely on the management of large, pre-defined patches. ¶ Given the potential limitations of the fragmentation model, a new, more holistic landscape model was developed. The “continuum model” was derived from continuum theory as developed for plant ecology. The continuum model recognises (1) spatial continua of environmental variables, and (2) species’ individualistic responses to these variables. For animals, key environmental variables may be related to the availability of food, shelter, sufficient space, and suitable climatic conditions. Unlike the fragmentation model, the continuum model is inherently process-based and thus may help to link the perceived gap between patterns and processes in landscape ecology. ¶ Three general conclusions arise from this thesis: 1. Some heterogeneous production landscapes support many native species, and therefore represent important conservation opportunities. 2. In some modified landscapes, the fragmentation model does not capture the complexity of animal distribution patterns. In those landscapes, conservation recommendations derived from the fragmentation model may be overly simplistic. 3. The continuum model may be a useful extension of the fragmentation model. It provides a process-based conceptual basis for empirical work on animal distribution patterns.