Based upon literature search, field investigation, examination of the halophyte collection at the Environmental Research Laboratory in Tucson, Arizona, and an expert survey, 97 salt-tolerant plants have been identified for the use as landscape plants. The plants were screened at salinity levels between 5 and 40 ppt which are critical for plant survival. The globally gathered data revealed information about landscape criteria, such as aesthetic value, climate amelioration, soil stabilization, color, form and drought tolerance, which were organized into architectural, engineering and design categories. These landscape halophytes range from herbaceous ground covers to woody trees. The data were filed in a computer data base, including detailed information about the description and performance of every plant, habitat requirements, maintenance, landscape values and economic uses. The data base, which is continually up-dated, is a search-type, which functions as an information storage system and is designed to provide plant selection by needed characteristics or intended use.

Graduation date: 1992

Thesis (Ph. D.)--University of Adelaide, 1987. / Includes bibliographical references (leaves 146-152).

Thesis (M. Sc. Agric(Pasture science))--University of Pretoria, 2005. / Includes bibliographical references. Available on the Internet via the World Wide Web.

High structural diversity amongst plant species in the arid winter rainfall region of southern Africa is common to other arid regions of the world. Details of the range and combinations of structural attributes in species, and within plant communities, are not widely known for southern African arid ecosystems. Hence little is known of how plant-form distributions vary within and between arid ecosystems. and of the environmental factors that may be responsible for any variations found. Nor is much information available on how phenological behaviour relates to the structural attributes of plant species. This study examined structural attributes of perennial plant species on the Worcester Veld Reserve, southwestern Cape. in relation to their distribution and phenology. Species were grouped on the basis of above ground structural and anatomical criteria. The distribution of species and plant-forms was assessed through a phytosociological survey along an environmental gradient in the area, and their phenology determined through qualitative and quantitative monitoring of species phenophases over a two year period. The results show that species of the predefined structural groups i) Co-occur throughout the area. but relative abundances vary from site to site and in relation to topography. aspect and the presence of Mirna-like mounds; ii) Phenological patterns for the flora overall are strongly seasonal, but the timing and periods of phenophases differ between, but are relatively uniform within, the defined structural plant groups. Since phenological patterns are indicative of resource-use patterns. these results indicate that the different plant-forms have different functional responses to the conditions of limited water availability and summer drought. This conceivably facilitates the co-existence of species in this water-limited environment. Habitat variability. which can be related to land-form patterns. is also implicated as a factor facilitating the co-existence of a diversity of species and plant-forms. Structural functional relationships known for arid region plants help to explain the relative uniformity of behaviour within plantform groups. and aid in understanding the ecological significance of distribution patterns of plant-forms in the arid landscape. The conclusion is reached that because of the close coupling between photosynthesis and water-use. the water storage potential and photosynthetic organ type are plant attributes which can serve as useful criteria by which arid region species can be grouped into ecologically meaningful categories or functional guilds. The formulation of a practical and meaningful functional classification is necessary to facilitate the understanding of complex vegetation patterns and processes within arid ecosystems. and allow for meaningful inter-ecosystem comparisons.

Northern Africa has experienced major shifts towards aridity and extensive C4 vegetation over the late Cenozoic, but due to a scarcity of spatially and temporally extensive paleoenvironmental records, the timing, patterns, and causes of these shifts are still under debate. Both long-term aridification and large amplitude orbital-scale climate variability have been recognized, with little understanding of how these two patterns relate to each other over time. African’s climate and environmental history of the last 7 Myr is of particular interest because hydrological and vegetation variability is considered the driving selection mechanism for human evolution. In addition, the age of the initiation of desert conditions in the modern Sahara desert, Earth’s largest warm desert and the largest source of dust to the modern atmosphere, is unknown. The stable isotope ratios of carbon and hydrogen in sedimentary plant leaf wax biomarker compounds have recently been shown to quantitatively track source vegetation photosynthetic pathways and the hydrogen isotope composition of plant source water, which is dominantly controlled by the amount of precipitation in Africa. These proxies have been applied to reconstruct long-term vegetation changes in East Africa and SW Africa over the last 14 Ma, as well as orbital-scale variability from various locations around the African continent, but they have not been extended further back in time or combined in tandem to robustly assess both long-term and orbital-scale climate and vegetation variability and how they relate to each other. In this thesis, I have utilized quantitative plant leaf wax stable isotope proxies to examine both orbital-scale and long-term changes in North African aridity and vegetation from a variety of regions over the last 25 Ma, with particular emphasis on the last 4.5 Ma. In Chapter 2, I investigated the evolution of hydrological and vegetation gradients from the equator to the sub-Sahara in NW Africa over the last 25 Myr using leaf wax stable isotopes at two marine sediment core locations, producing the longest existing leaf wax stable isotope record in Africa to my knowledge, and one of the longest such records globally. In this study I found that NW African environments were remarkably similar at both latitudes from 25 – 10 Ma, but at 10 Ma C4 vegetation abruptly expanded in the north, indicating sudden aridification in the Sahara region at that time. The hydrogen isotope record was stable long-term, with variability similar to that of known orbital-scale cyclicity in the Pliocene and Pleistocene, possibly suggesting that orbital-scale cyclicity or other factors obscured or were larger than any long-term changes in the hydrogen isotope ratio of precipitation. Saharan aridification at 10 Ma is consistent with climate model predictions of aridity due to the closure of the Tethys Seaway connection between the Indian Ocean and Mediterranean Sea near that time. The 10 Ma expansion in C4 vegetation is earlier than most other regions globally. To examine long-term changes in orbital-scale variability in the Eastern Sahara and Mediterranean Sea, I constructed a record of eastern Mediterranean sedimentary leaf wax carbon and hydrogen isotopes, leaf wax abundance, lignin biomarkers, and oxygen isotope ratios of planktonic foraminifera G. ruber during two 100-kyr periods of equal eccentricity near 3.0 and 1.7 Ma (Chapter 3). I found that precession-scale variability dominates the record during both periods, and Eastern Saharan precipitation and the vegetation assemblage, which was C4-dominated, do not change on average between the two periods. Chapter 4 extended the eastern Mediterranean record of Chapter 3 by sampling leaf wax stable isotopes in sapropel sediments (deposited during North African humid periods) at ~0.25 Myr resolution back to 4.5 Ma, placing the orbital-scale Chapter 3 results in long term context. I found that Eastern Saharan environments were persistently C4-dominated (>68%) throughout the entire interval, and that long-term hydrogen and carbon variability were similar in magnitude to orbital-scale cycles back to 4.5 Ma, strongly indicating that orbital-scale variability has been the dominant environmental control in NE Africa since the early Pliocene. This record contrasts sharply with observations of a transition from C3-C4 mixed vegetation to abundant C4 grasslands in East Africa over the same period of time. The results may suggest that long-term precipitation shifts did not occur in NE Africa since the Pliocene, or that the resolution of this approach is not sufficient to detect long-term shifts. It is likely that NW Africa also experienced similarly large hydrological variability over the same period of time, which may explain the unclear long-term hydrological signal in Chapter 2. The results emphasize that East Africa has not been representative of northern Africa as a whole since the Pliocene.

The world we live in is changing rapidly. Ecological, economic and social aspects and understandings are all undergoing paradigm shifts. Communities, farmers and individuals in arid zones are experiencing climate changes, more so than city dwellers. A better understanding of the current thinking in range ecology and management, especially of arid environments, is critical to the management of these delicate, complex systems. Ecosystems in equilibrium or in disequilibrium react differently to management and reclamation efforts. An understanding of the basic principles and how they evolved is important in order to apply these principles correctly in the management of arid zones. The use of keystone species and simple technologies, such as water harvesting and mulching, can all be used to reclaim and manage the arid zones. Understanding the differences between systems in equilibrium and disequilibrium can be used as a guide for planning appropriate future research in the arid zones. Searching for new indigenous species to help in the reclamation of arid zones is of the utmost importance. An ecological criterion was used to identify potential plant species for reclamation of degraded arid rangelands of southern Africa. Tripteris sinuatum and Sutherlandia microphylla were identified as possible candidates. Germination studies, with seeds harvested from naturally occurring plants, were conducted for both species. Treatments were based on the natural seed dispersal mechanisms for both species. S. microphylla has the potential to become an important plant species for reclamation purposes in arid zones of southern Africa. Not only potential new species should be sought, but also the management of species, already in use, are of critical importance. The relative palatability and survival of 16 different Atriplex species and accessions were determined at two different localities in the arid Northern Cape Province of South Africa. Significant differences were found between species at both localities. It is believed that because of the variety of species in the Atriplex genus, relative palatability and survival should be used to determine which of the different species could be useful under specific climatic and soil conditions. The establishment and reaction of Atriplex nummularia and Cassia sturti were tested for season of planting and the use of a stone mulch at two locations in the Northern Cape Province of South Africa. A. nummularia reacted the best to a moderate pruning treatments, while Cassia sturtii reacted best to severe pruning. These results should aid in the management of planted fodder plantations. Numerous attempts at improving natural veld have failed in the past. An examination of landscape function and the potential to harvest water in localized areas should drive veld improvement in arid zones. Seeds from two different plants species (Tetragonia calycina and Tripteris sinuatum) were used to inter-seed a bare patch in the Northern Cape Province of South Africa. The two species, with two treatments, (brush packing or not) in two different locally occurring eco-topes, were used to determine plant establishment. The water run-on eco-tope showed a significantly higher plant establishment percentage than the water run-off eco-tope. The establishment of perennial grass species was also found on the water run on eco-tope, three years after establishing the reclamation site. Such sites could form an important link in biodiversity conservation. / Dissertation (MSc Agric (Pasture Science))--University of Pretoria, 2007. / Plant Production and Soil Science / unrestricted

Thesis (MSc)--University of Stellenbosch, 2002. / ENGLISH ABSTRACT: Arid lands comprise some 47.2 % of the world's land surface. Between 32 % and 35 % of these drylands are subject to some form of desertification and land degradation. South Africa is an arid country with water being a major limiting resource. The Nama-Karoo biome is an extensive tract of semi-arid vegetation comprising some 22.7 % of South Africa, characterised by low rainfall and high temperatures. Grazing is the most common form of land use in the country, and particularly in this biome. Aridity and grazing are two factors that make large areas of South Africa (including the Nama-karoo) susceptible to land degradation. The Nama-Karoo biome has been neglected in past research and is an understudied part of South Africa's vegetation. This study is a component of a bigger umbrella project entitled "Restoration of degraded Nama-karoo rangelands: the role of conservation islands". The aim of this project was to assess the role played by isolated hills and mountains (mesas, inselbergs, etc.) in conserving the remnant biological diversity of the Nama-karoo rangelands of Namibia and South Africa. Vegetation composition, seed banks, seedling recruitment, pollination and grazing patterns and intensity on and off isolated mesas in the eastern parts of the Nama-karoo were investigated in the South African component of this project. I report on patterns of 1) plant species diversity and 2) plant communities across the Middelburg District, Eastern Cape, South Africa. Three isolated mesas (Tafelberg, Folminkskop and Buffelskop) and their surrounding plains were selected for this study. Permanent plots were established in broad transects extending from the plains to southeastern slopes, plateaux, north-western slopes and north-western plains of each mesa. Plots were 25 m2 in size, and ten 1 m2 blocks within the 25 m2 plot were randomly selected and sampled. Species composition was recorded and cover values determined for all plants in every subplot. In the eastern Nama-karoo mesas were not found to be higher in Alpha Diversity than their surrounding plains. Plains habitats were mostly equally as diverse as mesa habitats, with some plains habitats being higher in diversity than mesa habitats. Greater differences between mesas and their surroundings were found in a parallel study in the northern parts of the Nama-karoo in Namibia, indicating that mesas are more distinct islands of diversity with an increase in aridity. In the Middelburg District, a greater presence of woody phanerophytes occurred on mesa habitats in comparison with plains habitats, a possible result of the more moist microhabitats on mesas. Cooler, moister conditions on the south-eastern slopes of the mesas led to higher Alpha Diversity there in comparison with the warmer and drier north-western slopes, a phenomenon commonly found in arid areas. In terms of Alpha Diversity, the largest mesa sampled, Tafelberg (450 m above surroundings), was not significantly more diverse than the two smaller mesas (Folminkskop and Buffelskop both being 200 m above surroundings). Tafelberg and Buffelskop were equally high in Beta Diversity, with Folminkskop being much less diverse. Increased Beta Diversity for Tafelberg was explained by the increased size of the mesa (providing a more diverse microhabitat) relative to Folminkskop, which has the same shape and geology but is much smaller in size. High Beta Diversity values for Buffelskop were explained by the presence of degraded communities on the northwestern slopes and plains, while the south-eastern slopes were some of the most diverse habitats sampled in the landscape. Mesas and plains shared few species. Proportionally fewer species were shared between the plateau and slopes of the larger Tafelberg in comparison with the smaller two mesas. Isolation of the plateau could possibly become more distinct with an increase in size of the mesa. However, Tafelberg had more species in common with the surrounding plains than did the smaller mesas. It is suggested that bigger mesas support a more diverse array of microhabitats enabling plains species to occupy selective sites on the slopes of the mesa. Average Shannon-Wiener Alpha Diversity Indices for Middelburg were higher than that of the Nama-karoo in general. This could be explained by the higher precipitation in the eastern parts of the Nama-karoo relative to the rest. Diversity in Middelburg compared favourably with that in other arid lands of North America and Australia and was found to be similar in patterns and determinants of diversity compared to arid lands such as those of the New World. Regarding plant community composition, mesas were found to be distinctly different from their surrounding plains, with no shared communities between mesas and plains. The distribution of communities across the landscape was mainly attributed to a soilmoisture gradient. Mesas, compared to plains, have shallower, rockier soils coupled with very little run-off and naturally higher precipitation due to elevation. This results in a more mesic habitat. Plains, compared with mesas, have little rock cover, high percentages of bare soil and higher run-off rates than mesas, resulting in a more xeric habitat. Habitat differences such as these probably existed before the impact of domestic livestock but overgrazing has probably exacerbated the differences. The dominance structure of plant communities on the plains has probably changed in favour of toxic, spinescent or unpalatable plant species due to selective grazing by livestock. For the two dolerite-capped mesas (Tafelberg and Folminkskop) aspect and the expected cooler, moister conditions on south-eastern slopes as factors determining community composition were overridden by soil type and associated nutrient status. On Buffelskop (sandstone mesa), aspect and slope overrode soil type and associated nutrient status as determinants of community compositions. The potential to use mesas as a source of seeds and propagules to restore degraded plains habitats is low. Approximately 28 % of species were shared between mesas and their surroundings, and not all of these species could be regarded as good colonisers. Generalist, palatable species occurring across the landscape, such as Eragrostis obtusa, Felicia muricata, F. ovata, Fingerhuthia africana, Jamesbrittenia tysonii, Limeum aethiopicum, Pentzia incana and Selago albida could have some potential for future restoration attempts. Mesas are distinct in composition and can be regarded as islands of one vegetation type (mesas) in a sea of a different vegetation type (plains). Mesa habitats are not generally used for grazing by livestock, due to their general inaccessibility, and are not threatened by development. Plant communities in the Middelburg area were very similar In composition to communities identified in other parts of the Nama-karoo. Results from this study suggest that the plains are at present more degraded than the mesa habitats, but it is impossible to conclude whether or not these rangelands have stabilised given current stocking rates and climatic conditions. The Braun-Blanquet classification system proved an effective method to describe plant communities in the semi-arid Nama-karoo. All the plots used during the duration of the study were marked by metal stakes and are therefore of a permanent nature, so that they can be resampled in future. This might possibly shed some light on questions related to resilience, stability and degradation of the karoo. Restoring rangeland in the future is an important option, however, few farmers would, in the short term, be able to afford costly restoration techniques. It is recommended that farmers inspect the condition of their veld on a regular basis, and adjust stocking rates accordingly. Veld should be rested on a regular basis, and assessment should precede the movement of stock to a camp. Most farmers plough denuded areas of veld to encourage restoration, and it is recommended that this practice be combined with reseeding, followed by rest during and shortly after germination of the seed. / AFRIKAANSE OPSOMMING: Ariede areas beslaan tans 47 % van die aarde se land oppervlak. Tussen 32 % en 35 % van hierdie areas is onderworpe aan een of ander vorm van land degredasie. Die Nama- Karoo bioom is 'n ekstensiewe semi-ariede area wat naastenby 47 % van Suid-Afrika beslaan. Die karoo word gekenmerk deur hoë temperature en lae reënval. Suid-Afrika is 'n ariede land en water is 'n beperkende faktor. Weiding is die algemeenste boerderypraktyk, en meer so in ariede areas soos bv. in die karoo. Die droë klimaat en hoë voorkoms van weiding as boerderypraktyk maak ekstensiewe areas (insluitende die karoo) van Suid-Afrika vatbaar vir land degredasie. Hierdie studie is deel van 'n breër projek genaamd: "Restorasie van gedegradeerde Nama-karoo veld: Die rol van bewaringseilande". Die doel van hierdie projek was om die rol te bepaal van ge-isoleerde koppies en berge (mesas, inselberge, ens.) in die bewaring van biologiese diversiteit in die Nama-karoo van Suid-Afrika en Namibië. Plantegroeisamestelling, saadbanke, saailing oorlewing, bestuiwing en weidingsintensiteitlpatrone op koppies en hulle omringende vlaktes in die oostelike dele van die Nama-karoo is ondersoek in die Suid-Afrikaanse komponent van die studie. Ek rapporteer oor patrone van 1) plant diversiteit en 2) plantegroeigemeenskappe in die Middelburg Distrik, Oos-Kaap, Suid-Afrika. Drie ge-isoleerde koppies (Tafelberg, Folminkskop en Buffelskop) en hulle omliggende vlaktes is geselekteeer vir die doel van die studie. Permanente persele is uitgelê in 'n breë lyn vanaf die suid-oostelike vlaktes na die suid-oostelike hang, oor die plato's, noord-westelike hange en noord-westelike vlaktes vir al drie koppies. Persele was 25 m2 in grootte, waarvan tien 1 m2 sub-persele geselekteer is binne die groter 25 m2 blok. Spesies samestelling en bedekkingswaardes is bepaal vir alle plante in al die sub-persele. Mesas was nie hoër in Alpha Diversiteit as hulle omringende vlaktes nie. Vlaktes was meestal net so hoog in diversiteit as die mesas, en somtyds selfs hoër. Groter verskille in diversiteit is gevind in 'n paralelle studie in die noordelike dele van die Nama-karoo in Namibië. Dit dui aan dat mesas moontlik meer definitiewe eilande van diversiteit is in droër gebiede. Meer houtagtige fanerofiete het voorgekom op die mesas in vergelyking met die omliggende vlaktes, 'n moontlike gevolg van 'n natter habitat op die mesas. Suid-oostelike hange was hoër in alfa-diversiteit in vergelyking met noord-westelike hange. Dit is toegeskryf aan die koeler, natter mikrohabitat van die suid-oostelike hange. Die groter mesa (Tafelberg - 450 m bo die vlakte), was nie meer divers as die twee kleiner mesas nie (Folminkskop en Buffelskop is sowat 200 m bo die vlakte). Tafelberg en Buffelskop was ewe hoog in Beta Diversiteit, terwyl Folminkskop heelwat laer was. Hoër Beta Diversiteit vir Tafelberg kan verklaar word deur die groter oppervlak en hoogte van die mesa (voorsien 'n meer diverse mikrohabitat) in vergelyking met Folminkskop. Folminkskop het dieselfde vorm en geologiese geskiedenis as Tafelberg, maar is veel kleiner. Buffelskop was hoog in Beta Diversiteit as gevolg van 'n gedegradeerde gemeenskap op die noord-westelike hang en vlaktes, terwyl die suidoostelike hang baie hoog was in diversiteit. Min spesies kom op beide mesas en vlaktes voor. Minder spesies is gedeel deur die plato en hange van Tafelberg in vergelyking met die plato's en hange van Folminkskop en Buffelskop. Isolasie van die plato kan moontlik hoër wees in groter mesas. Tafelberg (slegs mesa) het meer spesies in gemeen met sy omliggende vlakte as die ander twee mesas. Groter mesas soos Tafelberg kan moontlik 'n meer diverse spektrum van mikrohabitatte bevat, wat sekere vlakte spesies in staat stel om te oorleef op mesa hange. Gemiddelde alfa diversiteit vir die Middelburg Distrik was hoër as die van die res van die karoo. Dit kan verklaar word deur die hoër reënval in die oostelike dele van die karoo relatief tot die meer westelike dele. Diversiteit in Middelburg vergelyk goed met die van ander ariede lande in Noord-Amerika en Australie, terwyl patrone en faktore wat diversiteit bepaal soortgelyk was aan die van gemeenskappe in ariede lande van die Nuwe Wêreld. Mesas en vlaktes het drasties verskil in die samestelling van hulle onderskeie plantegroeigemeenskappe, alhoewel hulle soortgelyk was in diversiteit. Geen plantegroeigemeenskappe het voorgekom op beide mesas en vlaktes nie (gedeelde plantegroeigemeenskappe). Die verspreiding van plantegroeigemeenskappe oor die landskap was toegeskryf aan 'n grond-water gradient. Mesas het, oor die algemeen, vlakker, meer rotsagtige grond en 'n hoër reënval met minder afloop as vlaktes. Vlaktes het minder rotse, 'n hoër persentasies kaal grond en vinniger afloop tempo's na reën as mesas. Hierdie faktore het tot gevolg dat mesas 'n natter habitat verteenwoordig in vergelyking met die meer ariede vlaktes. Habitatsverskille soos dié het heel moontlik klaar bestaan voor die aankoms van vee, maar oorbeweiding het die verskille in habitat tussen mesas en vlaktes groter gemaak. Selektiewe beweiding deur vee het waarskynlik tot gevolg gehad dat plante wat giftig, doringagtig of onsmaaklik is, toegeneem het in plantegroeigemeenskappe, ten koste van meer smaaklike spesies. Grondtipe en grondsamestelling het aspek en verwagte koeler kondisies op die suidoostelike hange onderdruk as bepalende faktore vir plantegroeigemeenskap samestelling op Tafelberg en Folminkskop (beide bedek met 'n doleriet laag). Aspek en steilte van die hange het grondtipe en grondsamestelling onderdruk as bepalende faktore vir plantegroeigemeenskappe op Buffelskop. Die potensiaal om mesas te gebruik as bronne van saad vir die rehabilitasie van die vlaktes was laag. Mesas en hulle omliggende vlaktes het naastenby 28 % van spesies in gemeen gehad, maar nie al hierdie spesies was goeie koloniseerders nie. Sekere smaaklike plante wat op beide mesas en vlaktes voorgekom het, kan potensiaal hê vir toekomstige restorasie doeleindes, bv. Eragrostis obtusa, Felicia muricata, F. ovata, Fingerhuthia Africana, Jamesbrittenia tysonii, Limeum aethiopicum, Pentzia incana en Selago albida. Mesas is uniek in samestelling en kan beskou word as een plantegroeitipe in 'n see van 'n ander tipe (vlaktes). Mesas word nie oor die algemeen bewei nie (as gevolg van hulle steil hange en bergagtigheid) en word gevolglik nie bedreig deur huidige ontwikkeling nie. Plantegroeigemeenskappe in Middelburg is soortgelyk aan die van ander dele van die Nama-Karoo bioom. Resultate van die studie dui aan dat die vlaktes huidiglik meer gedegradeer is as mesas, maar dit is onbekend of die veld gestabiliseer het of steeds besig is om verder te degradeer in huidige weidings- en klimaatskondisies. Die Braun-Blanquet klassifikasiemetode IS effektief vir die beskrywing van plantegroeigemeenskappe in ariede areas. Al die persele in die studie area is gemerk met metaal paaltjies en is dus permanent. Dit maak dit moontlik om in die toekoms die persele weer te ondersoek. 'n Langtermyn datastel kan moontlik lig werp op die stabiliteit en degredasie van karoo veld. Restorasie van veld is 'n moontlikheid vir die toekoms, maar min boere kan duur restorasie metodes bekostig in die kort termyn. Boere moet hulle lande gereeld ondersoek en drakrag aanpas by die kondisie van die veld. Baie boere ploeg kaal kolle in die veld op 'n gereelde basis, en daar word voorgestel dat die praktyk gekombineer word met die saai van geskikte saad, gevolg deur 'n rusperiode tydens en na ontkieming daarvan.

Bibliography: leaves 146-152.

Bibliography: leaves 146-152 / 152 leaves : ill ; 30 cm. / Title page, contents and abstract only. The complete thesis in print form is available from the University Library. / Thesis (Ph.D.)--Dept. of Plant Pathology, Waite Agricultural Research Institute, University of Adelaide, 1987