<p> This dissertation examines the competitive and cooperative strategies of male mantled howlers (<i>Alouatta palliata</i>) living within a fragmented habitat at La Pacifica (LP), Costa Rica. Mantled howlers exhibit considerable intraspecific variation in their social behavior, and there remains much to learn regarding the nature of their sociality and the proximate mechanisms that underlie it, particularly among males living in multimale groups. In contrast to males at other mantled howler study sites, males at LP had higher rates of within-group agonism, and exhibited clear dominance hierarchies. Both groups inhabited a fragmented forest, which may exert greater pressure to compete for both food and reproductive resources than mantled howler males living in more intact forests. However, data from this dissertation also suggest that males may adjust competitive relationships as forest fragment characteristics within the same population vary. The group inhabiting the larger forest fragment experienced higher intergroup competition but lower intragroup competition than the group inhabiting the smaller fragment, allowing higher-ranking males to be more tolerant of subordinates accessing key resources, including females, in order to cooperate in group defense. Despite clear hierarchies, males at LP nevertheless form both preferred and avoided associations with specific males, which suggest that cooperative relationships among males have adaptive value. Neither rank nor age emerged as clear drivers shaping competitive and cooperative relationships. Collectively, these results suggest that male mantled howlers show considerable flexibility in their social relationships that are likely dependent on a complexity of ecological, social, and demographic factors.</p>

<p> During the 1970s to the 1990s, Steller sea lions (<i>Eumetopias jubatus</i>) in Alaska declined severely, resulting in the western stock's listing as endangered under the United States Endangered Species Act. The localized depletion of their main food source around rookeries and haul out sites was the primary cause of their depletion. The western stock has still not recovered to historical levels. Therefore, scientists have continued to study Steller sea lions diet and foraging habits. One approach has been through stable isotope analysis of their various tissues. Analysis of their vibrissae, or whiskers, provides an exact record of an individual's diet from birth to the present. However, scientists so far have only measured stable isotope values of a single vibrissa from an individual, assuming no significant difference among vibrissae within or between cheeks. They have not accounted for the possibility of variable stable isotope accumulation in an individual's different vibrissae. In this study, I tested the assumption of no significant individual variability among vibrissae by examining the &delta;¹³C and &delta;¹⁵N values in all of the vibrissae on five Steller sea lion pups recovered in Alaska. &delta;¹⁵N values did not differ between vibrissae on a cheek or between vibrissae from left and right cheeks. The &delta;¹³C values did not differ between vibrissae within cheeks; however, &delta;¹³C values were significantly different between the left and right cheeks of Lowrie Island Pup and PWS 29 Pup. This between cheek variation is most likely due to small sample size rather than different stable isotope incorporation. The lack of &delta;¹⁵N value variation within or between cheeks, as well as the lack of &delta;¹³C variation within cheeks, supports the assumption that stable isotope accumulation is similar between all vibrissae and validates sampling protocols of previous studies collecting just the longest, thick vibrissa from a Steller sea lion. More studies similar to this one are needed to verify these findings and to look at variability in stable isotope incorporation of juvenile and adult Steller sea lion vibrissae as well as other mammal species.</p>