<p>Ecologists have long been interested in the relationships between climate change and forest biodiversity. For centuries, the scientific problems remain understanding the patterns of climate variation, forest geographic distribution, and demographic dynamics. Besides scientific merits, these questions will also help forest managers and policy makers to anticipate how forests respond to global change. This dissertation tackles these problems by using statistical modeling on climate and forest inventory data in the eastern United States.</p><p>In Chapter 1, we ask the question on the observed tree range distributions in response to contemporary climate change in the eastern United States. Tree species are expected to track warming climate by shifting their ranges to higher latitudes or elevations, but current evidence of latitudinal range shifts for suites of species is largely indirect. In response to global warming, offspring of trees are predicted to have ranges extend beyond adults at leading edges and the opposite relationship at trailing edges. Large-scale forest inventory data provides an opportunity to compare present latitudes of seedlings and adult trees at their range limits. Using the USDA Forest Service's Forest Inventory and Analysis data, we directly compared seedling and tree 5th and 95th percentile latitudes for 92 species in 30 longitudinal bands for 43,334 plots across the eastern United States. We further compared these latitudes with 20th century temperature and precipitation change and functional traits, including seed size and seed spread rate. Results suggest that 58.7% of the tree species examined show the pattern expected for a population undergoing range contraction, rather than expansion, at both northern and southern boundaries. Fewer species show a pattern consistent with a northward shift (20.7%) and fewer still with a southward shift (16.3%). Only 4.3% are consistent with expansion at both range limits. When compared with the 20th century climate changes that have occurred at the range boundaries themselves, there is no consistent evidence that population spread is greatest in areas where climate has changed most; nor are patterns related to seed size or dispersal characteristics. The fact that the majority of seedling extreme latitudes are less than those for adult trees may emphasize the lack of evidence for climate-mediated migration, and should increase concerns for the risks posed by climate change.</p><p>In Chapter 2, we ask the question on tree abundance within geographic range responding to climate variation in the eastern United States. Tree species are predicted to track future climate by shifting their geographic distributions, but climate-mediated migrations are not apparent in a recent continental-scale analysis (Chapter 1). To better understand the mechanisms of a possible migration lag, we analyzed relative recruitment patterns by comparing juvenile and adult tree abundances in climate space. One would expect relative recruitment to be higher in cold and dry climates as a result of tree migration with juveniles located further poleward than adults. Alternatively, relative recruitment could be higher in warm and wet climates as a result of higher tree population turnover with increased temperature and precipitation. Using the USDA Forest Service's Forest Inventory and Analysis data at regional scales, we jointly modeled juvenile and adult abundance distributions for 65 tree species in climate space of the eastern United States. We directly compared the optimal climate conditions for juveniles and adults, identified the climates where each species has high relative recruitment, and synthesized relative recruitment patterns across species. Results suggest that for 77% and 83% of the tree species, juveniles have higher optimal temperature and optimal precipitation, respectively, than adults. Across species, the relative recruitment pattern is dominated by relatively more abundant juveniles than adults in warm and wet climates. These different abundance-climate responses through life history are consistent with faster population turnover and inconsistent with the geographic trend of large-scale tree migration. Taken together, this juvenile-adult analysis suggests that tree species might respond to climate change by having faster turnover as dynamics accelerate with longer growing seasons and higher temperatures, before there is evidence of poleward migration at biogeographic scales.</p><p>In Chapter 3, we ask the question on the demographic dynamics of density dependence at the individual tree level in eastern US forests. Density dependence could maintain diversity in forests, but studies disagree on its importance. Part of the disagreement results from the fact that different studies evaluate different responses (per-seedling or per-adult survival or growth) of different stages (seeds, seedlings, or adults) to different inputs (density of seedlings, density or distance to adults). Most studies are conducted on a single site and thus are difficult to generalize. Using USDA Forest Service's Forest Inventory and Analysis data, we analyzed over a million seedling-to-sapling recruitment observations of 50 species for both per-tree (adult) and per-seedling recruitment rates, controlling for climate effects in eastern US forests. We focused on per-tree recruitment as it is most likely to promote diversity at the population level, and it is most likely to be identified in observational or experimental data. To understand the prevalence of density dependence, we quantified the proportion of species with significant positive or negative effects. To understand the strength of density dependence, we determined the magnitude of effects among conspecifics and heterospecifics, and how it changes with overall species abundance. We found that the majority of the 50 species have significant density dependence effects, mostly negative, on both per-tree and per-seedling recruitment. Per-tree recruitment is positively associated with conspecific seedlings, saplings, and heterospecific saplings, negatively associated with heterospecific seedlings, conspecific and heterospecific trees. Per-seedling recruitment is positively associated with conspecific and heterospecific saplings, but negatively associated with conspecific and heterospecific seedlings and trees. Furthermore, for both per-tree and per-seedling recruitment, density dependence effects are stronger for conspecific than heterospecific neighbors. However, the strength of these effects does not vary with species abundance. We conclude that density dependence is pervasive, especially for per-tree recruitment, and its strength among conspecifics and heterospecifics is consistent with the predictions of the Janzen-Connell hypothesis.</p> / Dissertation

A monthly resolved, 133 year record of coral Sr/Ca variations has been developed from a massive Porites coral that was drilled in the shallow waters of a submerged carbonate platform (Sabine Bank, 15.9°S, 166.14°E) located ~50 km west of Espiritu Santo, Vanuatu. This truly open-ocean site, at which daily measurements of temperature and salinity are available for ~ 6 years, permits the reconstruction of local environmental variability using variations in coral skeletal geochemistry. Coral Sr/Ca-SST variations are well matched to variations in local SST, but bear little relation to changes in local SSS indicating little or no influence of salinity on coral Sr/Ca. The complete coral Sr/Ca-SST time series is characterized by abundant inter-annual variability, a strong trend towards warming (i.e., lower Sr/Ca values) from ~1980-2006. Interannual SSTA variations at Sabine Bank correspond reasonably well to SSTA variations from the central Pacific cool tongue (Niño 3.4 region), indicating that coral Sr/Ca variations record ENSO variability in the region. / text

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Two of the most pressing questions involving ancient bristlecone pines are how microsite factors lead to differences in tree responses to climate at high-elevation sites, and how global change has impacted growing season events. Disparities in climate response at treeline and subalpine locations have been associated with local environmental characteristics while the increasing growth trend found at treeline has been linked to warming. In this study, environmental conditions were considered in order to identify microsite differences between trees growing at two different elevations on four aspects of a conical-shaped mountain in the White Mountains, California. Dendrochronological, environmental, correlational, and spectral methods were employed to explore differences in ring-width chronologies. Albedo, soil thickness, and percent slope led to ring-width variability. Northwestern upper site was most highly correlated with precipitation, while the Southeastern lower site showed a strong negative correlation with temperature. This work indicates that selection of climate-sensitive trees a priori necessitates the consideration of local environmental factors, and these microsite differences resulted in different climate responses between nearby trees. By monitoring growing season events at the historic phenology site from the 1962-64 (Fritts 1969), natural variations and responses to climate change can be identified. Morphological and physiological phenophases, dendrometer traces, and environmental data were collected throughout the summers of 2007 and 2008. Duration and timing of cambial activity (tracheid lifespan) in the present study were similar to those recorded in the Fritts (1969) investigation, while pollination onset and bud opening occurred earlier in this study. No change was found in duration and timing of cambial activity suggesting that changes in cambial phenology are not an explanation for the increasing growth trend found at upper forest borders. On the other hand, changes in bud opening and pollination onset may be related to recent warming. To monitor diurnal and seasonal stem variability as part of phenologic studies on several trees, a point potentiometer dendrometer was designed. The newly designed point potentiometer dendrometer was tested in multiple environments and found to be versatile, cost-effective, and portable, working well in semi-arid and arid environments.

This item was digitized as part of the Million Books Project led by Carnegie Mellon University and supported by grants from the National Science Foundation (NSF). Cornell University coordinated the participation of land-grant and agricultural libraries in providing historical agricultural information for the digitization project; the University of Arizona Libraries, the College of Agriculture and Life Sciences, and the Office of Arid Lands Studies collaborated in the selection and provision of material for the digitization project.

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