The late Neogene represents an exceptionally dynamic period in Earth history during which the Northern Hemisphere has transitioned from a warmer, more equable climate to a cooler, more transient state characterized by waxing and waning continental ice sheets. While geographical distal, the tropical ocean has played a significant role in shaping the evolution of the climate system, as the opening and closing of low latitude (LL) ocean gateways and reorganization of oceanic and atmospheric circulation structure have helped shape the climate system into its present form. This study provides a reconstruction of sea surface temperature (SST), ocean circulation, and thermal structure of the LL eastern Pacific and North Atlantic based upon the compilation of proxy data derived from planktic foraminifer assemblages and geochemical techniques. This research begins with a paleoceanographic reconstruction of the eastern tropical Pacific (ETP) and subtropical Northwest Atlantic (NWA) during the early stages of uplift of the Central American Isthmus and associated shoaling of the Central American Seaway (CAS). In the subtropical NWA (DSDP 103 and ODP 1006), the 5.2 to 5.1 Ma interval is characterized by an increase in SST and sea surface salinity, indicating a strengthening of the Florida Current (FC) and Gulf Stream (GS). Sea surface temperature in the ETP Warm Pool (DSDP Site 84) remained relatively stable between 6.9 and 5.1 Ma, during which El Niño-like conditions persisted. A slight cooling is observed after this interval (with synchronous warming in the NWA), followed by the onset of major cooling at ~3.2 Ma, both of which are preceded by a shallowing of the thermocline. Stepwise cooling is attributed to enhanced Atlantic meridional overturn circulation (AMOC), which caused a shoaling of the main tropical thermocline, thereby strengthening the Walker Circulation and weakening the Pacific North Equatorial Counter Current. During the mid-Piacenzian warm period (MPWP; ~3.3-3.0 Ma), SST in the Panama Basin was ~0.8°C cooler than today, while the subtropical NWA was only ~1.1°C warmer. This corroborates evidence for reduced meridional SST gradients during the mid-Pliocene as well as the hypothesis that more vigorous ocean circulation--particularly in the NWA--was critical during this period. The timing of SST changes in the ETP and NWA (~5.1 Ma) suggest that the termination of permanent El Niño and enhanced AMOC did not contribute significantly to the onset of major Northern Hemisphere glaciation (NHG), as both of these events occur well before the beginning of the glacial cycles. However, these processes may have contributed to the development of the small ice sheets of the late Miocene and early Pliocene, but were most likely only preconditioning factors for the onset of major NHG. In contrast, changes in SST and relative thermocline position suggest that high latitude (HL) processes and global cooling may have influenced thermal structure in the ETP. The SST estimates provided indicate that even in its early stages, the shoaling of the CAS had significant implications for low-latitude ocean circulation and thermal structure, as well as for some of the most significant global climate events of the late Neogene, including the MPWP. During the MPWP, mean global surface temperatures were similar to those predicted for the next century (2-3˚ C warmer) while atmospheric CO2 concentrations, paleogeography, and paleobiology were similar to today. As such, the MPWP has been studied in detail as a potential (albeit imperfect) analog for future climate change and has provided a natural and unique test-bed for the integration of proxy data and general circulation models. Central to this research effort is the Pliocene Research, Interpretation, and Synoptic Mapping (PRISM) project, an iterative paleoenvironmental reconstruction of the MP focused on increasing our understanding of warm-period climate forcings, dynamics, and feedbacks by providing three-dimensional data sets for general circulation models. A mainstay of the PRISM project has been the development of a global sea surface temperature (SST) data set based primarily upon quantitative analyses of planktic foraminifer assemblages, supplemented with geochemical SST estimates wherever possible. In order to improve spatial coverage of the PRISM faunal and SST data sets in the LL North Atlantic, this study provides a description of the MP planktic foraminifer assemblage and multiproxy SST estimates from five Ocean Drilling Program sites (951, 958, 1006, 1062, and 1063) in the North Atlantic subtropical gyre (NASG), a region critical to Atlantic Ocean circulation and tropical heat advection. Assemblages from each core provide evidence for a temperature- and circulation-driven 5-10° northward displacement of MP faunal provinces, as well as regional shifts in planktic foraminifer populations linked to species ecology and interactions. General biogeographic trends also indicate that, relative to modern conditions, gyre circulation was stronger (particularly the Gulf Stream, North Atlantic Current, and North Equatorial Current) and meridionally broader. Overall, SST estimates suggest that surface waters in this region were not significantly warmer (1-2˚ C) than today and that mean annual SSTs along LL western boundary currents were indistinguishable from modern. Multiproxy SST data also provide evidence for enhanced northward transport of warm, salty, oligotrophic surface waters via a vigorous western boundary current system with warmer (cooler) cold-season (warm-season) temperatures. Collectively, this reconstruction of SST and ocean circulation provides support for a model of an enhanced Atlantic meridional overturn circulation (AMOC) system, with particularly vigorous LL western boundary currents and thus, more efficient northward heat transport. These trends therefore suggest that more vigorous thermohaline circulation, in conjunction with elevated atmospheric CO2 concentrations, played a significant role in shaping the global surface temperature distribution during the MPWP. A strengthening of the AMOC under warmer-than-modern conditions has significant implications for future climate change. The current generation of climate models suggests that HL warming and associated ice-sheet melting will induce a freshening of the North Atlantic and thus, to a reduction in the strength of the AMOC, thereby buffering surface temperature increases in the Northern Hemisphere. However, if after this transient period of climate system adjustment, Earth returns to a more Pliocene-like climate state the AMOC system may strengthen, thereby exacerbating the HL warmth caused by elevated atmospheric CO2 concentrations. Thus, through the reconstruction of warm-period SST and ocean circulation, this research provides insight into the potential operation of the LL North Atlantic and its associated impact on broad-scale Northern Hemisphere climate.

The Neogene history of Caribbean deep-sea benthic foraminifera was investigated by calculating changes in paleoproductivity, diversity and paleobiogeography ~26 to 2 Ma, which includes the progressive closure of the Central American Seaway. Three hypotheses were tested: (1) Paleoproductivity values prior to closure of the Central American Seaway are similar in both the Caribbean and equatorial Pacific and then diverge by the time of early shoaling events; (2) Diversity values of benthic foraminifera prior to the closure of the Central American Seaway were similar in the Caribbean and EEP, and had changed by the time of early shoaling; and (3) during the Miocene and into the Pleistocene, the progressive constriction of the CAS affected deep-sea benthic foraminiferal assemblages by increasing their dissimilarity between the Caribbean and equatorial Eastern Pacific. These hypotheses were tested with 104 samples from five Caribbean and EEP deep-sea cores by calculating paleoproductivity with multiple proxies, determining diversity indices and calculating biogeographic similarity coefficients. The data supported the first two hypotheses: The greatest change in paleoproductivity occurred at ~8 Ma during seaway constriction, when values diverged between the Caribbean and EEP. After complete seaway closure at ~4 Ma, the Caribbean became oligotrophic, noted by a decrease in high-organic flux species, and an increase in Nuttalides umbonifera, an indicator species. The largest changes in species-level diversity occurred with the barrier to deep-water flow at ~12 Ma, and Caribbean diversity increased at ~8 Ma with seaway constriction. However, the third hypothesis was rejected: Increases in assemblage similarity actually occurred during most major paleoceanographic events, with the only decrease in Caribbean-EEP similarity occurring at ~12 Ma, coincident with a drop in diversity and emplacement of the Panama isthmian sill. Thus, the barrier to deep-water flow at ~12 Ma affected the composition of tropical American benthic foraminifera more than the largest change in paleoproductivity at ~8 Ma, or closure of the Central American Seaway at ~4 Ma.