Typescript. Includes bibliographical references (leaves [279]-287) This thesis examines the growth ring structure of Eucalyptus camaldulensis Dehnhardt and investigates links between ring features and the climatic conditions under which growth took place. Samples of E. camaldulensis from the Barmah Forest (near the River Murray in northern Victoria) were used in the study. E. camaldulensis growth is linked to periodic flooding, and the Barmah Forest contains sites that are frequently flooded. Wood samples were obtained from sites subject to different average flooding frequencies. Trees used in the study grew out of natural regeneration in the 1920's and 1930's and from regeneration trials in the early 1960's. Initial investigation of E. camaldulensis samples revealed ring-like features that were able to be traced on samples by eye. Microscopic investigation showed that there existed considerable variations in the properties of individual rings at different positions on the samples, and that the boundaries between rings were often indistinct. Further examination of E. camaldulensis microstructure was performed on samples from two trees that grew on sites with significantly different flooding regimes. These samples possessed features that formed rings, with rings being successfully matched between samples taken from different heights in the trees. As the complex microstructure of E. camaldulensis did not lend itself to standard dendroclimatological techniques, methods were developed to facilitate the comparison and matching of rings. These methods were also used in the successful matching of ring patterns with the output from a simple climate-based tree growth model. Two sets of E. camaldulensis samples (BS1, with 33 samples, and BS2, with 39 samples) were studied to assess the level of individual variability in ring patterns, and to provide a representative ring pattern for climate comparison. Following the development of methods for identifying samples with similar ring patterns, a subset of similar samples was selected from the BS1 set. A member of this subset was selected to provide a ring-width pattern upon which a representative pattern of ring features for BS1 was based. The rings of the BS2 samples possessed poorer ring definition than the BS1 samples and provided no new or different information. Consequently, the representative ring pattern for BS1 was used in a dendroclimatological investigation for the site. The ring pattern was matched with the output from two tree growth event models. Although rings were matched with growth events over a 27 year period, the high variability of individual ring features prevented matching of particular types of ring features with particular types of climatic events. An investigation of numerical methods for matching ring patterns with ring or growth event patterns, and for identifying samples with similar ring patterns, was performed using signal smoothing and filtering techniques and a dynamic time-warping procedure. Ring matching and identification of similar ring patterns was found to be most successful on samples where the ring patterns, expressed as continuous signals, had similar mean and amplitude values. The techniques were unsuccessful in the matching of signals of different form, such as continuous ring pattern signals and discontinuous growth event signals.

The purpose of this investigation was to evaluate the usefulness of white oak tree-ring sensitivity to local climatic information. White oak core samples were taken from Christy Woods in Muncie, Indiana. Tree-ring measurements were standardized and summarized using computer programs from the Laboratory of Tree-Ring Research, Tucson, Arizona.Low mean sensitivity in each series, and a variance component for the total group chronology of 30.23% indicated that annual growth increments of white oak were less than moderately limited by annual mean growing season precipitation and temperature. It may be inferred that a variety of factors have limited growth diminishing the affects of macroclimatic variations on growth at this site.By use of stepwise multiple regression analysis annual growth increment for the mean chronology was positively correlated to mean growing season precipitation. Annual growth increment was negatively correlated to mean growing season temperature. Only 24% of the total variability in the indices was attributable to the dependence on precipitation and temperature.Ball State UniversityMuncie, IN 47306

The first objective of this thesis was to predict the future success of selected tree species under low (B1, 550 CO2 ppm) and moderate (A1B, 720 CO2 ppm) climate change scenarios as defined in the Special Report on Emissions Scenarios (SRES). This was accomplished through the creation of radial-growth forecasts for eastern hemlock (Tsuga Canadensis (L.) Carr.), sugar maple (Acer saccharum L.), white spruce (Picea glauca (Moench.) Voss), and white pine (Pinus strobus L.) in the Grand River Watershed of Ontario, Canada. The forecasts were founded on historic growth-climate relationships between standardized regional dendrochronologies for each species and past climate data from the Guelph OAC weather station. These species-specific growth-climate relationships were then extended to 2100 using modeled climate data from the Third Generation Coupled Global Climate Model (CGCM3) to project radial-growth under both emissions scenarios. Results indicated that eastern hemlock radial-growth will remain stable throughout the 21st-century, sugar maple and white spruce growth will start to decline, and white pine growth will increase. While the radial-growth forecasts were limited by the length of the past climate data, the accuracy of the modeled climate data, and the number and type of variables used in the forecast model, the results were statically significant and strongly supported in the literature. The second thesis objective was to assess the potential impact of the radial-growth forecasts on environmental planning policy and forest management strategy in the Grand River Watershed. Examples of how the forecasts could influence basic management strategies in the watershed were provided to display the conceptual linkages between the results and policy formulation. Next, the radial-growth forecasts were presented to four forest managers working in the watershed to gage the practical implications, perceptions and limitations of the radial-growth forecasting method. While the managers found the radial-growth forecasts interesting, they also noted that the results were of limited use since they could not account for other factors important to the future success of the study species, such as seedling dispersal and establishment rates, as well as the potential effects of pathogens, insects and invasive species. Therefore, it was recommended that future research should work to extrapolate the results of the radial-growth forecasts to other tree species and types in the region, as well as incorporate more variables into the models, so that more accurate and applicable growth projections could be constructed in the watershed.

The first objective of this thesis was to predict the future success of selected tree species under low (B1, 550 CO2 ppm) and moderate (A1B, 720 CO2 ppm) climate change scenarios as defined in the Special Report on Emissions Scenarios (SRES). This was accomplished through the creation of radial-growth forecasts for eastern hemlock (Tsuga Canadensis (L.) Carr.), sugar maple (Acer saccharum L.), white spruce (Picea glauca (Moench.) Voss), and white pine (Pinus strobus L.) in the Grand River Watershed of Ontario, Canada. The forecasts were founded on historic growth-climate relationships between standardized regional dendrochronologies for each species and past climate data from the Guelph OAC weather station. These species-specific growth-climate relationships were then extended to 2100 using modeled climate data from the Third Generation Coupled Global Climate Model (CGCM3) to project radial-growth under both emissions scenarios. Results indicated that eastern hemlock radial-growth will remain stable throughout the 21st-century, sugar maple and white spruce growth will start to decline, and white pine growth will increase. While the radial-growth forecasts were limited by the length of the past climate data, the accuracy of the modeled climate data, and the number and type of variables used in the forecast model, the results were statically significant and strongly supported in the literature. The second thesis objective was to assess the potential impact of the radial-growth forecasts on environmental planning policy and forest management strategy in the Grand River Watershed. Examples of how the forecasts could influence basic management strategies in the watershed were provided to display the conceptual linkages between the results and policy formulation. Next, the radial-growth forecasts were presented to four forest managers working in the watershed to gage the practical implications, perceptions and limitations of the radial-growth forecasting method. While the managers found the radial-growth forecasts interesting, they also noted that the results were of limited use since they could not account for other factors important to the future success of the study species, such as seedling dispersal and establishment rates, as well as the potential effects of pathogens, insects and invasive species. Therefore, it was recommended that future research should work to extrapolate the results of the radial-growth forecasts to other tree species and types in the region, as well as incorporate more variables into the models, so that more accurate and applicable growth projections could be constructed in the watershed.

Prepared for The National Park Service, United States Department of Interior, Yellowstone, Wyoming / The goal of this investigation has been the reconstruction of past seasonal climate for the period 1750-1910 (161 years) in the Yellowstone National Park region based upon tree-ring data. Tree-ring series are useful in the reconstruction of past climate owing to the availability of large numbers of trees, the great longevity of trees, and the critical fact that the climatic information they contain is accumulated over specific years. In this project a number of tree-ring series from the region around Yellowstone National Park have been calibrated against short-term (1912-1971) seasonal temperature and precipitation data for Bozeman, Moran, Red Lodge, and Yellowstone Park. From these calibrations, long-term seasonal temperature and precipitation records have been reconstructed for each of the four stations. A major reason for these reconstructions has been the need for long-term climatic data that can be used to indicate potential variations in the climate of the park region. Knowledge of these climatic variations may facilitate estimates of natural food supplies or availability of forage in winter as related to snow depth. Previously such estimates have had to be based upon relatively short-term climatic data which undoubtedly do not encompass all possible climatic variations. With this in mind, a series of precipitation and temperature maps have been produced to indicate some of the seasonal extremes that have probably been experienced since 1750 within a given year or group of years as indicated by the tree -ring data. It is hoped that these maps will be useful to various types of researchers involved in planning within Yellowstone National Park.

Little research has been conducted on the sensitivity of earlywood vessel in ring-porous tree species in response to flooding. The impact of flooding and climate on vessel characteristics in black ash (Fraxinus nigra Marsh.) was studied in the boreal region of Lake Duparquet, northwestern Quebec. In addition to standard tree-ring measurements, numbers and cross-sectional area of earlywood vessels were examined and measured using an image analyzing program. Interestingly, among all Spearman rank correlations between chronologies and hydrologic/climatic variables, the strongest associations were found between earlywood vessel chronologies from floodplain trees and spring river discharge data. High water discharge in the spring was negatively correlated to earlywood vessel area and inversely correlated to number of vessels. The mean earlywood vessel area chronology developed from floodplain trees was found to be the best proxy for high magnitude flood events and it has potential to be used for flood reconstructions.

Stable carbon isotope ratios (δ13C) in tree-ring cellulose are modified by environmental conditions occurring during carbon fixation. Researchers have however not reached a consensus as to whether extractives, lignin and/or hemicelluloses, all with specific isotopic signatures, should be removed prior to dendroisotopic analysis. The topic of the first paper dealt with the comparison of Thuja occidentalis L. wood components and their suitability for subsequent dendroisotopic analyses. It was recommended that holocellulose be isolated since an alpha-cellulose yield may be too low for subsequent mass spectrometer analysis, especially when narrow rings are encountered and multiple stable isotope analyses are to be performed per sample. The second paper investigated the associations between the ring-width and δ13C chronologies with climate variables. The δ13C chronology spanned from 1650 to 2006 A.D. and incorporated dead and living T. occidentalis trees selected from two sites in central Manitoba, Canada. Compared to the δ13C values, ring width was more often associated with climate conditions in the year prior to ring formation. However, moisture stress was limiting for both radial growth and carbon assimilation. During the year of ring-formation, ring width was associated with spring and early summer conditions whereas, δ13C was more indicative of overall summer conditions. Nonetheless, each of ring width and δ13C contained individualistic climate information which could be used in tandem for long-term climate reconstruction.

Little research has been conducted on the sensitivity of earlywood vessel in ring-porous tree species in response to flooding. The impact of flooding and climate on vessel characteristics in black ash (Fraxinus nigra Marsh.) was studied in the boreal region of Lake Duparquet, northwestern Quebec. In addition to standard tree-ring measurements, numbers and cross-sectional area of earlywood vessels were examined and measured using an image analyzing program. Interestingly, among all Spearman rank correlations between chronologies and hydrologic/climatic variables, the strongest associations were found between earlywood vessel chronologies from floodplain trees and spring river discharge data. High water discharge in the spring was negatively correlated to earlywood vessel area and inversely correlated to number of vessels. The mean earlywood vessel area chronology developed from floodplain trees was found to be the best proxy for high magnitude flood events and it has potential to be used for flood reconstructions.

Stable carbon isotope ratios (δ13C) in tree-ring cellulose are modified by environmental conditions occurring during carbon fixation. Researchers have however not reached a consensus as to whether extractives, lignin and/or hemicelluloses, all with specific isotopic signatures, should be removed prior to dendroisotopic analysis. The topic of the first paper dealt with the comparison of Thuja occidentalis L. wood components and their suitability for subsequent dendroisotopic analyses. It was recommended that holocellulose be isolated since an alpha-cellulose yield may be too low for subsequent mass spectrometer analysis, especially when narrow rings are encountered and multiple stable isotope analyses are to be performed per sample. The second paper investigated the associations between the ring-width and δ13C chronologies with climate variables. The δ13C chronology spanned from 1650 to 2006 A.D. and incorporated dead and living T. occidentalis trees selected from two sites in central Manitoba, Canada. Compared to the δ13C values, ring width was more often associated with climate conditions in the year prior to ring formation. However, moisture stress was limiting for both radial growth and carbon assimilation. During the year of ring-formation, ring width was associated with spring and early summer conditions whereas, δ13C was more indicative of overall summer conditions. Nonetheless, each of ring width and δ13C contained individualistic climate information which could be used in tandem for long-term climate reconstruction.

It has become increasingly clear that climate fluctuations during the Holocene interval were unusually frequent and rapid, and that our current understanding of the temporal and spatial distribution of these oscillations is incomplete. Little paleoenvironmental research has been undertaken on the windward side of the central Coast Mountains of British Columbia, Canada. Very high annual orographic precipitation totals, moderate annual temperatures regulated by the Pacific Ocean, and extreme topographic features result in a complex suite of microclimate conditions in this largely unstudied area. Dendroclimatological investigations conducted on a steep south-facing slope near Confederation and Franklin glaciers suggest that both mountain hemlock (Tsuga mertensiana) and subalpine fir (Abies lasiocarpa) trees at the site are limited by previous year mean and maximum summer temperatures. A regional subalpine fir chronology for the central and southern Coast Mountains indicates that subalpine fir trees at the study site experience physiological stress with warm summer temperatures, despite the high annual precipitation totals experienced there. This response is likely a result of the extreme gradient and the aspect of the slope at the sampling location, underscoring the importance of site characteristics on annual radial tree growth. Local (AD 1820-2008) and regional (AD 1700-2008) tree ring width chronologies were used to reconstruct previous July mean and maximum temperatures, explaining between 13% and 36% of the variance in climate. The proxy record features cool intervals that are comparable to other paleoenvironmental research from the region, and cyclical oscillations in temperature commonly associated with the El Niño Southern Oscillation and Pacific Decadal Oscillation. Century-scale fluctuations may be connected to changes in solar irradiance. Dendroglaciological investigations were undertaken at the confluence of the Confederation and Franklin glaciers with the intention of exploring the Holocene behaviour of low-elevation maritime glaciers in this region. These glaciers are suspected to be sensitive to variations in the mean position of winter freezing level heights and warm winter temperatures, and may respond differently to changes in climate than more continental glaciers. Buried wood samples were radiocarbon-dated and cross-dated to construct three floating chronologies. Float A (r = 0.467) suggests an early Little Ice Age advance of the two glaciers, and Float B (r = 0.466) suggests an early Tiedemann advance of Confederation Glacier. Float C (r = 0.519) is dated to the Garibaldi Phase of glacier expansion, but may not have been killed by glacial activity. The temporal synchronicity of these findings with glacial events documented throughout the region suggests a spatially coherent response of maritime and continental glaciers to the dominant climate–forcing mechanisms operating in Pacific North America throughout the late Holocene. The dendroclimatological and dendroglaciological findings of this study help to fill a spatial research gap in the current understanding of Holocene climate variations in British Columbia. Because of the complex and at times topographically-controlled response of conifers to climate in the study area, this region may provide a particular challenge in terms of reconstructing Holocene climate variability.