In this paper, we develop a stochastic differential equation model to simulate the movement of a social/subsocial spider species, Anelosimus studiosus, during prey capture using experimental data collected in a structured environment. In a subsocial species, females and their maturing offspring share a web and cooperate in web maintenance and prey capture. Furthermore, observations indicate these colonies change their positioning throughout the day, clustered during certain times of the day while spaced out at other times. One key question was whether or not the spiders spaced out "optimally" to cooperate in prey capture. In this paper, we first show the derivation of the model where experimental data is used to determine key parameters within the model. We then use this model to test the success of prey capture under a variety of different spatial configurations for varying colony sizes to determine the best spatial configuration for prey capture.

Niche partitioning is a major component in understanding community ecology and how different species divide limited environmental resources, enabling them to coexist. Temporal niche partitioning has been widely studied in a broad sense, such as in species that forage on similar nutritional sources dividing activity along diurnal and nocturnal classifications. Here, we approach this temporal niche partitioning with higher resolution to investigate partitioning between species within the same broad temporal and foraging niche. Two species of diurnal orb-weaving spiders (Araneae: Araneidae), Verrucosa arenata and Micrathena gracilis, both construct their orbs in spatially similar locations throughout the understory of deciduous forests in the morning, forage on flying insects throughout the day, and retreat in the evening. However, despite consisting of what appear to be roughly similar total lengths of adhesive silk in the capture spiral, overall orb structure is starkly different: V. arenata orbs are relatively large in diameter and sparse with capture threads; M. gracilis orbs, condensed in diameter and tightly coiled. What other differences might distinguish foraging strategy within this same niche? With extensive observation in their natural environment, we have found that these two species employ two distinct strategies by modulating behavior and orb structure: V. arenata construct orbs earlier in the day, resulting in a longer foraging period. However, V. arenata webs are more likely to be destroyed during the day such that there is a higher variance in foraging duration in V. arenata. We also found that V. arenata actively capture and consume more large prey and that M. gracilis more passively capture and consume small prey more reliably. These data suggest that these species have evolved different foraging strategies with V. arenata being risk-prone and M. gracilis being risk-averse. This study provides a more nuanced analysis of niche partitioning between species occupying otherwise similar temporal, habitat, and foraging niches.

Vigilance behavior can aid in the detection of predators and may also play a role in observation of conspecifics, in food acquisition, and in the prevention of kleptoparasitism. However, in most occasions, vigilance is most important as an antipredator function. Generally, factors that increase the risk of predation also increase the amount of vigilance. We examined whether the reintroduction of the large predator, the wolf, in Yellowstone National Park (YNP) would influence coyote vigilance and foraging ecology. From December 1997 to July 2000, we collected 1743 h of coyote activity budgets. Coyote home ranges occurred within wolf territories (termed high-use or nonbuffer zone areas) and also between them in buffer zones. In high wolf use areas as well as when wolves were present, coyotes fed on carcasses much more; however, they increased the amount of vigilance and decreased rest to prevent predation. Wolf kills may provide a quick source of food and be energetically advantageous to coyotes; however, costs include increased vigilance, decreased rest, and a higher predation risk. Vigilance and avoidance behavioral responses to the reintroduction of large predators may ultimately be more common outcomes than actual killing by competing carnivores of prey. Keystone carnivore reintroductions have a variety of cascading effects throughout the ecosystem and can be driven by both numeric responses (trophic cascades) and behavioral responses ("behavioral cascades"). Behavioral cascades resulting from increased vigilance or spatial changes may lead ultimately to numeric changes and trophic cascades.

No description available.

Turbidity is an important measurement of water quality, considering it describes water clarity and is an indirect indicator of light transmittance in the water column. Turbidity may impact fish that rely on vision to forage by affecting ability to search for prey. Largemouth bass (LMB; Micropterus salmoides) is a visual predator that may have lower foraging success resulting in reduced growth, reproduction, or survival under conditions of high turbidity. I conducted a field experiment in Wolf Lake, Mississippi to assess impacts of turbidity on diet and distribution and a laboratory experiment with manipulated turbidity levels (0 – 250 NTU) to assess foraging efficiency of LMB. There were no significant relationships between turbidity and diet or distribution in the field study. However, there were significant effects of turbidity on foraging efficiency in laboratory trials. These results indicate that assessing turbidity is important to manage LMB in systems susceptible to high turbidity levels.

During 2008 – 2009, we quantified foraging behavior of adult Atlantic puffins (Fratercula arctica) by deploying time-depth recorders (TDRs) on 18 adults and collected morphological measurements from 40 adults nesting on Petit Manan Island, Maine. Dive data were successfully retrieved from 5 birds foraging for 14 days in 2008, and 8 birds foraging for 18 days in 2009. Pooling across all birds, a total of 8,097 dives were recorded, with peaks in activity during 0400-0800 and 1600-2000, and no diving between 2100 and 0400. Mean (± SD) dives/bird/day was 276.4 (± 84.7), with dives grouped into bouts lasting 17.8 (± 31.5) minutes, consisting of 8.9 (± 3.4) dives. Dive depth was less than 15 m for 86% of the dives. Mean maximum dive depth across birds was 9.7 (± 1.7) m, with the deepest dive being 40.7 m. Females made fewer deep dives (27-41 m), had more midday dives (1000 - 1559), and their dives were spread across a greater number of bouts per day than males. Given a mean foraging trip length of 60.1 (± 38.3) minutes for 26 birds observed in 2009, we estimate that adult puffins foraged, on average, within 31 km of the colony. Morphological measures were recorded by a single observer and included body mass, wing chord, bill depth, bill length, culmen, and head-bill length, and represented 19 males and 21 females, based on blood sample analysis. Data were analyzed using classification trees, and our final tree used culmen length and bill depth to correctly classify 34 of 40 (85%) birds (kappa = 0.695, P < 0.01). Use of our model can greatly improve the ability of biologists to identify sex of puffins in the field at this colony site, but variability in morphological data we collected at addition colonies indicates that future work is needed to determine its applicability throughout the Gulf of Maine. During 2009, we measured burrow characteristics of alcids and empty burrows. Breeding success and burrow characteristics were measured for nests of 104 puffins, 58 guillemots and 4 razorbills, with burrow characteristics measured for an additional 12 guillemot and 56 empty burrows. Mean diameter of burrow openings of puffins, guillemots, razorbills and empty burrows were significantly different, and artificial puffin burrows had significantly smaller openings than natural while artificial and natural guillemot burrows had similar opening diameters. Hatch, chick, and nest success of puffins was similar among burrow types, but guillemots had higher hatch and nest success in artificial burrows. The variables we used to create models for predicting hatch and nest success for puffins and guillemots had inadequate discriminatory power to predict success.

No description available.

No description available.

Aspen is a key resource in the Rocky Mountain Region for wildlife forage and habitat, lumber products, scenery, and plays important roles in fire ecology and hydrological processes. There is evidence of aspen decline over much of the Intermountain West for approximately 100 years. In Dixie and Fishlake National Forests, UT, aspen distribution has decreased by nearly half. Causes of this decline are not well understood, although wildlife browsing by ungulates has been implicated as playing a major role. The objective of this research was to examine what soil or plant factors might be involved in wildlife browse choice in aspen. Twenty-two pairs of moderately and intensively browsed sites were studied to identify factors related to browse preferences over two field seasons. In the summer of 2008, sites were sampled in June, July, and August, and in the summer of 2009 sites were sampled in August only. Soils were analyzed for pH, EC, total nitrogen and carbon, and mineral nutrients. Leaf tissue samples were analyzed for defense chemical (tannin and phenolic glycoside) concentrations, mineral nutrients (via acid digestion), acid-detergent fiber, water content, carbon:nitrogen ratio, and non-structural carbohydrate (sugar) concentration. No significant difference in phenolic glycoside concentrations between moderately and intensively browsed sites was found. Tannins were highest in sites with intensive levels of browsing. Iron was significantly higher and zinc lower in intensively than moderately browsed sites. Leaf moisture was also significantly lower in intensively browsed sites. In the absence of differences in phenolic glycosides, ungulates may be selecting browse sites based on iron requirements. Seasonal changes in the studied factors could be identified in 2008. Over the course of the summer, we found significant decreases in nitrogen, phosphorus, potassium, sulfur, zinc, iron, copper, phenolic glycosides, and moisture concentration. Seasonal increases in calcium, sodium, tannins, sugars, acid-detergent fiber, and carbon:nitrogen ratios were observed. The need for large ungulates to obtain specific nutrients may indicate that aspen is in higher demand as a forage at different times of the year, particularly in areas with forages low in these nutrients. Our data suggest that aspen high in iron may be at risk since other factors explaining browsing choice were not significantly different in our study. This information can help identify clones that are at risk and direct resources where and when they are needed most.

Human encroachment into wilderness areas can influence the persistence of wildlife populations by decreasing and degrading habitat, displacement, and decreasing survival. For bighorn sheep (Ovis canadensis), some human activities are detrimental, causing both physiological stress and habitat abandonment. Between 1979 and 2000, human recreation has increased over 300% in areas occupied by desert bighorn sheep (O. c. nelsonii) in southeastern Utah. We investigated if an increase in human activity in areas used by bighorns affected their behavior. We observed 34 bighorn sheep using focal-animal sampling for >14 hrs to compare time spent grazing and scanning between areas of high and low human use. We identified group size, presence or absence of a lamb, distance to escape terrain, and human use (high versus low) as potential explanatory variables that influenced grazing and scanning times, and created an a priori list of models based on these variables. We used Akaike's Information Criterion adjusted for small sample sizes (AICc) to rank models, and used model selection to find a best approximating model (lowest AICc value) for both behaviors. Desert bighorn sheep spent less time grazing and more time scanning in high human use areas (22% grazing, 29% scanning) than in low human use areas (54% grazing, 8% scanning). Caution should be taken when considering which areas or trails should be opened during these important seasons to minimize and reduce additional stresses to bighorns caused by human activity. Bighorn sheep populations experienced significant declines after European settlement in North America. Today, the primary practice of bighorn sheep conservation is through population restoration and augmentation from remnant source populations. We conducted a 9-year telemetry study for a source population of desert bighorn sheep in Canyonlands National Park, Utah. We captured and collared 58 bighorn sheep from 2002-2009. To estimate annual and seasonal survival, we used known-fate analysis in Program MARK 4.1. We used model selection to test hypotheses for bighorn survival, including sex, age, human use, year, and month, as possible explanatory variables. There were 20 mortalities during the study. Annual survival ranged from 83% - 88% with no significant variation among any of the years. Model selection results showed that the top six models included a temporal variable (e.g. season or month), and carried 92% of the AICc weight. Population persistence for bighorn sheep can be compromised by high levels of predation, habitat fragmentation, and disease transmitted from domestic sheep. We suggest that land managers continue to maintain the separation of domestic sheep from bighorns in CNP. We also recommend that survival studies continue to ensure that future translocation projects do not occur at the expense of the source population.