Aspects of the biology of the calanoid copepod Pseudodiaptomus hessei are described, with particular reference to its vertical migratory behaviour. The present investigations were carried out largely by means of Nansen-type plankton nets, but several new pieces of apparatus were developed and. are described herein. The daytime vertical distribution of P. hessei varies according to developmental stage and depth of water. In shallow areas of the lake the entire population is benthic or in very close association with the lake bed. In the deepest part of the lake (40m) the naupliar and early (i.e. C I - C III) copepodite stages are essentially pelagic, but the adult and late (i.e. C IV - C V) copepodite stages are predominantly benthic and may be quiescent or infaunal. During the hours of darkness, the calanoids are distributed through the water column. The nauplii are consistently abundant in the surface waters but the distribution of the other stages is not regular. The distribution can be related to lunar intensity in many cases, with the post-naupliar stages frequently occurring deeper in the water column on bright moonlight nights and in the surface waters on overcast moolnless nights. The dusk ascent and dawn descent of the calenoids is clearly related to changes in light penetration in most individuals. The movements of a fraction of the adults occur in the apparent absence of adequate light cues. This behaviour is shown more extensively by the adult females. An endogenous activity rhythm has been shown in the species under laboratory conditions and it is suggested that this may play a part in the migratory movements. A basic examination of the feeding methods, feeding appendages and food sources of adult and late copepodite stages has been made. Changes in feeding intensity through twenty-four have been examined in the field and under laboratory conditions. Using as an index of feeding intensity, the proportion of animals with food in their guts, it has been shown that a pronounced diel difference occurs in adult calanoids in the lake. Feeding is almost entirely restricted to the nocturnal presence of the calanoids in the water coloumn. This difference exists in the pre-adult copepodite stages, but is not nearly as striking. The absence of feeding during daylight is not readily accounted for in terms of food availability and it is attributed to the quiescent or possible infaunal existence of the adults. A periodicity in egg hatching has been shown Naupliar release from the parental egg sac is predominantly a nocturnal phenomenon in the hot and cool seasons. It is suggested that this may be important in attaining a favourable vertical distritution for the nauplii, and may be important in the distribution of the species. A preliminary acccunt of the seasonal cycle and population dynamics of P. hessei is given, based on data collected over two years at a single station. Seasonal changes in calanoid abundance are intermediate between those recorded in truly tropical areas and in temperate latitudes. Potential food sources show relatively little change. The vertical migration of P. hessei is considered in relation to its apparent ecological significance and comparisons are drawn with observations made on the same species or other pseudodiaptomids in estuaries and lagoons elsewhere.

MSc., Faculty of Science, University of the Witwatersrand, 2011 / Sodic soils are salt-affected soils which are high in sodium in relation to magnesium and calcium. Commonly called sodic sites in the Kruger National Park (KNP), these patches exhibit unique functional characteristics due to the high levels of sodium which cause surface crusting, cracking and the dispersion of clay particles. The aim of this study is to use satellite imagery to map sodic sites in the KNP at different spatial and spectral scales, giving the best option for a repeatable, semi-automated classification. The resultant map of sodic sites for the KNP will be used as a management tool and for future research projects. A field test for sodicity was necessary to collect sufficient ground truth samples for robust accuracy assessment of the image classification. Sodic soils are identified by measuring EC, pH and SAR which are highly variable within site and between testing methods, and therefore not useful for rapid ground truth classification of sodic soils in the field. The sodium level at which clay particle dispersion takes place varies between soils, but is measurable in the field using the Emerson dispersion test. Laboratory tested sodic soil sites from previous research re-tested in this study showed positive results for dispersion of clay particles in water. The physical properties of sodic sites described in the literature and observed in the field were applied to classify sodic sites in the KNP in the field using a decision tree, together with results from the dispersion test and the observed presence of the grass species Sporobolus iocladus. Landsat 7 and SPOT 5 imagery cover the whole park, with ASTER, CAO hyperspectral, LiDAR and black and white orthophotos available for selected areas. The topography elements of crest and footslope were derived from the STRM 90m digital elevation model (DEM). Image preprocessing to top of atmosphere reflectance was performed where necessary and visual enhancement techniques and transformations were applied to derive the normalised difference vegetation index (NDVI) and other indices. Spectral signatures were checked against spectral signature libraries, and the class separation was tested using the cluster analysis of spectral signatures. MODIS NDVI averages placed the imagery in phenological context. Object-based image analysis using eCognition was applied to classify the sodic sites of the Shingwedzi and Ripape River catchments. The input imagery was segmented into ecologically meaningful patches and classification accuracy was assessed using the field samples collected using the decision tree to identify four classes: sodic sites (bare and woody), river sand, riverine vegetation and savanna areas. Comparison of the accuracy assessments for the Shingwedzi study site showed that the Landsat 7 and SPOT 5 classification algorithms gave an overall kappa index accuracy of 89% and 78% respectively, and a sodic site kappa index of 90% and 89%. Validation results using the ground truth samples gave an overall kappa index accuracy of 61% for Landsat 7 and 52% for SPOT 5, with a sodic site kappa index of 49% and 39% respectively. The classification algorithms were applied to the Ripape study site for Landsat 7 and SPOT 5 with repeatable results for the SPOT 5 imagery of 88% overall kappa index and 81-93% kappa index for sodic sites using similar seasonal imagery in the wet to early dry season. The Landsat 7 classification algorithm was applied to the entire KNP based on the repeatability results of 56% overall kappa index and 60% sodic site kappa index for the Ripape site. The quest for a repeatable algorithm to classify sodic sites from satellite imagery has been met by the SPOT 5 imagery using scenes acquired at similar seasonal stages. The late wet season or early dry season imagery was used to apply the classification algorithm with the best success. Changes in size or shape of sodic sites over time requires very high resolution imagery and further studies to understand where the edge of sodic sites are detected from imagery, and how the phenology of the vegetation growing on these sites affects detecting any change in size of the sodic site.