La dernière période glaciaire est marquée par la présence de deux grandes calottes boréales recouvrant l’actuel Canada et le nord de l’Eurasie. Ces calottes constituent un élément actif du système climatique en interagissant avec les différentes composantes du système Terre. L’objectif de cette thèse est de déterminer par quels mécanismes les changements de circulation atmosphérique lors du dernier cycle glaciaire induisent potentiellement une téléconnexion entre les paléo-calottes de l’hémisphère nord. L’utilisation d’un modèle couplé climat-calotte simplifié m’a permis de tester séparément l’influence de la topographie et de l’albédo des calottes sur les champs de température et de précipitation lors du dernier cycle glaciaire, et de mettre en évidence le rôle de la circulation atmosphérique dans la synergie entre les paléo-calottes de l’hémisphère nord. Pour étudier plus en détail les mécanismes de cette interaction, l’utilisation d’un modèle de circulation générale s’est avérée nécessaire. J’ai tout d’abord effectué une inter-comparaison des modèles ayant participé à l’exercice PMIP3 pour le dernier maximum glaciaire (DMG). Cette inter-comparaison a permis d’illustrer l’impact des conditions glaciaires sur le décalage du courant-jet en Atlantique Nord et d’établir un lien entre ce décalage et les précipitations au sud de l’Europe. Enfin, à l’aide d’expériences idéalisées menées avec le modèle atmosphérique LMDZ, j’ai pu étudier le rôle de chacune des calottes dans les changements de circulation atmosphérique observés auDMG. Cette étude montre en particulier l’influence notable de la calotte nord-américaine sur le bilan de masse de surface de la calotte eurasienne. / The last glacial period is characterized by the presence of two large ice sheets covering Canada and North Eurasia. These ice sheets are a key element of the climatic system by interacting with all the components of the Earth system. The aim of this thesis is to determine by which mechanisms changes in atmospheric circulation may have induced a teleconnexionbetween the Northern hemisphere paleo-ice sheets. The use of a simplified coupled climate-ice sheet model allowed to test separately the influence of the ice-sheet topography and albedo on temperature and precipitation fields throughout the last glacial cycle and to highlight the role of atmospheric circulation within the synergy of past boreal ice sheets.To investigate in more details the underlying mechanisms, the use of a general circulation model was necessary.I therefore carried out an inter-comparisonof the PMIP3 models to examine the GCM responsesto glacial conditions. This work allowed to determinethe role of glacial conditions on the shift of the NorthAtlantic jet stream position and to establish a relationshipbetween this shift and the amount of precipitationover southern Europe. The last part of this thesisis devoted to the respective role of each ice sheeton atmospheric circulation changes observed underglacial conditions. To achieve this, I performed idealizedexperiments with the atmospheric circulationmodel LMDZ. The results highlight the key influenceof the North American ice sheet on the Eurasian icesheet surface mass balance.

<p>This thesis relates modern speciation theory to the effects of sudden changes in the range and size of populations. Special reference is made to the climatic oscillations during the Quaternary ice ages. A meta-analysis of laboratory experiments showed that support for allopatric speciation is weak, especially for the peripatric type of allopatric speciation. Furthermore, factors traditionally believed to increase the likelihood of speciation have had little effect on the generation of reproductive isolation in speciation experiments. However, the method of testing reproductive isolation appeared important, in the sense that experimentally derived sister populations were likely to demonstrate reproductive isolation from each other but not from the unaffected mother population. Raw data from mating tests showed that the poor isolation between mother and daughter populations was an effect of asymmetric mate preferences towards males from the mother population. This suggests that peripatric speciation can be effective in generating reproductive isolation between sister populations. The proposed mechanism is that males become less attractive to females by losing certain secondary sexual traits during population bottlenecks, and that females shift their preferences towards other male traits. Support for this mode of speciation is found in the widespread bird genus <i>Motacilla</i> (wagtails). This genus is characterised by extensive plumage variation and contains a large number of widely distributed taxa in the northern parts of its distribution. This thesis shows that taxonomic diversity of wagtails is inversely related to complexity in song and to diversity in molecular and mitochondrial markers. The northern taxa seem to be descendants of southern populations, which were subjected to bottlenecks during expansions into re-opened habitats after the last ice age. The bottlenecks would not only reduce genetic diversity but also inhibit cultural transmission of song to the leading edge of colonisers, allowing sexual selection on other traits, such as plumage. Rapid plumage differentiation among wagtail taxa appears to be a recurrent process and has lead to convergent evolution, making the currently recognised species <i>Motacilla flava</i> (Yellow Wagtail) polyphyletic.</p>

This thesis relates modern speciation theory to the effects of sudden changes in the range and size of populations. Special reference is made to the climatic oscillations during the Quaternary ice ages. A meta-analysis of laboratory experiments showed that support for allopatric speciation is weak, especially for the peripatric type of allopatric speciation. Furthermore, factors traditionally believed to increase the likelihood of speciation have had little effect on the generation of reproductive isolation in speciation experiments. However, the method of testing reproductive isolation appeared important, in the sense that experimentally derived sister populations were likely to demonstrate reproductive isolation from each other but not from the unaffected mother population. Raw data from mating tests showed that the poor isolation between mother and daughter populations was an effect of asymmetric mate preferences towards males from the mother population. This suggests that peripatric speciation can be effective in generating reproductive isolation between sister populations. The proposed mechanism is that males become less attractive to females by losing certain secondary sexual traits during population bottlenecks, and that females shift their preferences towards other male traits. Support for this mode of speciation is found in the widespread bird genus Motacilla (wagtails). This genus is characterised by extensive plumage variation and contains a large number of widely distributed taxa in the northern parts of its distribution. This thesis shows that taxonomic diversity of wagtails is inversely related to complexity in song and to diversity in molecular and mitochondrial markers. The northern taxa seem to be descendants of southern populations, which were subjected to bottlenecks during expansions into re-opened habitats after the last ice age. The bottlenecks would not only reduce genetic diversity but also inhibit cultural transmission of song to the leading edge of colonisers, allowing sexual selection on other traits, such as plumage. Rapid plumage differentiation among wagtail taxa appears to be a recurrent process and has lead to convergent evolution, making the currently recognised species Motacilla flava (Yellow Wagtail) polyphyletic.