Thesis (Ph. D.)--University of Wisconsin--Madison, 1968. / Typescript. Vita. eContent provider-neutral record in process. Description based on print version record. Includes bibliographical references.

This dissertation expands current theoretical and practical knowledge of variability in the technological strategies practiced by past forager societies. Specifically, it examines the interplay between raw material innate and working properties, and tool design as they relate to skeletal media and technologies. Data were synthesized from existing biomedical literature on the structure and mechanical properties of technologically-relevant osseous media, including bird and cetacean bone, and antler. Original laboratory tests were then conducted to determine the mechanical properties of Young's modulus (intrinsic stiffness), intrinsic strength, and fracture resistance of the compact tissue of reindeer antler, cervid long bones, and the limb bones of the California sea lion. Cervid compact limb tissue is stiff, strong, and brittle, while reindeer antler is flexible and highly fracture-resistant (tough). Air-drying hardens all skeletal tissues, and greatly increases investment times for creating tool blanks of both antler and cervid limb bone. Water -soaking can soften dry antler, but may have little effect on the workability of previously-dried land mammal limb bone. Finally, data on the mechanical and working properties of osseous tissues were applied to an analysis of the raw material selection and tool design strategies practiced by protohistoric Alutiiq foragers of Alaska's Kodiak region. Drawing on a sample of over 300 osseous tools and tool blanks, the engineering designs of five tool types were investigated: unbarbed arrows, barbed sea mammal harpoons, fishing harpoon tips, woodworking wedges, and awls. By employing multiple analytical scales, the study points to multiple design pathways toward a generalized goal of maximizing tool longevity, or circulation time. Tool fracture potential can be reduced through raw material selection and stress-reducing structural design. Alutiiq designs for longevity include nested fish harpoon valves, and the off-set line holes on unilaterally barbed harpoons. Also, both tool types were created most frequently from tough but non-local antler. Tool recycling and conservation to avoid drying and fracture can likewise increase tool use-lives. For osseous tools, maximizing longevity might offset high initial tool production investments. The results are applicable to processes of technology transfer in many protohistoric contexts and the Upper Paleolithic of Eurasia.

Thesis (Ph. D.)--University of Wisconsin, 1966. / Typescript. Vita. eContent provider-neutral record in process. Description based on print version record. Includes bibliographical references.

The Kodiak National Wildlife Refuge is home to one of the highest concentrations of brown bears in Alaska. As the public demand for bear viewing opportunities continues to increase, managers are faced with the challenge of accommodating this new kind of visitor use on a refuge that was traditionally managed for the sustainable hunting of bears. To inform the public use management planning process, the Kodiak Refuge allocated funding to support social science research that objectively assessed the current nature of bear viewing opportunities and the factors that influence the quality of those opportunities. Ecologist Aldo Leopold claimed that the outstanding advance of modern ecology would be the “inevitable fusion” of the social and natural sciences. Therefore, a conjoint constitution framework inspired by Freudenburg, Frickel, and Gramling (1995) enabled this study to examine the active interplay of social and environmental factors in a bear viewing experience. Two seasons of research were conducted in partnership with Utah State University. The first season employed qualitative research methods to conduct detailed interviews with a wide variety of bear viewing stakeholders in Kodiak. This process informed the creation of a survey measurement tool that was administered to bear viewers the following summer. Survey results suggest that seeing a larger number of bears and seeing big bears are trip characteristics associated with higher satisfaction among visitors, while closer proximity to bears is associated with learning more about bear behavior. The environmental sociology principle of “conjoint constitution” guided both phases of research by helping to examine how social and physical factors interact with one another to create trip outcomes. Just as there are ongoing biological inventory and monitoring processes that inform refuge management, there should be inventory and monitoring of human activity and the fluent sociological factors influencing the nature of that activity. As the Kodiak Refuge continues its public use planning process, the ongoing integration of both biological and social science data will be critical.

Thesis (Ph. D.)--Bryn Mawr College, 1995. / Includes bibliographical references (p. 748-770).

Seabirds commonly nest colonially on cliffs or inaccessible islands, so that oceanic effects on the quality or quantity of prey fed to chicks more often determine nest success than does terrestrial predation. However, when predators can access nests, impacts can be dramatic. In Kittlitz’s murrelet (KIMU), a rare and recently declining seabird, nests are widely dispersed in poorly accessible areas, growth rates are exceptionally high, and nestling periods are very short. This unique strategy may offset the vulnerability of their ground nests to predators but demands adequate deliveries of high-energy prey. We investigated whether variations in energy content of prey fed to chicks could alter growth rates and resulting duration of exposure to predators, and whether extending predator exposure had important effects on nest success. From 2009‒2016, we measured fish length with cameras, modeled prey energy content from chick energy needs, and measured chick growth rate and nest survival. We monitored an average 17 nests/year (139 total), of which 49% were depredated (almost all by red fox) and 25% survived to fledging. Prey were mainly Pacific sand lance (80%) and capelin (19%), with capelin having 2.3× higher energy per unit length. In a year of slow growth, sand lance energy density needed to increase by 31% from 4.29 to 5.64 kJ/g (within published values), or the proportion of capelin in the diet needed to increase from 5.6% to 27.2%, to achieve maximum chick growth. Adults could supply the energy required for maximum growth by delivering only 1.9 capelin/day versus 5.5 sand lance. Slow growth increased time to fledging by 5 days. This extended exposure to predation could decrease overall nest survival from 14.2% to 13.1%, a relative decrease of 7.7%. With reported breeding propensity of KIMU averaging only 20% and sometimes much lower (range 5 to 45%), even small effects on nest success may be consequential. Although direct predation was the main limitation to nest success, effects of ocean conditions on prey quantity and quality can have overriding effects in some years, and apparently small but substantive indirect effects in other years by mediating exposure to predation. Continued climate warming that adversely affects availability of high-energy fish and allows range expansion of terrestrial predators may have disproportionate effects on species such as KIMU with accessible nests and demands for energy-rich prey.

Thesis (Ph. D.)--University of Washington, 2003. / Vita. Includes bibliographical references (leaves 294-327).

The Kittlitz's murrelet (Brachyramphus brevirostris) is a rare member of the seabird family Alcidae that breeds in coastal areas of Alaska and Beringian Russia. The species belongs to the genus Brachyramphus, an unusual seabird taxon in which all three extant species nest non-colonially, situating their nests up to 75 km inland from coastal marine waters. This nesting strategy is different from that of most seabird species, which tend to nest colonially on remote islands or sea cliffs, where terrestrial predators are generally absent or cannot easily access nests. Within the genus Brachyramphus, Kittlitz's murrelet is notable because a majority of the global population appears to nest on the surface of the ground in rocky alpine habitat near inland or tidewater glaciers, foraging in adjacent marine waters influenced by glacial outflows. The unusual nesting habits of Kittlitz's murrelet have made the study of its nesting ecology difficult, and gaps therefore exist in our understanding of the species' breeding biology. Kittlitz's murrelet populations have declined substantially in core areas of its range, causing the U. S. Fish and Wildlife Service to designate the species as a candidate for protection under the Endangered Species Act. A better understanding of Kittlitz's murrelet nesting ecology is crucial for determining potential causes of these declines and for future management of the species. To this end, I studied Kittlitz's murrelet breeding ecology and nest site selection during 2008-2011 on Kodiak Island, Alaska, in an unglaciated area that was recently found to have large numbers of accessible nests. I and my colleagues found 53 active Kittlitz's murrelet nests in inland scree-dominated habitats and placed remote, motion-sensing cameras at 33 nests. Adults exchanged incubation duties at the nest every 24 or 48 h, almost exclusively during early morning twilight. Following hatching of eggs, parents provisioned their single nestling with an average of 3.9 to 4.8 fish per day, depending on the year. Parental visits to the nest during chick-rearing occurred primarily after sunrise in the early to mid-morning hours, and during evening twilight. Fish were delivered singly to the chick, and Pacific sand lance (Ammodytes hexapterus), a high-lipid forage fish, accounted for about 92% of all identifiable chick meal deliveries. Chick growth rates were high relative to confamilial species, consistent with the high quality of chick diets; the logistic growth rate constant (K) was 0.291, greater than that for any other semi-precocial alcid. Chicks fledged an average of 24.8 d after hatching and asymptotic chick body mass averaged about 135.5 g, approximately 58% of adult body mass. Age at fledging, asymptotic chick body mass (% adult mass), and the number of meal deliveries required to fledge a chick were all lower than or as low as any other species of semi-precocial alcid. The average estimated nest survival rate during 2008-2011 was 0.093 (95% CI = 0.01–0.30), which is extremely low compared to other species in the family Alcidae, and is almost certainly insufficient to sustain a stable population. The primary causes of nest failure were depredation (47% of total nest fates), mostly by red foxes (Vulpes vulpes), and unexplained nestling mortality on the nest (21% of nest fates). Saxitoxin and/or pathogenic endoparasite burdens were observed in five of six necropsied chick carcasses, suggesting possible causes for chick mortality not directly attributable to predation. Habitat characteristics of Kittlitz's murrelet nest sites differed significantly from unused sites at several scales. At a small scale (within 5 m of the nest), nest sites had a lower percent coverage of vegetation and higher percent coverage of intermediate-sized rocks (5–30 cm diameter), compared to randomly selected unused sites. Nest sites were also located on steeper, more north-facing slopes compared to randomly selected sites. Nest sites also had a lower percent coverage of vegetation than randomly-selected sites at larger scales (within 25 m and 50 m of the nest site). Nest sites were located significantly farther from the edge of densely-vegetated habitats than random sites. There was no evidence that nest sites were different from randomly-selected sites in terms of elevation, proximity to ridgelines, or proximity to the open ocean, although a low degree of variation within the study area for these habitat characteristics may have precluded detection of potential differences. Nest survival rates did not co-vary with slope, percent vegetation coverage, distance from vegetated edges, or percent cover of intermediate-sized rocks; however, this result may be an artifact of a limited sample size. The results of this thesis will provide managers with a better understanding of the factors that may limit Kittlitz's murrelet nesting success, such as nest predation and forage fish availability, as well as factors that may influence the quality and distribution of Kittlitz’s murrelet nesting habitat in the future, given on-going and progressive climate change. / Graduation date: 2013