The extant venomous Gila monster and beaded lizards, species of Heloderma, live today in southwestern USA and south along the Pacific coastal region into Central America, but their fossil history is poorly understood. Here we report helodermatid osteoderms (dermal ossicles) from the late Miocene-early Pliocene Gray Fossil Site, eastern Tennessee USA. Twenty-three species of mammals are known from the fauna including abundant Tapirus polkensis, as well as fishes, anurans, salamanders, turtles, Alligator, birds, and snakes. Beaded lizards belong to the Monstersauria, a clade that includes Primaderma + Paraderma + Gobiderma + Helodermatidae (Estesia, Eurheloderma, Lowesaurus, and Heloderma). Osteoderms of lizards in this clade are unique within Squamata; they typically are circular to polygonal in outline, domed to flat-domed in cross-section, have a vermiculate surface texture, are not compound structures, and do not have imbricate surfaces as on many scincomorph and anguid lizards. We review and characterize the osteoderms of all members of Monstersauria. Osteoderms from the cranium, body, and limbs of Heloderma characteristically have a ring-extension (bony flange) at least partly surrounding the dome. Its presence appears to be a key character distinct to all species of Heloderma, consequently, we propose the presence of a ring-extension to be an apomorphy. Three osteoderms from the Gray Fossil Site range from 1.5 to 3.0 mm in diameter, have the circular shape of helodermatid osteoderms with a domed apical surface, and have the ring-extensions, permiting generic identification. Macrobotanical remains from the Gray Fossil Site indicate an oak-hickory subtropical forest dominated by Quercus (oak) and Carya (hickory) with some conifer species, an understorey including the climbing vines Sinomenium, Sargentodoxa, and Vitis. Plant and mammal remains indicate a strong Asian influence.

Deformation of the Eastern Cordillera, as a double-verging thrust belt that separates the Magdalena Valley from the Llanos Basin, is a defining moment in the history of the northern Andes in South America. Here we examine the age and depositional setting of the youngest stratigraphic unit in three sectors of the Eastern Cordillera: (i) the Santa Teresa Formation (western flank), (ii) the Usme Formation (southern central axis), and (iii) the Concentración Formation (northeastern central axis). These units were deposited prior to the main Neogene deformation events. They represent the last preserved record of lowland conditions in the Eastern Cordillera, and they are coeval with a thick syn-orogenic deposition reported in the Llanos Basin and Magdalena Valley. Based on palynological data, we conclude that the upper Usme Formation was deposited during the Bartonian-earliest Rupelian? (Late Eocene-earliest Oligocene?); the Concentración Formation was deposited during the Late Lutetian-Early Rupelian (Middle Eocene to Early Oligocene), and the upper Santa Teresa Formation was accumulated during the Burdigalian (Early Miocene). These ages, together with considerations on maximum post-depositional burial, provide important time differences for the age of initial uplift and exhumation along the axial zone and western foothills of the Eastern Cordillera. The switch from sediment accumulation to erosion in the southern axial zone of the Eastern Cordillera occurred during the Rupelian-Early Chattian (Oligocene, ca 30 to ca 26 Ma), and in the northeastern axial zone occurred prior to the latest Chattian-Aquitanian (latest Oligocene-Early Miocene ca 23 Ma). In contrast, in the western flank, the switch occurred during the Tortonian (Late Miocene, ca 10 Ma). In addition, we detected a marine transgression affecting the Usme and Concentración formations during the Late Eocene; coeval marine transgression has been also documented in the Central Llanos Foothills and Llanos Basin, as evidenced by the similarity in floras, but not in the western foothills. Our dataset supports previous sedimentological, geochemical and thermochronological works, which indicated that (i) deformation in the Eastern Cordillera was a diachronous process, (ii) the sedimentation along the axial zone stopped first in the south and then in the north during the Oligocene, (iii) depositional systems of the axial zone and central Llanos Foothills kept partly connected at least until the Late Eocene, and (iv) Miocene strata were only recorded in adjacent foothills as well as the Magdalena and Llanos basins.

The Miocene climate evolution in North China is preliminarily discussed by means of comparisons in seven climate parameters quantitatively reconstructed by the Coexistence Approach on 34 selected macro- and microfloras over North China. The Miocene temperatures show no great difference in the western and eastern part of North China. Temperature fluctuations, particularly in mean annual temperature, are found within floras from several sites. The fluctuation pattern, from a climate optimum in the Mid Miocene to cooling decline in the Late Miocene, is generally consistent with the global trend of Miocene temperature change. The reconstructed precipitation from all the sites studied shows much wetter conditions in North China during the Miocene than at present, which corroborates the results from paleoprecipitation proxy of fossil mammals. Like the situation in paleo-temperature, the Miocene precipitation from North China shows no distinct difference between the western and eastern regions. It is suggested that North China, particularly in the western part, was by no means under an arid or semi-arid environment during the Miocene. North China is an ideal region for study of the impact of the East Asian monsoon system, however, the pattern of precipitation change derived from the monsoon index (MSH) and mean annual precipitation (MAP) shows contradictory results. Therefore, there appears no definite conclusion on when the East Asian summer monsoon intensified. Possible reasons for inconsistency in temperature and precipitation changes are discussed. Directions of future work to improve the resolution of climate evolution are also pointed out.

Pinus prekesiya Xing, Liu et Zhou sp. nov. was described as a new species on the basis of two well preserved ovulate cones from the upper Miocene of central Yunnan, southwestern China. It is the first fossil record of three dimensionally preserved Pinus ovulate cones from China. Morphological comparisons with 15 previously published Cenozoic cones and seven related extant pine species reveal that the fossil cones are identified as a new species, P. prekesiya sp. nov., which belongs to subsection Pinus of subgenus Pinus. The new species shows a combination of characters of P. kesiya and P. yunnanensis, but has a closer affinity with P. kesiya which occurs in the humid region of Yunnan and therefore suggests a more humid climate in central Yunnan during the late Miocene than today. The general cooling trend during the late Neogene and topographic change due to the dramatic Tibetan uplift might have caused a vicariant origin of P. kesiya and P. yunnanensis from the ancestral P. prekesiya.

Through the Cenozoic, paleoclimate records show general trends of global cooling and increased aridity, and environments in North America shifted from predominantly forests to more open habitats. Paleobotanical records indicate grasses were present on the continent in the Eocene; however, paleosol and phytolith studies indicate that open habitats did not arise until the late Eocene or even later in the Oligocene. Studies of large mammalian herbivores have documented changes in ecomorphology and community structure through time, revealing that shifts in mammalian morphology occurred millions of years after the environmental changes thought to have triggered them. Smaller mammals, like rodents and lagomorphs, should more closely track climate and habitat changes due to their shorter generation times and smaller ranges, but these animals have received much less study. To examine changes in smaller mammals through time, we have assembled and analyzed an ecomorphological database of all North American rodent and lagomorph species. Analyses of these data found that rodent and lagomorph community structure changed dramatically through the Cenozoic, and shifts in diversity and ecology correspond closely with the timing of habitat changes. Cenozoic rodent and lagomorph species diversity is strongly biased by sampling of localities, but sampling-corrected diversity reveals diversity dynamics that, after an initial density-dependent diversification in the Eocene, track habitat changes and the appearance of new ecological adaptations. As habitats became more open and arid through time, rodent and lagomorph crown heights increased while burrowing, jumping, and cursorial adaptations became more prevalent. Through time, open-habitat specialists were added during periods of diversification, while closed-habitat taxa were disproportionately lost in subsequent diversity declines. While shifts among rodents and lagomorphs parallel changes in ungulate communities, they started millions of years earlier than in larger mammals. This is likely a consequence of the smaller mammal' greater sensitivity to environmental changes and more rapid evolution. These results highlight the importance of examining understudied members of vertebrate faunas for understanding the evolution of terrestrial communities through time.

Continental scale studies on ungulate crown heights in relation to climate and habitat changes have revealed a correlation between increasing hypsodonty and a shift to more arid environments. Small mammals have been shown to adapt to changing habitats millions of years earlier than larger mammals. In this study I examined fossil localities throughout the last 37 Ma across North America. Diversity of rodents and lagomorphs were analyzed through this time period, with examination of community structure characterized by relative percentages of taxa with different crown heights. Overall, a decrease in precipitation and temperature was found across North America from 37 Ma to the present. The Mid Miocene Climatic Optimum (around 15 Ma) was an pronounced period of warming, comparable to the warming we see today. Nebraska and California showed increases in crown height before Oregon. Overall, these findings help demonstrate how communities react at different time scales to climate change.

The impact of large-scale volcanic eruptions on landscapes can affect many processes ranging from interrupting or redirecting regional soil forming processes and hydrological systems to generating temporary changes in global climate. Though more studies exist every year, less is known of the direct impact of large-scale volcanic eruptions on ecosystems and extinction, while even less is known of their impact on speciation. In deposits throughout the Pacific Northwest a special combination and association of volcanic magnitude with palaeoecological yield frequently presents unique prospects for inquiry. In this study, particular attention is given to large, late Miocene ash-flow tuffs of central and eastern Oregon, the Rattlesnake Tuff among them. Additionally, a scenario is tested whereby populations become isolated from one another across the expanse of ash-flow tuff and enclosed by nearby physical barriers. Exploration into this scenario and the effects of ecological recovery on speciation in mammals are made through a cellular automaton created here and speciation modeling by H.A. Orr. The cellular automaton is validated by a small data set from New Zealand with results of all models compared to studies in ecology, the fossil record, genetics, and island evolution. Results suggest ecosystem recovery is much faster than mammalian species origination rates. Though evolution in mammals is known to occur over relatively short intervals of time (10 - 10 yrs), based on model comparisons, primary productivity generally returns to its carrying capacity two orders of magnitude (10 - 10 vs. ≥ 10 yrs) faster than mammalian speciation.

Widespread, mid-Miocene rhyolite volcanism of eastern Oregon that are coeval or slightly postdate flood basalts of the Columbia River Basalt Province allows for mapping crustal domains using radiogenic and stable isotopes. Rhyolites are thought to be derived in large part by partial melting of the crust and thus yield direct information on the composition of the crust. Silicic volcanism is expressed in the form of numerous domes and tuffs exposed over a wide area (~300 km in N-S dimension and ~200 km in E-W dimension) west of the presumed craton boundary, which runs parallel but mostly east of the Oregon-Idaho state border as delineated by geophysical characteristics and isotopic transitions, including the 87Sr/86Sri = 0.7060 line (MSL) and 87Sr/86Sri = 0.7040 (CSL). 87Sr/86Sri of twenty-seven silicic units are variable and some are high. Sr isotopic ratios are inconsistent with the location of the traditional MSL and CSL boundaries. A primary control on the 87Sr/86Sri isotope variations may reflect changes in the crustal make-up of Paleozoic accreted terranes of a particular area rather than arising from a westward-dipping decollement that moved cratonic lithosphere below accreted terranes in eastern Oregon. A secondary control on observed isotopic ratios may be related to the amount and composition of basalt involved in the generation of rhyolites. This could lead to higher or lower 87Sr/86Sri relative to the surrounding crust because de facto coeval mafic magmas of the Columbia River Basalt Group have a wide range of Sr isotopic signatures. While Pb isotope data is incomplete for all samples of this study, the available data indicate a significant range in Pb isotopes. Yet, data of individual regions tend to plot close to one another relative to the entire data distribution. Comparison of samples from this study in a more regional view indicates the samples generally fall within the previously defined lead isotope boundaries of the main-phase Columbia River Basalt Group lavas. [lowercase delta]¹⁸O values range from below 2 parts per thousand to above 9 parts per thousand. In addition, there is a crude trend of rhyolites having lower [lowercase delta]¹⁸O and more radiogenic ⁸⁷Sr/⁸⁶Sr[subscript i] ratios. The lowest oxygen ratios (< 2 parts per thousand) are found in rhyolites ~80 km west of the cratonic margin, potentially reflecting remelting or assimilation of hydrothermally altered crust. Low [lowercase delta]¹⁸O of selected rhyolite flows cannot be explained by remelting of Cretaceous plutons of the Idaho Batholith and appear irreconcilable with remelting of altered silicic rocks at centers of multiple, confocal caldera cycles- both processes that have been proposed to explain low [lowercase delta]¹⁸O of rhyolites of the Snake River Plain-Yellowstone area.

The chalicothere Moropus was a rare perissodactyl present in the Great Plains region of North America through much of the Miocene. A temporal gap in named species of Moropus is present in the early Hemingfordian North American Land Mammal Age. This gap is filled by specimens currently referred to as Moropus sp. from the Batesland Formation in southwest South Dakota, and unnamed specimens of Moropus in the Runningwater Formation in northwestern Nebraska. A comparison of the fossils of Moropus nsp. from the Batesland Formation with those of previously described chalicothere species from the Greats Plains region, such as Moropus elatus, Moropus hollandi, Tylocephalonyx skinneri, Moropus merriami, and Moropus matthewi, in addition to Metaschizotherium bavaricum and Metaschizotherium fraasi from the Miocene of Southern Germany, illustrates that there are substantial differences in morphology between equivalent skeletal elements. Based on these findings, I propose that the specimens of Moropus from the Batesland Formation belong to a new species.

No description available.