Prairie rivers of the Great Plains, USA, provide important habitat for the federally threatened piping plover (“plover”, Charadrius melodus). Plovers nest on open to sparsely vegetated river sandbars, and their demographic rates are closely linked to habitat availability and quality, as well as river flow. The Niobrara River in northern Nebraska has supported 22–41% of the state’s plovers since species listing in 1986, but the population and habitat are relatively understudied, and both have declined since 2010. The objectives of this study were to understand plover demography, habitat, and the role of the Niobrara in the regional plover population. Periods of high river flow promote creation and maintenance of suitable sandbar nesting habitat, but increased river flow during the plover breeding season can decrease nest and chick survival. We estimated the effect of daily peak river flow on survival rates of 115 nests and 66 chicks on the Niobrara River, 2010–2016, using logistic exposure and Cormack-Jolly-Seber models, respectively. We monitored 1,874 banded hatch-year and adult birds across the regional population (Niobrara River, Lewis and Clark Lake, Gavins Point Reach segment of the Missouri River), and used multi-state mark-recapture models to estimate survival and inter-annual dispersal probabilities among sites relative to habitat availability. We developed land cover datasets from high-resolution aerial imagery to quantify suitable habitat and compare the relative effects of habitat characteristics on nest-site selection and nest success for a subset of years (2010, 2012, 2014, and 2016) using logistic regression models. We included data from a sympatric nester with similar nesting habitat needs, the interior least tern (“tern”, Sternula antillarum athalassos), to improve precision of our models. We compared 63 plover and 92 tern nests to 292 random unused points, and 73 successful (hatched ≥1 egg) to 79 failed nests. Low nest and chick survival and high emigration from the Niobrara appear to be important factors contributing to population decline. Daily nest and chick survival were negatively related to river flow. Nest-site selection was based primarily on distance to the river bank (i.e., the nearest potential source of predators), yet flooding (eggs submerged or washed out of the nest bowl during increased river flow) caused at least as many nest failures as predation. Nests predominantly were surrounded by dry sand habitat, indicating some degree of flood avoidance, but were no farther from water than random, and drier nest sites were no less likely to fail. Dispersal occurred throughout the regional population, but plovers were more likely to leave the Niobrara than to enter it. Expansive flood-created sandbars on the Missouri River, concurrent with a trend towards more vegetated and saturated habitat on the Niobrara, may have drawn birds from the Niobrara population, especially those that dispersed to the Niobrara during sustained Missouri River flooding 2010–2011. The outsized negative effect of flooding on nest success, the lack of protection afforded by dry sand nest sites, and selection for nesting habitat based more strongly on predator avoidance than flood avoidance suggest that plovers may have face more frequent and intense levels of breeding season flooding than is typical. Identifying and promoting the processes that contribute to creation and maintenance of high-elevation sandbars on the Niobrara is an important next step towards effective management of nesting birds. / Master of Science / Prairie rivers of the Great Plains, USA, provide important habitat for a federally threatened shorebird, the piping plover (“plover”, Charadrius melodus). During their breeding season (April–August), plovers nest and raise chicks on open to sparsely vegetated areas of river sandbars, and their survival and reproductive success are closely tied to amount and quality of sandbar habitat. The number of plovers on the Niobrara River in northeast Nebraska has declined since 2010, and in this study our objectives were to monitor nests, chicks, and adult birds to document plover survival, movement among neighboring sites, reproductive success, and habitat changes on the Niobrara from 2010–2016. Because plovers nest on the sand, they can lose nests and chicks when river flow increases and washes over sandbars (“flooding”). We found that nest and chick survival sharply decreased when river flow was high. Plovers chose nest sites in dry sand areas, which are typically the tallest part of a sandbar and most protected from flooding, but these nests were no more likely to hatch than those in areas with wetter (lower elevation) sand. Plovers primarily chose nest sites far from the river bank, which was the nearest potential source of predators, yet flooding caused at least as many nest failures as predation. Plovers moved between the Niobrara and nearby Missouri River, but overall were much more likely to leave the Niobrara than to enter it. Expansive sandbar creation on the Missouri River, concurrent with a trend towards lower-quality (more vegetated and saturated) habitat on the Niobrara, may have drawn birds from the Niobrara population, especially those displaced from the Missouri during sustained flooding 2010–2011. Movement to the Missouri River and low nest and chick survival due to flooding contributed to population decline on the Niobrara. The outsized effect of flooding on nest success, the lack of protection afforded by dry sand nest sites, and selection for nesting habitat based more strongly on predator avoidance than flood avoidance suggests that plovers face more frequent and intense levels of breeding season flooding than is typical. Identifying and promoting the processes that contribute to creation and maintenance of high-elevation sandbars on the Niobrara is an important next step towards effective management of nesting birds.

Prompted by increased interest in understanding microporosity, recent efforts at describing and classifying pore types in mudstones have focused primarily on siliceous, gas producing unconventional reservoirs with little attention being paid to carbonate, mixed oil-and-gas producers. The Niobrara Formation in the Denver-Julesburg Basin is a self-sourced resource play producing oil and natural gas from low permeability chalks. Key reservoir lithologies consist of chalk, chalky marl and marl. These lithologies contain flattened chalk fecal pellets which play a significant role in providing porosity. Integration of depositional fabric with pore-type distribution emphasizes the unique textural and depositional nature of chalk and provides a starting point for evaluation of diagenetic porosity modification. Chalk depositional textures comprise two main subdivisions. The first, called rainstone, includes chalks that form largely from settling of planktonic skeletal remains and fecal pellets as marine snow. New terms related to pelagic chalk textures are pelagic mudstone, pelagic wackestone, and pelagic packstone. The second, called allochthonous chalk, consists of chalks formed from syndepositional tectonic disruption of the seafloor, resulting in mass-movement and redeposition of chalk as turbidites and slide sheets. New terms related to allochthonous chalk textures are allomudstone, allofloatstone, and allorudstone. A chalk porosity classification consisting of four major pore types is presented that can be used to quantify Niobrara chalk pores and relate them to depositional texture, porosity networks, diagenetic history, and pore distributions. Interparticle porosity occurs largely between coccoliths and coccolith fragments, and decreases with burial ranging from 27-38% to 5-17%. Intraparticle porosity occurs within chalk pellets, coccospheres, coccolith plates and foraminifera tests, and also decreases with burial. Organic matter pores are intraparticle pores located within organic matter and are related to hydrocarbon generation. Channel pores, where present, can have significant influence on hydrocarbon storage and permeability networks. In the Niobrara, burial diagenesis in the form of mechanical compaction, chemical compaction, and syntaxial cement overgrowths, modifies pore shape and abundance. Porosity distribution is controlled by the abundance of chalk pellets and the mineralogy of the matrix. Permeability is a function of matrix lithology (micrite-rich vs. silt- and clay-rich). Understanding chalk depositional and diagenetic processes, and how they relate to porosity formation and pore evolution provide a foundation for more accurately predicting the occurrence and distribution of hydrocarbon source and reservoir rocks within the Niobrara.