A global extinction event has been documented in protozoan foraminifera in the late Pliocene to Pleistocene. The timing of the extinction event varied depending on location, however for Ocean Drilling Project Site 1125, disappearances occurred between 2.5 and 0.57 Ma, with the major decline approximately 1.1 Ma. In order to determine if this event affected benthic organisms other than protozoans, this study was undertaken to determine how podocopid ostracods (Crustacea) recovered Ocean Drilling Program Site 1125 responded. The present study was hindered by the small number of valves recovered; the fact that a large proportion of taxa found were undescribed and new to science; and the current state of taxonomic scheme that is under significant revision. These factors meant that a comprehensive comparison could not be achieved. Despite this, counts of ostracod valves and assessments of diversity from this study reveal a significant increase in both parameters from approximately 900-600 ka. Three possible causes were investigated to account for this increase, sediment type and sample size; affects of taphonomy, mainly dissolution; or an actual biotic 'event'. Statistical analyses showed that although sample size did have some effect, it was not the sole reason for the increase in ostracod numbers. Dissolution had an expected affect on the percentage of juveniles but no correlations were found with other sample characteristics. Sedimentation rate was investigated but this also proved unrelated. Therefore, it is suggested that the increase in total ostracod valves and diversity which occurs between 900 and 600 ka was in fact a natural, biotic 'event'. This preliminary evidence suggests that an oceanographic event that has negatively impacted on the foraminifers has had the reverse affect on the ostracod assemblage, in the sense that both population size and diversity increase during that time.

Stable isotope values of ostracod calcite provide a record of variation in ¥ä18O(H2O) values and water temperature from the late glacial to mid-Holocene in Western Ireland. Lough Monreagh, located in County Clare, Western Ireland, contains marl sediment that includes pristine ostracod calcite whose ¥ä18O and ¥ä13C values were evaluated. These values were used relative to modern ecological requirements to derive a paleoenvironmental record for Lough Monreagh that includes water temperature, eutrophication, water depth, as well as terrestrial vegetation and weathering within the lake¡¯s watershed. ¥ä13C values of ostracod calcite presented herein suggest a significant increase in terrestrial vegetation beginning during the Aller©ªd (13,600 cal year B.P.) and extending through to the mid-Holocene (6,997 cal year B.P.). Marl and ostracod ¥ä18O values record variability in temperature and precipitation ¥ä18O(H2O) values that are in turn forced by variation in atmospheric and oceanic circulation. Water temperatures presented herein were calculated from ¥ä18O values of ostracod calcite and marl, constrained by temperature preference and tolerance ranges of ostracod species, yielding the highest resolution temperature record covering this period to date. Over 4,700 ostracods representing all three freshwater superfamilies were counted and identified to evaluate the trophic stage of the lake. The lake was characterized as a clear-, cold-water (~8¨¬C summer water temperature), low-nutrient environment during the Aller©ªd, then freezes abruptly during the Younger Dryas as evidenced by black clay deposits aged 12,800 to 11,300 cal yr B.P. Following the Younger Dryas, transitional warming and increasing terrestrial vegetation are evidenced by decreasing ¥ä13C values of ostracoda and faunal transition to phytophyllic species. Summer water temperatures warm to >16¨¬C, with ostracod species suggesting a shallow-water, fen- and macrophyte-rich environment with abundant plant life in and around the lake by ~8,000 cal yr B.P.

Stable isotope values of ostracod calcite provide a record of variation in ¥ä18O(H2O) values and water temperature from the late glacial to mid-Holocene in Western Ireland. Lough Monreagh, located in County Clare, Western Ireland, contains marl sediment that includes pristine ostracod calcite whose ¥ä18O and ¥ä13C values were evaluated. These values were used relative to modern ecological requirements to derive a paleoenvironmental record for Lough Monreagh that includes water temperature, eutrophication, water depth, as well as terrestrial vegetation and weathering within the lake¡¯s watershed. ¥ä13C values of ostracod calcite presented herein suggest a significant increase in terrestrial vegetation beginning during the Aller©ªd (13,600 cal year B.P.) and extending through to the mid-Holocene (6,997 cal year B.P.). Marl and ostracod ¥ä18O values record variability in temperature and precipitation ¥ä18O(H2O) values that are in turn forced by variation in atmospheric and oceanic circulation. Water temperatures presented herein were calculated from ¥ä18O values of ostracod calcite and marl, constrained by temperature preference and tolerance ranges of ostracod species, yielding the highest resolution temperature record covering this period to date. Over 4,700 ostracods representing all three freshwater superfamilies were counted and identified to evaluate the trophic stage of the lake. The lake was characterized as a clear-, cold-water (~8¨¬C summer water temperature), low-nutrient environment during the Aller©ªd, then freezes abruptly during the Younger Dryas as evidenced by black clay deposits aged 12,800 to 11,300 cal yr B.P. Following the Younger Dryas, transitional warming and increasing terrestrial vegetation are evidenced by decreasing ¥ä13C values of ostracoda and faunal transition to phytophyllic species. Summer water temperatures warm to >16¨¬C, with ostracod species suggesting a shallow-water, fen- and macrophyte-rich environment with abundant plant life in and around the lake by ~8,000 cal yr B.P.

京都大学 / 新制・課程博士 / 博士(理学) / 甲第23714号 / 理博第4804号 / 新制||理||1687(附属図書館) / 京都大学大学院理学研究科地球惑星科学専攻 / (主査)教授 生形 貴男, 准教授 成瀬 元, 准教授 佐藤 活志 / 学位規則第4条第1項該当 / Doctor of Science / Kyoto University / DGAM

A global extinction event has been documented in protozoan foraminifera in the late Pliocene to Pleistocene. The timing of the extinction event varied depending on location, however for Ocean Drilling Project Site 1125, disappearances occurred between 2.5 and 0.57 Ma, with the major decline approximately 1.1 Ma. In order to determine if this event affected benthic organisms other than protozoans, this study was undertaken to determine how podocopid ostracods (Crustacea) recovered Ocean Drilling Program Site 1125 responded. The present study was hindered by the small number of valves recovered; the fact that a large proportion of taxa found were undescribed and new to science; and the current state of taxonomic scheme that is under significant revision. These factors meant that a comprehensive comparison could not be achieved. Despite this, counts of ostracod valves and assessments of diversity from this study reveal a significant increase in both parameters from approximately 900-600 ka. Three possible causes were investigated to account for this increase, sediment type and sample size; affects of taphonomy, mainly dissolution; or an actual biotic 'event'. Statistical analyses showed that although sample size did have some effect, it was not the sole reason for the increase in ostracod numbers. Dissolution had an expected affect on the percentage of juveniles but no correlations were found with other sample characteristics. Sedimentation rate was investigated but this also proved unrelated. Therefore, it is suggested that the increase in total ostracod valves and diversity which occurs between 900 and 600 ka was in fact a natural, biotic 'event'. This preliminary evidence suggests that an oceanographic event that has negatively impacted on the foraminifers has had the reverse affect on the ostracod assemblage, in the sense that both population size and diversity increase during that time.

Sexual dimorphism is exhibited throughout various organisms and takes an extensive variety of forms, although rarely does it take the form of eye-reduction within a population. This poses the question of whether this dimorphism arises due to sexual selection in order to increase reproductive success or rather from ecological selection as a result of sex-specific niche partitioning. In this case, we study an organism that displays a drastic form of sexual dimorphism, in that the males have large complex lateral eyes whereas the females lack eyes and merely have rudimentary lateral eye-spots. Euphilomedes carcharodonta is a small crustacean native to the coast of California. Studies on the dimorphism between these individuals have mainly been related to genetics and embryology. However, minimal studies have been performed questioning the evolutionary driving force behind their dimorphism. One study suggests that this driving force is ecological selection, more specifically, predator evasion as a result of the reproductive role hypothesis. In this study, we focus on sexual selection as a driving force and ask whether the eyes play a role in the ability of males to find mates. We blinded and mock-blinded male individuals, placed them in tank settings with females in the absence of predators, and observed their behavior. If sexual selection is involved in the eye development of males, we expected to see changes in male and/or female behavior in response to obstructing the vision of the males. We did not see changes in behavior of the males or the females. This leads us to conclude that the development of complex eyes in male Euphilomedes carcharodonta arises from ecological selection rather than sexual selection. With the previous work on predator evasion, these experiments and findings are the first to experimentally test the reproductive role hypothesis and are the beginning steps for further behavioral studies of these organisms and in discovering more about the unique dimorphism exhibited by Euphilomedes carcharodonta.

The Sulaiy Formation, which is the oldest unit in the Lower Cretaceous succession, is conformably overlain by the Yamama Formation and it is a challenge to identify the precise age of the two formations using foraminifera and other microfossil assemblages. In the eastern side of Saudi Arabia, the Sulaiy Formation and the base of Yamama Formation are poorly studied. The main objectives of this study is to enhance the understanding of the Sulaiy Formation sequence stratigraphical correlation, regional lateral variations and palaeoenvironmental investigation. Lithological and semi-quantitative micropalaeontological analysis of 1277 thin sections taken from core samples from nine cored wells providing a geographically representative distribution from the Saudi Arabian Gulf. These cores intersected the base of the Yamama Formation and the Sulaiy Formation in the total thickness of cored wells of 843.23 meters (2766.5 feet). On the evidence provided by the foraminifera, the Sulaiy Formation is considered to represent the Berriasian to the lowermost Valanginian. The investigation of the micropalaeontology has provided considerable insights into the biocomponents of Sulaiy and the base of Yamama formations in order to identify their biofacies. These microfossils include rotalid foraminifera, miliolid foraminifera, agglutinated foraminifera, calcareous algae, calcispheres, stromatoporoids, sponge spicules, problematica (e.g. Lithocodium aggregatum), molluscs, corals, echinoderms and ostracods. Systematics of planktic and benthic foraminifera is accomplished using the foraminiferal classification by Loeblich and Tappan (1988) as the main source. The assemblage contains foraminifera that recorded for the first time in the Sulaiy Formation. Other microfossils were identified and recorded to help in the identification of the sedimentary environments. The investigation of the micropalaeontology and the lithofacies analysis have provided evidence the identification of the various lithofacies. About twenty four microfacies were identified on the basis of their bio−component and non-skeletal grains. The lithofacies and the bio−component results have provided the evidence of the sedimentary palaeoenvironmental model namely the Arabian Rimmed Carbonate Platform. This palaeoenvironmental depositional model is characterised by two different platform regimes. They are the Platform Interior and the Platform Exterior each of which have unique sedimentary lithofacies zones that produce different types of lithofacies. Each lithofacies is characterised by special depositional conditions and palaeobathymetry that interact with sea level changes and the accommodation space. The important palaeoenvironments are intertidal, restricted lagoon (subtidal), open marine, deeper open marine, inner shoal, shoal and platform margin. Generating, and testing, a depositional model as a part of formulating a sequence stratigraphical interpretation of a region is a key to understanding its geological development and – ultimately – reservoir potential. The micropalaeontology and sedimentology of the Sulaiy Formation in the subsurface have indicated a succession of clearly defined shallowing−upwards depositional cycles. These typically commence with a deep marine biofacies with wackestones and packstones, capped with a mudstone-wackestone maximum flooding zone and an upper unit of packstone to grainstones containing shallow marine biofacies. The upper part of the Sulaiy Formation is highstand-dominated with common grainstones that host the Lower Ratawi reservoir which is capped by karst that defines the sequence boundary. This karst is identified by its abundant moldic porosity that enhanced the the reservoir quality by increasing its porosities into greater values. Integration of the sedimentology and micropalaeontology has yielded a succession of shoaling−upwards depositional cycles, considered to be 4th order sequences, that are superimposed on a large scale 3rd order system tract shallowing−upwards, highstand-associated sequence of the Sulaiy Formation. The Lower Ratawi Reservoir is located within the latest high-stand portion of a third-order Sulaiy Formation sequence. The reservoir consists of a succession of several sequences, each of which is sub-divided into a lower transgressive systems tract separated from the upper highstand systems tract by a maximum flooding surface (MFS/Z). The last of these depositional cycles terminates in beds of porous and permeable ooid, or ooidal-peloidal, grainstone. The reservoir is sealed by the finer-grained sediments of the Yamama Formation.