Pollen analysis of organic sediments from vleis in upland areas of the Central and marginal Karoo has provided a vegetation history spanning the late Quaternary in the Winterberg, Sneeuberg, Nuweveldberg and Cederberg Mountains. Fossil pollen studies indicate moist conditions at the beginning of the Holocene, followed by a drier period. The second half of the Holocene (4 600 BP) is characterised by moister conditions, becoming drier towards the present and resulting in a decline in grasslands and an increase in Karoo-bushes. In the western Cape, the late Pleistocene (15 000 to 10 000 BP) was characterised by conditions moister than the present. This was followed by a drier period which ameliorated in the late Holocene. Although climatic fluctuations occurred, fynbos has been maintained in situ throughout the sedimentation period (14 600 BP), highlighting the dominant role of the substrate and secondary importance of climate. Climatic changes caused a shuffling of plant communities within the Fynbos Biome - this long uninterrupted history may be one reason for the high species diversity of fynbos. San hunter-gatherers, who occupied southern Africa prior to 4 000 BP, had a negligible impact on the vegetation. Khoi herders, who were first documented in the fossil record about 1 800 BP, had a slight impact on the vegetation, particularly by altering the fire regime. However, the arrival of European Trekboers some four hundred years ago had a significant effect on the environment. Although the general climatic trend in the Karoo is one of a drier phase, the decline in grasses and eastward movement of xeric Karroid elements has been accelerated due to mismanagement and sedentary farming techniques. An extensive contemporary pollen rain study has been undertaken to examine the representivity of these fossil pollen data. The investigation shows that contemporary pollen rain is a good reflection of the vegetation communities in the Karoo and Cederberg. Multiple discriminant analysis compares fossil pollen assemblages with contemporary pollen spectra and proved useful for determining whether modern analogues exist for the fossil pollen assemblages. TWISA confirmed the zones derived from discriminant analysis. Environmental changes have occurred in the Karoo during the late Quaternary, resulting in fluctuations and changes in vegetation patterns which have been accelerated in the recent past by human activity. Bibliography: pages 293-318.

Bibliography: pages 171-197. / Avian remains in coastal archaeological samples from Eland's Bay Cave, Die Kelders Cave 1 and Nelson Bay Cave in the Cape Province, South Africa, cover the periods between 80 000 and 40 000 B.P. and 18 000 and 300 B.P. Results of modern comparative surveys indicate that beached birds provide a predictable food supply. Kolmogorov-Smirnov two-sample nonparametric tests confirmed the close resemblance between the relative proportions of seabirds in archaeological and beached assemblages and earlier assumptions that the composition of seabird samples in archaeological sites could not otherwise have been achieved. It is shown that this simple but effective practice has a history going well into the Middle Stone Age. Kolmogorov-Smirnov tests for differences between the relative proportions of skeletal elements of Cape cormorants preserved in archaeological and modern jackal accumulations provided a useful means of drawing attention to possible activity of jackals and/or domesticated dogs. Recognition that diagenesis in some earlier samples may mimic the characteristics of modern jackal samples has established the need to extend the comparison of skeletal elements to additional species and to study the relative durability of avian skeletal elements. Similar comparison with the proportions of modern mammalian and avian predator prey species and size (mass) categories provided no indication that black or martial eagles might have contributed to the samples. Similarly, present knowledge of Cape eagle owls argues against their being likely inhabitants of caves suitable for occupation by people. It is concluded that people were the primary accumulators of the assemblages studied and that the role of small food items in prehistoric subsistence can be addressed with greater confidence. Correspondence analysis was used to determine the existence of seasonality in the modern beached seabird samples. The profiles of the archaeological samples are plotted in relation to months in which they were most likely to have been collected. Seasonal evidence from species not subjected to the correspondence analysis supported these results. The results obtained closely supported the hypothesis for seasonal exploitation of the coast. It was also possible to indicate that visits were probably of short duration and that their timing varied. Exploitation of seabirds did not coincide with the period of maximum availability of beached birds. Comparison of the avian evidence with that from seals, Cape dune mole rats and steenbok/grysbok suggested that small food items comprised part of a seasonal strategy that made maximum use of a range of seasonal resources. Evidence for significant local environmental change in addition to, and in support of, existing information has been obtained. Fluctuations in marine, freshwater and terrestrial birds at Eland's Bay Cave have been related to evidence for changes in terminal Pleistocene and Holocene sea levels and the position of the coast, and in the morphology of Verlorenvlei. At Die Kelders Cave 1 between 80 000 and 40 000 B.P., previously drier conditions were ameliorating and mixed scrub and grass and freshwater existed on the coastal foreland in the vicinity of the cave. Fluctuations in frequencies of seabirds indicate that the sea level rose slightly and then receded during the period of deposition. At Nelson Bay Cave samples indicate the approach of the coast after the Last Glacial Maximum, the disappearance of grassland and its replacement by scrub and bush as significant elements of the vegetation. Freshwater birds did not respond as expected, however, indicating that their interpretation at Nelson Bay Cave is complex and not consistent with evidence for wetter or drier conditions. A possible link has been shown to exist between fluctuations of albatrosses, petrels and shearwaters at Nelson Bay Cave and the intensity of wind patterns which are related to oceanic and atmospheric circulation. Further investigation should establish whether seabirds would provide an index of climatic conditions without support from other sources.

Palaeoenvironmental reconstruction of the late Pleistocene and Holocene geological periods for central and southern Africa has been hampered by the erratic distribution of suitable sites, incomplete and inaccurately dated sequences and the limited nature of published data. One geomorphological feature which has supplied valuable evidence for fluctuations in past environmental conditions, is the vlei or dambo. The type-site of these waterlogged features is in south central Africa, but similar features have been described on other continents. The clastic and organic sediments contained within these features are affected by, and therefore reflect to some degree, the environment under which they were formed. The characteristics of the sediments supply information as to their transport and mechanisms of deposition. From these processes, the environmental conditions at the time of vlei formation can be inferred. The environmental history of the Eastern Cape region has been considerably neglected, and is far less well understood than other countries such as Malawi, Zambia and Zimbabwe. A study site in the Winterberg Range (Eastern Cape) was selected which permitted the comparison of two vleis, the objective being to establish an accurate late Pleistocene sediment chronology for the entire plateau area. Radiocarbon dates from organic layers indicate that these sediments span the last 12 000 years BP, suggesting that organic accumulation at this site began at roughly the same time as at sites further afield. The vlei sediments are analysed in terms of their morphology, particle slze distribution, and other physical and chemical characteristics. These data facilitate the construction of detailed stratigraphic diagrams and a chronological summary of sediment accumulation, from which the period and governing processes of vlei development under changing environments may be described. It is found that the Winterberg vleis contain sediments which respond to changes in the prevailing environment. This makes these sediments useful indices from which to trace such changes during the late Pleistocene and Holocene times. These features are found to be similar in many respects to those described elsewhere in Southern Africa. The study attempts to provide greater understanding of contemporary vlei processes and emphasises the necessity of their preservation, as finite and valuable resources, by future generations