The Main Karoo Basin of South Africa is renowned for its exceptional palaeontological record and while its vertebrate fossils have been extensively researched, Karoo floras have received considerably less attention. Poor yields of palynomorphs from the Beaufort and “Stormberg” groups have complicated the task of erecting a comprehensive palynozonation scheme for the Karoo Supergroup. For this study, 65 palynologically productive samples from the Dwyka, Ecca, Beaufort and “Stormberg” groups allowed for systematic descriptions of all palynomorphs, as well as the ranges of the different taxa through the entire Karoo stratigraphic succession. Taxa with restricted ranges are useful for biostratigraphic correlation and these palynomorphs were used to delineate microfloral zones for the Karoo basin. The Dwyka Group contains high numbers of acritarchs and is generally low in species diversity. Useful biostratigraphic taxa for the Ecca Group include Cannanoropollis, Hamiapollenites, Platysaccus and Striatopodocarpites. Aratrisporites is a marker for the Latest Permian / Early Triassic Beaufort Group, while Cyathidites, Dictyophyllidites, Equisetosporites and Uvaesporites are indicators of the Late Triassic / Early Jurassic “Stormberg” Group. Palynostratigraphic zones correlate largely with the Karoo vertebrate biozones and severe and sudden extinction events are recognised among Karoo palynomorphs in the upper Tapinocephalus and Dicynodon assemblage zones. The first comprehensive palynological biozonation scheme for the Main Karoo Basin is proposed and the study provides a broad overview of Gondwanan Carboniferous - Jurassic floras. This study demonstrates that palynology is useful in correlating age equivalent lithostratigraphic units in the proximal and distal sectors of the Karoo Basin. Microfloras from previous South African studies are integrated within the proposed palynostratigraphic scheme, and palynological signatures of the various Karoo formations are shown to be consistent. Despite the constraints of floral provincialism, South African microfloras can be correlated to selected Gondwanan biozonations from Australia, Africa, Antarctica, New Zealand and South America. Future studies should focus on sampling more intensively over smaller stratigraphic intervals, which will assist in the correlation of time equivalent lithostratigraphic units in the different sectors of the basin, thus aiding in refinement of basin development models. Key words: palynology, Karoo, vertebrate biozones, stratigraphy, Gondwana

A dissertation submitted to the Faculty of Science, University of the Witwatersrand, Johannesburg, in fulfilment of the requirements for the Degree of Master of Science. Johannesburg, 2016. / Pareiasaurs were abundant, large, herbivorous parareptiles of the middle and late Permian which had a global distribution. The most basal pareiasaurs are found only in the middle Permian of South Africa, suggesting a Gondwanan origin for the group. Despite their relative abundance, most pareiasaurs are poorly known, especially the large middle Permian South African taxa that went extinct at the end of the Tapinocephalus Assemblage Zone. Historic taxonomic confusion was reduced with studies by Lee (1995, 1997a) that addressed the alpha-taxonomy of all pareiasaurs. He reduced the middle Permian South African pareiasaurs from 11 to four species: Bradysaurus baini, B. seeleyi, Embrithosaurus schwarzi and Nochelesaurus alexanderi. However this revision did not include detailed anatomical descriptions of any of the middle Permian South African Tapinocephalus Assemblage Zone taxa. The first detailed cranial description of Embrithosaurus schwarzi is presented in this contribution. Within the middle Permian pareiasaurians, Embrithosaurus has unique wide, short and stubby teeth with nine marginal cusps arranged more regularly. A cranial taxonomic reassessment of all middle Permian pareiasaurs has confirmed the validity of the four taxa identified by Lee and produced updated cranial diagnoses for each species. Diagnostic features noted by Lee and retained include an autapomorphic large distinctive maxillary boss on a strongly buckled or bent maxilla for B. baini, distinctive horizontally flat and pointed bosses on the posterior margin of the quadratojugal for Nochelesaurus and the smallest cheek flanges for B. seeleyi. Using the updated diagnoses, re-identification of 39 specimens out of 108 studied has produced updated biostratigraphic ranges for the middle Permian species. B. seeleyi is the first to make an appearance, in the middle Koornplaats Member of the Abrahamskraal Formation. B. baini first appears in the upper iii Koornplaats Member, Nochelesaurus in the Swaerskraal Member, and Embrithosaurus in the lower Moordenaars Member of the Abrahamskraal Formation. All taxa disappear by the top of the Karelskraal Member of the Abrahamskraal Formation, confirming that all the middle Permian pareiasaur species went extinct at the end of the Tapinocephalus Assemblage Zone. / MT2017

A study of the Abrahamskraal Formation in the area around Merweville, in the southwestern corner of the Karoo Basin has revealed the presence of traceable lithological units with lateral continuity throughout the study area. The stratigraphic section measured in this part of the basin matches the section measured by Jordaan, (1990) south of Leeu Gamka, with a basal arenaceous unit overlain by a predominantly argillaceous succession. The thickness of the Abrahamskraal Formation in this part of the Karoo Basin in 2565m, charactersized by a braided depositional environment in the lower 2075m and a meandering depositional environment in the upper 490m. Biostratigraphically the succession comprises a basal Eodicynodon Assemblage Zone which constitutes the lower 1104m and this is overlain by a 1461m thick Tapinocephalus Assemblage Zone whose upper limit is 21m below the Poortjie Member of the Teekloof Formation. This study has also corroborated the work by earlier authors who proposed a northeasterly palaeoflow direction as well as contributing to the global correlation of the Middle Permian terrestrial tetrapod faunas where the Eodicynodon Assemblage Zone correlates with the fauna from the Russian Ocher & Ischeevo; fauna of China’s Xidagou Formation and Rio da Rosto fauna of Brazil while the Tapinocephalus Assemblage Zone fauna corrletaes with fauna from Mezen and Ischeevo in Russia, Posto Queimado fauna in Brazil and those from the Madumabisa strata of Zimbabwe.