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Zooarchaeologists are beginning to produce more data on age profiles and sex ratios in archaeological faunal assemblages, but often lack the ecological basis to interpret these data. This thesis reviews the ethological literature on four main prey species found in southwest Asia faunal assemblages during the Late Pleistocene and Early Holocene: gazelle (Gazella sp.), Fallow deer (Dama dama mesopotamica), wild goat (Capra ibex) and wild sheep (Ovis sp.). This ethological review is used to develop models which predict the age and sex composition of archaeological faunal assemblages that were produced during different seasons and by different hunting techniques. Finally, a review of the archaeological record from the Pleistocene/Holocene transition in southwest Asia puts the age and sex ratios from archaeofaunas into the context of economic intensification and domestication.

The systematics of two hundred and eighty-nine prosobranch gastropods from the early to middle Pleistocene Moin Formation of Costa Rica are treated. The paleoecology of most extant and some extinct species of these mollsucs is discussed, and a paleoecological reconstruction of the Punta Limon peninsula is made. The presence of recognizable communities on differing substrates within the ecological zonation of a fringing reef is shown, from the intertidal zone down to the inner shelf at the base of the reef system itself, where ultimate deposition of the Moin Formation occurred. The faunal composition of the Moin indicates the occurrence of species-rich faunas evolving during the Pleistocene interglacial times, each involving speciation events following the extinction events of intervening glacial pulses. Evidence is given for the presence of a western Caribbean subprovincial unit during the Pleistocene, which continues on to the present day / acase@tulane.edu

Distinguishing marine and freshwater animals in the fossil record is a long-standing problem in paleontology. The physiological tolerances of extinct animals usually are inferred from environmental indicators and/or on the physiology of nearest living relatives. These types of evidence are often ambiguous and may be confounded by factors such as post-mortem transport and polymorphism in the living relatives. A solution to this problem is to combine these types of data with analyses of the oxygen isotope compositions of the phosphate (δ¹⁸O(p)) and the carbon isotope compositions of the carbonate (δ¹³C(sc)) of teeth and bones, to determine whether the ingested water and diet, respectively, were fresh or marine. The power of this approach is illustrated here in a study of the early evolution of cetaceans (whales, dolphins and porpoises). Changes in δ¹⁸O(p) and δ¹³C(sc) of the teeth and bones of early cetaceans documented here indicate that fully marine cetaceans existed by the middle Eocene and that some species exploited both marine and freshwater environments. This isotopic approach requires the avoidance of isotopically altered specimens. For this reason, the second component of this work deals with criteria for recognizing isotopically altered fossils. In contrast to one recent study, I found a positive correlation between δ¹⁸O(p) and δ¹⁸O(sc) not only in mammals but also in fish and reptiles. This correlation can be used as a test of whether the original isotopic composition is preserved in fossil specimens. Another approach to this problem is to make analyses of samples taken along growth transect of a fossil tooth or bone. Growth-transect analyses could resolve whether within-species isotopic variation represents differences in preservation or ontogenetic shifts in diet or habitat. In support of this goal, a new method for the analysis of phosphate oxygen is presented. This new method differs from all previous methods in that it involves no chemical reaction, but rather high-temperature (>725°C) equilibrium oxygen isotope exchange between CO₂ and Ag₃PO₄ As the amount of CO₂ is controlled by the analyst, small phosphate samples may be analyzed, making this method potentially useful for growth-transect analyses.

The four elements of the lower hind foot (calcaneus, metatarsal, naviculo-cuboid, and tibia) were tested for use as age-independent proxies of body size in white-tailed deer using known aged specimens from Ft. Hood Texas. Statistical analysis indicates that the calcaneum and the tibia are good proxies of age-independent body size in white-tailed deer. In addition to expanding the list of elements that can be used for studies of age-independent body size, these elements can also be used to age faunal remains to an ordinal scale of juveniles and adults. This is useful for research regarding prehistoric prey populations; as a single element can be used to determine prey body size and age simultaneously, which are the two variables used to assess changes in human subsistence practices via the archaeological remains of their prey.

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The human-abetted introduction of commensal species (i.e. those that opportunistically exploit the anthropogenic environment for food and shelter, e.g. rats, cockroaches etc.) to new areas has occurred throughout history. This has resulted in detrimental ecological changes worldwide but, from a viewpoint of human knowledge, a beneficial corollary of these translocations is that the species in question can be used as proxies to study the movement of the humans who transported them. I reconstruct colonisation histories of three widespread commensal mammalian species in the Western Indian Ocean, the black rat Rattus rattus, house mouse Mus musculus and Asian house shrew Suncus murinus, through phylogeographic studies (the geographic distribution of genetic lineages) of maternally-inherited mitochondrial markers, and zooarchaeological data. The DNA analyses are conducted on samples largely derived from museum specimens collected up to 110 years ago, and from archaeological bones (in the case of rats). I show considerable cryptic diversity in all three species, particularly in mice for which we find a potential major new lineage. Certain lineages within each species predominantly reveal long-distance translocations within the Indian Ocean, but high resolution geographic and genetic clustering is also evident, particularly in Asian house shrews. Phylogeographic structuring of the three species in East Africa and the southern Indian Ocean region (e.g. Madagascar, Reunion, etc.) indicate connections with Arabia, the Middle East, and India in the Islamic period from the first millennium AD, and later European connections during the Age of Exploration. Closer to the origins of the three species (the Indian subcontinent in all cases), range expansions in Eurasia and nearby islands relate to early to mid Holocene human populations, but also with signals of later secondary colonisations. Through ancient DNA studies I found genetic continuity between temporally separated populations of black rats suggesting population persistence, and high levels of diversity in Songo Mnara, a Swahili stonetown in Tanzania. Knowledge of the colonisation history and genetic diversity of an introduced species is essential to understand their resilience in novel landscapes, and to identify pathways of invasion and, by proxy, human trade and exchange networks that facilitated their dispersal. My research contributes significantly to that end for three socially, economically and ecologically important species that are well-established in the Indian Ocean region and beyond.

Archaeological and paleontological datasets are used in conservation to add time-depth to ecology. In central Texas several top carnivores including prehistoric Native American hunters have been extirpated or have had their historic ranges restricted, which has resulted in pest-level white-tailed deer (Odocoileus virginianus texana) populations in some areas. Predator extermination has dramatically reduced the average body size of members of the extant predator guild, and large carnivores most capable of hunting white-tailed deer are extirpated. Character release in the remaining “large” predatorsmesocarnivoresis a predicted outcome related to the adaptive vacuum at the top of the trophic hierarchy. Differences in body size of deer between prehistory and modernity are expected given that a lack of predation likely has increased intraspecific competition for forage among deer resulting in smaller body size today. In fact modern deer from settings without harvest pressure are significantly smaller than those from harvested areas and from prehistoric deer. From a natural history perspective, this research highlights potential evolutionary causes and effects of top-predator removal on deer populations and related components of biological communities in central Texas.