South Africa lags behind the rest of the world with regard to the availability of allometric biomass information. There is a complete lack of site specific allometric data for Pinus radiata in the southern Cape region, impeding investment in the renewable energy sector. This shortcoming was addressed by developing up-scalable, single tree biomass models. These models quantify the aboveground biomass of rotation age P. radiata grown in the southern Cape across a range of site conditions. The models use diameter at breast height (DBH) to predict the aboveground component biomass. A nutrient loss risk potential was assigned to each biomass component. Nineteen trees were destructively harvested using a full fresh weight sampling approach. Basic density was determined using a water displacement method, while Newton’s volume equation was used for stemwood volume determination. Log linear models were simultaneously regressed through seemingly unrelated regression (SUR) using the “Systemfit” R statistical package to force component additivity. A categorical variable was applied to the models, grouping the data into two Site Index (SI) based categories, namely “Low” SI and “Medium to High” SI, to account for inter-site variability. The corrected Akaike Information Criteria (AICc) and coefficient of determination (R2) was used to determine the goodness of fit of the models. The McElroy R2 for the SUR system was 0.95. Biomass models were developed that are able to predict various tree component masses at high levels of certainty within site and stand attribute ranges similar to this study. The importance of accurate, site specific wood basic density was demonstrated by its substantial weighting on stem and hence total biomass. Results showed that the stemwood mean basic density range was between 503 kg m-3 and 517 kg m-3 for the “Low” SI sites and 458 kg m-3 for the “Medium to High” SI sites. Site quality can have a major impact on the models, particularly on poorer sites where stemwood production is proportionally less than other tree components. Total aboveground biomass was estimated to range between 58.61 odt ha-1 and 70.85 odt ha-1 for “Low” SI sites, and 185.31 odt ha-1 to 266.58 odt ha-1 for “Medium to High” SI sites. Stemwood biomass accounted for 65% of the total aboveground biomass for “Low” SI sites and 70% for “Medium to High” SI sites.

In South Africa, the limited extent of indigenous forests accelerated the development of fast-growing introduced species plantations, on which the country is now heavily reliant for its fibre and timber products. However, the plantation forestry industry is challenged with limited freshwater resources, the need for sustainable management of introduced tree species in commercial forest production systems, and a diversity of plant ecological production factors such as soils that have to be manipulated for the purposes of increasing production capacities. Additionally, plantations are established in the limited high rainfall regions of the country and the industry’s water use has been regulated since 1972. Conversely, natural forests also provide valuable goods, however, their slow growth rates have restricted their development as commercial tree production systems. In this regard the forestry industry is continuously seeking to provide for the country’s timber and fibre needs while ensuring the provision of other ecosystem services from tree production systems. Recent developments in the forestry industry have therefore been focused on water use efficiency in current and alternative tree production systems. There is also widespread unsubstantiated belief that indigenous tree species are efficient users of water. Against this background this study sought to explore the single tree water use and water use efficiencies of introduced commercial plantation species (Pinus radiata) and that of important indigenous species (Ilex mitis, Ocotea bullata and Podocarpus latifolius) for timber and fibre production in the southern Cape region of South Africa. Single tree water use measurements were carried out for a year using the heat pulse velocity method. Growth measurements for utilisable stem wood were done on a quarterly basis for the specimen trees and growth increments over the year were determined. Single tree water use efficiency was a calculated as a function of grams of stem wood gained per litre of water transpired. Measurements of daily weather conditions and soil water content were concurrently taken during the year. The relationships between the variables that express daily climatic conditions, soil water content and daily volumes of transpired water were explored. Empirical models for the prediction of daily transpiration as a function of climate and soil water content were then developed using multiple linear regression analysis. Pinus radiata trees exhibited higher totals of volumetric transpiration than the indigenous species. The water use patterns of Pinus radiata showed higher peaks of maximum and minimum daily and seasonal water use while the indigenous species showed temperate patterns of water use throughout the year. Podocarpus latifolius attained the lowest water use efficiency in the year (0.49g/L) while Pinus radiata, Ocotea bullata and Ilex mitis had comparable water use efficiency values ranging between 1g/L to 2.50g/L. Significant positive correlations existed between climatic variables and daily sapflow volumes of the tree specimens. Daily total solar radiation showed the highest positive correlation with daily transpiration and the average value of the correlation coefficient for solar radiation and water use for all tree species was 0.70 (p < 0.001). Empirical models for predicting daily water use integrated the variables expressing plant available moisture and energy. The study enhances our understanding of single tree water use, water use efficiency, the drivers of transpiration and the applicability of such studies in developing rapid estimation techniques for water use in current and alternative South African tree production systems.

Thesis (MScFor)--Stellenbosch University, 2012. / ENGLISH ABSTRACT: The aim of this project was to determine the site effect- especially water availability - and the effect of cambial age on selected anatomical properties of Pinus radiata, in order to be able to predict possible changes in wood quality due to an expected change in climate. A second objective was to correlate ring and fibre properties, in order to determine, if ring properties could be used as a proxy to describe wood quality. The samples consisted of 12 trees, sampled at an age of 1 to 16 from six selected sites in the Western Cape, which ranged from water stressed to moist. Apart from the water availability all other external factors, such as elevation etc. were kept as equal as possible. Anatomical wood properties such as fibre length and fibre diameter, lumen diameter, cell wall thickness, ring width and earlywood/latewood ratio were determined and their change with cambial age and water availability was evaluated. Fibre length, fibre diameter and cell wall thickness increased with increasing cambial age, and ring width and earlywood/latewood ratio decreased with increasing cambial age. No significant correlations were found between any of the ring or fibre properties and water availability. Most of the fibre properties were significantly correlated with ring width and earlywood/latewood ratio when age was not considered as covariate, but showed no correlation when the age effect was excluded. / AFRIKAANSE OPSOMMING: Die doel van die projek was om die perseel se effek te bepaal, veral water beskikbaarheid, ten opsigte van die effek van kambium ouderdom en geselekteerde anatomiese eienskappe van Pinus radiata, om sodoende die moontike verandering in hout kwaliteit as gevolg van verwagte klimaatsverandering te voorspel. ‘n Tweede doelwit was om die ring en vesel eienskappe te korreleer en ook te bepaal of ring eienskappe gebruik kan word om hout kwaliteit te beskryf. Die monsters het bestaan uit bome van ses geselekteerde persele in die Wes Kaap en het gevarieer van ‘n water tekort na klam. Behalwe vir water beskikbaarheid is al die ander eksterne faktore, soos hoogte ens., konstant gehou waar moontlik. Anatomiese hout eienskappe soos vesel lengte en deursnee, sel deursnee, selwand dikte, ring wydte en E/L verhouding was bepaal asook die verandering met kambium ouderdom en water beskikbaarheid. Vesel lengte , vesel deursnee en selwand dikte het toegeneem met toename in kambium ouderdom, en ring wydte en E/L verhouding het afgeneem met toename in kambium ouderdom. Toename in water beskikbaarheid het gelei to afname in vesel lengte en selwand dikte, waar vesel deursnee, sel deursnee, E/L verhouding en ring wydte toegeneem het. Meeste van die vesel eienskappe het betekenisvol gekorrelleer met ring wydte en E/L verhouding wanneer ouderdom nie as ko-variant gebruik is nie.