Thesis (Ph. D.)--University of Washington, 1983. / Vita. Bibliography: leaves [138]-147.

Thesis (M.E.S.)--The Evergreen State College, 2007. / Title from title screen viewed (2/14/2008). Includes bibliographical references (leaves 58-61).

Thesis (Ph. D.)--University of Sydney, 1999. / Title from PDF title page (viewed on May 4, 2005). Includes bibliographical references.

No description available.

Thirteen badgers and 20 foxes were radio-tracked in the Wytham Estate, Oxfordshire, between 1981 and 1983. Thirteen badger and 10 fox groups were identified from radio-tracking and bait marking. Badger groups (mean size 1982: 4.45, 1983: 5.82) occupied contiguous territories (size: 22-75 ha) with boundaries marked by latrines. Seasonal variation in marking intensity and choice of marking sites presumably were responses to changing intrusion pressure. Fox groups (mean size: 2.6) occupied stable territories (size: 22- 104 ha) with little overlap. Faeces deposition by foxes facilitated territory marking. Earthworms (Lumbricus terrestris) dominated the diet of badgers (63 % estimated dry weight EDW, faeces), followed by cereals, fruits and other Invertebrates. Diet was highly variable between groups and seasons. For foxes, lagomorphs (20 % EDW) and earthworms (33 % EDW) were the most important prey, followed by scavenge and fruits. Variation in diet between groups and seasons was marked in lagomorphs but not earthworms. Multlvarlate analyses of habitat parameters revealed a low-dimensional 'resource space' that could be divided into conventional habitat categories. Censuses of prey species indicated that resource presence varied consistently between habitat categories. Key habitats occurred at fairly constant proportions in territories of both species) their dispersion partly determined the configuration of territory boundaries. The proportions of specific habitats per territory were correlated with the proportions of certain prey items in diets. space use by individuals was analysed by spatial autocorrelation methods, variation in space use by foxes was attributed to variation in resource dispersion. In contrast, individual badgers were similar in their use of space. Here, small-scale heterogeneity in intensity of use may reflect local earthworm availability, in one studied fox group, males and females differed in range use. Individuals in one studied badger group coordinated their use of space probably to minimize foraging interference. It is suggested that group living in Wytham badgers is a response to defending resources, and a model is proposed to explain how the spatial and social organisation of male and female badgers relate to the characteristics of the resources they require.

M.S.

This thesis provides an insight into the biology and ecology of foxes Vulpes vulpes from a coastal habitat and free roaming dogs Canis lupus familiaris from an Aboriginal community, both in SE NSW. In particular, the study examines the home range and movement patterns of both canids from a view point of population and disease control. Additional base line information was collected for these species on diet, parasites and population biology. A total of 14 foxes and 10 dogs were collared and radio tracked to determine the home range and movement patterns of the animals. The data collected provides information on the behaviour of foxes in coastal habitat and represents the first detailed biological study of freeroaming dogs in Australia. The results provide a better understanding of dog movements, particularly the differences in behaviour between wandering and sedentary dogs. Home range sizes varied greatly between the dog groups from 1 ha to 2451 ha. There was evidence that dog forays were influenced by prey availability, particularly medium sized macropods. Occasional sallies, size and shape of fox and dog home range is discussed in relation to the richness of habitat at Jervis Bay with comparisons being made with other studies from the literature. Foxes in Jervis Bay had home ranges between 60-270 ha which is comparable to foxes living in the forests of Germany and in coastal habitat in the Netherlands This study also provides data on the diet of foxes and dogs presenting quantitative and qualitative data on food preferences. Foxes tended to be opportunistic in food selection although critical weight range mammals ie. ringtail possums were favoured. The high prevalence of the parasite Spirometra spp. in foxes indicated that considerable numbers of amphibians and reptiles are consumed by this species. A comprehensive inventory of the parasites of the two canid species showed a high proportion of hookworm in foxes and dogs and Spirometra spp in foxes. These results are discussed in light of the potential health implications for local Aboriginal people. Analysis of fox scats revealed that they have the potential to spread the seeds of bitou bush. A more detailed pilot study revealed the potential impact of foxes on the environment through the spread of weed seeds. Germination trials recorded high viability of seeds that were eaten and deposited by foxes. The management implications of this behaviour are discussed in context to weed control. This thesis has provided new information on foxes and dogs in Australia. Discussion on these species in the thesis has been framed around the benefits of this information for population and disease control.

The red fox occurs across a very broad range of habitats, and displays great behavioural flexibility under different environmental conditions. In Australia, mounting concern over the impacts of foxes on livestock and native fauna has highlighted a need for more information on fox ecology under Australian conditions as a fundamental step towards developing more strategic means of managing foxes. This study explores ranging behaviour, dispersal, use of dens, activity rhythms, population dynamics and diet in the absence of management in productive agricultural land in the central tablelands of New South Wales. The study was conducted from June 1994 to June 1997 on private property near Murringo, NSW Australia (34°15� S, 148°30� E). The site was primarily sheep and cattle grazing land and had a history of no fox management. Rainfall was considerably below average for much of the study. A total of 83 foxes were trapped over 3931 trapnights, of which 50 were fitted with radio-collars (23 adult and 6 juvenile females, 12 adult and 9 juvenile males) and 26 released with eartags only (all juveniles: 10 females, 16 males). Thirty-three foxes were radio-tracked using fixed towers between March 1995 and December 1996, with between 11 and 28 foxes tracked at any time. Mean home range size was 446.1 ha ± 69.8 se using 95% Minimum Convex Polygons (MCP), and 276.4 ha ± 36.3 se using 95% kernel utilisation distributions. Male home ranges defined by MCP were significantly larger than female ranges, but no significant difference was found using 95% kernels. Core ranges were estimated to be 133.4 ha ± 23.7 se using 50% MCP and 59.8 ha ± 6.1 se using 95% kernels, with no significant difference between sexes. No significant differences were found between range sizes of adults and juveniles or between years or seasons. While most home ranges were steady for the duration of the study, some foxes were observed to shift range location and 4 foxes displayed nomadic behaviour for at least some of the study. There was a high incidence of overlapping home ranges, most commonly between females or males and females but occasionally between males, but core areas were usually separate. Fully overlapping core areas were observed in 1995 but not in 1996. Juvenile foxes were significantly more likely to disperse than adults, and usually travelled further (juveniles 61.1 km 31.6 ± se; adults 5.9 km 1.1 ± se). Males and females were equally likely to disperse, and there was no significant difference in the distance travelled. The furthest distances were 285 km and 140 km, but mean distance of dispersal excluding these animals was 12.3 km ± 4.3 se (n = 13). Thorough surveys across a 16.4 km² area located 200 dens, with 68 of these active in 1995 and 96 active in 1996. Density of breeding foxes was estimated to be 0.55 and 0.52 adult foxes/km² in 1995 and 1996 respectively based on natal den counts. Density estimates based on active den counts, which include non-breeding foxes, were 0.91and 1.30 foxes/km² in 1995 and 1996 respectively. These estimates appear lower than other studies in similar habitats but this is likely due to using a half home range boundary strip around the surveyed area in the present study. Application of mark-recapture analysis found very high �recapture� rates of dens and gave a similar estimate of the total number of dens to that observed directly. Natal dens were regularly distributed across the study area, whereas active dens tended to be in clusters. There was a high turnover of which dens were used each year, but the total number of natal dens was similar across years (16 in 1995 and 17 in 1996). Natal dens were more likely to be used on repeat occasions than other dens, but not necessarily by the same vixen. Litter size based on sightings of emergent cubs was 2.8. Foxes were predominantly nocturnal, with a major peak in activity about an hour after sunset. A new method of analysing activity rhythm data using Fourier series to mathematically describe animal movements was developed, that allowed systematic identification of the cyclical components underlying overall movement patterns. General fox behaviour could be clearly described by a 24-hour and a 12-hour cyclical component when corrected for variation in daylength. The rising and setting of the sun appeared to be a major trigger underlying movement patterns. Seasonal and sex differences were observed in patterns of activity. The annual rate of increase of the fox population was found to vary around a mean of zero between June 1994 and June 1997. A major drop in fox numbers as estimated by spotlight counts occurred in the second half of 1995, but numbers recovered by the end of 1996. Kaplan-Meier analysis of radio-tagged foxes found annual adult survival was generally very high (0.56-0.96) with lowest survival between July and October. Causes of mortality were human-related outside the site and apparently of natural causes within the site. However foxes dying of natural causes outside the site were unlikely to be found. There was no overall movement of foxes into or out of the site. Immigration was detected following the drop in fox numbers in late 1995, but there was no evidence of immigration prior to this period although emigration occurred. A sensitivity analysis was conducted on the effects of a small change in life history parameters on finite rate of increase using published data as well as adult mortality data from the present study. The two most influential life-history parameters were adult and juvenile survival, while changes in fecundity and age at first reproduction had much less impact on finite rate of increase. In terms of management, in which fertility control is being considered as an alternative to lethal control, this implies that a small change in fecundity may cause less change in the rate of increase of foxes than lethal control. Foxes were culled in June 1997 on completion of the study. Estimated density using a Petersen estimate was 2.4-5.3 foxes/km² and index-manipulation-index was 1.4-3.2 foxes/km². The different methods used to cull foxes appeared to target different age groups within the population, and were generally biased in favour of younger foxes. Success at killing animals was low, leading to large standard errors in the population estimates. Stomachs of foxes shot in the Orange district were found to contain predominantly rabbit and carrion, with invertebrates present when abundant. These findings were not strictly representative of the diet of foxes in the study area, where rabbits were scarce. Foxes scavenged heavily on lamb carcasses within the study site. The quantity of fresh lamb carrion removed from a lambing paddock in winter 1996 was estimated to support 13-24 foxes, with available fresh lamb theoretically able to support 240-440 foxes. Density based on removal of fresh carcasses was estimated to be 0.83-1.5 foxes/km².

Host behavior, age, sex, diet, and condition, as well as variation in parasite specificity, drive variation in parasite infection, and ultimately determine the host parasite community. The objectives of this thesis were to 1) examine intraspecific variation in arctic fox parasites, 2) determine relationships between diet and parasites in sympatric arctic and red fox, and 3) compare wolf parasites and diet. Male arctic fox had more cestodes than females and juveniles had more nematodes than adults, likely due to diet and exposure. Red fox carried fewer parasites than arctic fox, likely due to diet, evolved resistance behaviors and higher immune investment, but diet affected cestode abundance in both species. Wolves that ate more white-tailed deer had more cestodes, suggesting increasing deer populations could enhance parasite transmission to moose. However, body condition was unaffected by parasites, suggesting northern canids may have not reached a threshold of infection.

Host behavior, age, sex, diet, and condition, as well as variation in parasite specificity, drive variation in parasite infection, and ultimately determine the host parasite community. The objectives of this thesis were to 1) examine intraspecific variation in arctic fox parasites, 2) determine relationships between diet and parasites in sympatric arctic and red fox, and 3) compare wolf parasites and diet. Male arctic fox had more cestodes than females and juveniles had more nematodes than adults, likely due to diet and exposure. Red fox carried fewer parasites than arctic fox, likely due to diet, evolved resistance behaviors and higher immune investment, but diet affected cestode abundance in both species. Wolves that ate more white-tailed deer had more cestodes, suggesting increasing deer populations could enhance parasite transmission to moose. However, body condition was unaffected by parasites, suggesting northern canids may have not reached a threshold of infection.