Seasonal trends in forage production and environmental parameters for five plant community types within a northeastern Oregon riparian zone were described and modeled using correlation and path analysis. Wet meadows produced the greatest amount of herbage biomass, followed by moist bluegrass meadows, gravel bars, forests and dry bluegrass meadows. Trends in soil moisture generally increased and then declined from spring to fall. Depth to the water table declined and then increased. Soil temperatures steadily increased. Variables driving seasonal forage production varied by community type. Soil moisture was most important in dry bluegrass meadows and least important in wet meadows. Depth to the water table was most important in wet meadows and least important in dry bluegrass meadows. The amount of herbage production which had already occurred was also an important variable in describing biomass production. Streamflow levels and the amount of production having occurred were driving variables in the gravel bar communities. Preference for grazing different riparian vegetation community types and forage intake by cattle was monitored over a three-week grazing period occuring at the end of summer. Concurrent to preference and intake, vegetative and nutritional characteristics of the forage available for grazing were monitored and relationships between these variables and both community preference and intake described through correlation and path analyses. Grazing cattle initially favored communities with highly digestibile forage, hence communities dominated by Kentucky bluegrass were most preferred. Late in the grazing period community preference was best associated with community abundance, indicating that cattle were grazing communities in proportion to their abundance in the pasture. Intake levels were greater during the first year of the study than the second (2.15 versus 1.81 percent of body weight). Daily grazing time declined as livestock neared the end of the grazing period. Intake was correlated with in vitro dry matter digestibility and the amount of time spent grazing, but poorly related to the amount of forage available. The indirect effect of the amount of forage available on intake was greater than the direct effect and functioned through increases in grazing time as a result of increased availability of highly digestible forage. / Graduation date: 1992

Because riparian canopy controls most energy inputs to stream ecosystems, it directly affects the structure of aquatic food webs and the ecological processes that govern interactions among trophic levels. This study addresses the interdependence among riparian canopy, benthic community structure, and the carrying capacity of high desert streams for salmonid fishes. In streams in the lower John Day River Basin in eastern Oregon, algal, invertebrate, and fish communities were compared in reaches with varying densities of riparian canopy. Water temperatures varied with the density and upstream extent of canopy. Densely canopied sites were cool, while sites with high irradiances had temperatures exceeding the upper lethal limit for salmonids. Periphyton and grazer biomasses were greater in well-lighted sites, but 90% of grazer biomass consisted of Dicosmoecus gilvipes, a large caddisfly inedible by juvenile trout. Warmer water increased metabolic demands for salmonids, while the overwhelming dominance of Dicosmoecus in open sites shifted energy flow away from trout and shrunk their food base. High water temperatures, however, provided suitable habitat for many warmwater fishes which would otherwise not enter tributaries of this size. At higher elevation study sites in Camp Creek, light levels were higher and less variable than at the lower sites. Periphyton and invertebrate abundances were not correlated with irradiance. Rather, periphyton was maintained at low levels by grazers, particularly Dicosmoecus and snails. Manipulations of fish densities in enclosures showed that trout and dace had no negative impacts on numbers of invertebrate prey, and that grazers played a larger role in regulating lower trophic levels than did fish. Dicosmoecus acted as a keystone species in the benthic food web of Camp Creek by simultaneously influencing the trophic level both below and above its own. When irradiance was experimentally reduced under artificial canopies, periphyton standing crops were not different from those in open control pools after 4 wks. However, grazers were more abundant in open pools. The cropping of periphyton to uniform levels in both sunlight and shade indicated that mobile grazers targeted sites of varying productivities. Comparisons between benthic communities in Camp Creek and in a densely canopied reference stream suggested that benthic community structure shifted to accommodate changes in energy resources that occur when canopy density is altered. / Graduation date: 1998

Pacific salmon populations in Johnson Creek, Portland, Oregon have been adversely impacted by urbanization, and by residential and agricultural land development. f Ecological impacts include loss or depletion of riparian vegetation features which directly influence stream and associated salmonid ecology. This research examines the nearstream riparian zone's contribution to instream habitat complexity for anadromous salmonids in Johnson Creek. Visual surveys were conducted on over half the stream length. Five features were assessed to determine the extent of riparian influenced stream habitat including, overhead enclosure, overhanging vegetation, undercut banks, and large and small woody debris and root wads. The stream survey showed that areas of riparian-influenced habitat are spatially intermittent and present in areas of the least stream disturbance. Suitable riparian habitat is limited to locations where there is minimal riparian disturbance, property management, and channelization. I Although salmonid populations have been reduced, insufficient riparian influenced salmonid habitat features are not the primary limiting factor on salmon populations. Other factors such as pollution, sedimentation, hatchery fish introduction, low flows, inadequate food supplies, high stream temperatures, repress wild salmon populations.

Two trials were conducted to evaluate changes in the quantity, quality, and moisture of available forage in the pasture, and shrub utilization by cattle during a 30-d late summer grazing period (Trial 1) and the effect of cow age (experience) on grazing distribution and diet composition (Trial 2) in mountain riparian areas. In the trial 1, a pasture (44.7 ha) in the Catherine Creek site at OSU���s Hall Ranch in northeast of Oregon was grazed with 30 yearlings and 30 mature cow/calf pairs from early August to early September in 2001, and from late July to late August in 2002. Sampling dates were d 0, d 10, d 20, and d 30 of the grazing period. The forage availability before grazing was 1058 kg/ha and declined to 323 kg/ha at the end of the grazing period (P<0.10). Grasses dominated the pasture, followed by forbs, grasslikes, and shrubs. Kentucky bluegrass was the most prevalent forage species followed by timothy, sedges, and common snowberry. The highest percent disappearances of forage species was (83.7-92.7%) observed with quackgrass, western fescue, California brome, redtop, and heartleaf arnica, though their initial contributions to the available forage were less than 5%. High levels of shrub utilization were observed from d 20 through the end of the grazing period (45% for willow and 59% for alder). Forbs and shrubs did not vary in moisture content between the 10 d intervals and across the years averaging 59% and 61%, respectively (P>0.10). In contrast, the moisture content of grasses were over 50% at the beginning of the grazing period but declined dramatically to 34% from d 10 to d 20. Likewise, forbs and shrubs were higher (P<0.05) than grasses in CP (11, 14, and 6%, respectively) and IVDMD (58, 49, and 42% respectively). In summary, our results suggest that cattle grazing late summer riparian pastures will switch to intensive shrub utilization when grasses decline in quality and quantity, and forbs decline in quantity. In the trial 2, thirty first calf heifers, and thirty mature cows were randomly assigned to four pastures (15 head per pasture, average 21.5 ha) in the Milk Creek site of Hall Ranch from late July to early September of 2000 and 2001. Botanical composition of diets was determined by analyzing the feces from 10 animals (5 per pasture) in each treatment during the fourth week of the trial using the microhistological procedure. Correction factors were calculated for the 22 major plant species. First calf heifers had higher portions of grasses (75% versus 71%; P<0.05), but lower portions of shrubs and trees (9% versus 13%; P<0.10) as compared to mature cow diets, respectively. On an individual species basis, ponderosa pine consumption was a major contributor with mature cows consuming greater quantities (P<0.10) than first calf heifers. In summary, mature cows seem to have selected diet less in the amount of grasses and more in the amount of shrubs and trees as compared to younger cows. / Graduation date: 2004

Twelve stream segments in the Silvies River drainage system were classified in the fall of 1993 and again in 1994, using a morphological classification of natural rivers (Rosgen 1994). Bankfull flow of stream channels is the key feature of this system. Measurement of bankfull is used in the calculation of entrenchment ratio and width/depth ratio. Analysis of the differences of the averages between years in bankfull measurements showed that despite being consistently repeated at the same locations along the channel, the measurements were found to be different (p=0.000163 for entrenchment ratios and p = 0.0208 for width/depth ratios). Bankfull was found to be a non-repeatable field measure, therefore, a poor benchmark for a classification scheme. Historical information collected for the study area indicated a history of domestic livestock use that dates back as far as the mid-nineteenth century. Although grazed by livestock for several years, settlement of the Bear and Silvies Valleys occurred mostly around the turn of the twentieth century after the stockgrower's homestead acts were passed. Ownership today is dominated by large (relative to the homestead days) ranches. / Graduation date: 1996

Riparian zones provide habitat for breeding birds in the semiarid western United States; however, there are few data available that address the effects of livestock grazing strategies on riparian habitats and avian communities. Documenting avian community composition in different riparian vegetation communities and relating vegetation communities to livestock grazing strategies may identify management alternatives that are sustainable from a wildlife habitat perspective, and may permit constructive coalitions between agricultural industry and environmental groups. I compared diurnal breeding bird abundance, individual species abundance, and species richness, and vegetation composition and structure among 12 streamside riparian areas of Bear and Silvies valleys in eastern Oregon during 1993 and 1994. Bird and vegetation data were collected along four replicate transects within each of three riparian vegetation communities characterized by vegetation structure: herbaceous, discontinuous willow (Salix spp.), and continuous willow. These riparian vegetation communities were grazed under summer season-long, summer short-duration, and fall short-duration livestock grazing strategies, respectively, >5 years before the study. Differences in riparian vegetation among communities were primarily related to shrub structure by experimental design. The continuous willow community had more shrub cover overall (P<0.001) and within each 1-m height interval from 0-4 m (P<0.013) than the herbaceous and discontinuous willow communities. The herbaceous community had no shrub cover >1 m in height. Willows extended farther (P=0.031) from the steam edge in the continuous willow community than in the discontinuous willow community. No willows were detected in the herbaceous community. I detected 4,016 birds representing 56 species along the transects. Total bird abundance was greater (P<0.001) in the continuous willow community than in the herbaceous and discontinuous willow communities. Species richness was inconsistent (P=0.034) between years within communities; it was greatest (P<0.037) in the continuous and discontinuous willow communities in 1993, and greatest (P<0.003) in the continuous willow community in 1994. Total bird abundance and species richness in 1994 increased with willow volume (r��>0.707, P<0.001). Of 23 bird species with >20 individual detections over both years, 13 species were most abundant in one or two vegetation communities (P<0.088). Seven species (yellow warbler [Dendroica petechia], song sparrow [Melospiza melodia], willow flycatcher [Empidonax traillii], American robin [Turdus migratorius], common snipe [Gallinago gallinago], bobolink [Dolichonyx oryzivorus], and Vaux's swift [Chaetura vauxi]) were most abundant in the continuous willow community. Three species (savannah sparrow [Passerculus sandwichensis], black tern [Chlidonias niger], and American wigeon [Anas americana]) were most abundant in the herbaceous community. Willets (Catoptrophorus semipalmatus) were most abundant in the herbaceous and discontinuous willow communities whereas cinnamon teal (A. cyanoptera) were most abundant in the herbaceous and continuous willow communities. Red-winged blackbirds (Ageliaus phoeniceus) were inconsistent (P=0.032) between years within communities: they were most abundant in the continuous willow community in 1993, and most abundant in the continuous willow and herbaceous communities in 1994. Hydrophytic woody vegetation within semiarid rangeland environments increases structural complexity and is associated with avian abundance and diversity. Yellow warblers, willow flycatchers, and song sparrows, which depend on hydrophytic shrubs for nesting almost exclusively in the semiarid West, are especially threatened by the elimination or simplification of woody riparian vegetation. I suggest that riparian vegetation structure and composition, which is associated with avian abundance, species richness, riparian associate bird species, and landscape-level biological diversity, be maintained where possible. Seasonal light (<30% use) fall short-duration grazing seemed to be compatible with the maintenance of woody riparian vegetation whereas summer season-long and summer short-duration grazing is likely incompatible. / Graduation date: 1995

The purposes of this study were: 1) to review the land use history of Oak Creek watershed since European settlement. And 2) to consider the human-caused impacts and their effects associated with different land uses. The study area was classified into three land use patterns: residential, agricultural, and forested areas. Since the European settlement in the mid 1840's, the Oak Creek watershed has experienced changes of land use simultaneous with the development of the city of Corvallis and its vicinity. Associated with this, different impacts have occurred on riparian areas. By and large, these impacts were inherent to each land use. / Graduation date: 1993

Graduation date: 2002

I hypothesized that vegetation and physical environmental characteristics would differ between the upper and lower extents of the annually flooded riparian zone on the John Day River, and that relationships between species and environmental variables would display differences between these two zones. Vegetation, environmental variables, and relationships between them were assessed for the entire annually flooded riparian zone, and for the proposed upper and lower zones. Data were collected from 60 one-square-meter quadrats: 30 in each the upper and lower zones. Sites were randomly selected and located so that flood duration was roughly equal at all sites within each zone. 34 plant species were encountered: 25 in the upper zone, 27 in the lower zone. Wetland obligate and facultative wetland species groups and eight individual species accounted for statistically different percentages ofquadrat cover between zones. ANOSIM analysis identified two statistically distinct vegetation communities between the two zones. Soil texture averaged 75.85% sand and 20.81% fines. Sand ranged between 36.69% and 95.55%. Fines ranged between 2.54% and 58.84%. A horizon depths and fine soil particle concentrations were greater in the upper zone. Coarser soils with more sand and gravel dominated the lower zone. All enviromnental variables studied, except pH, were highly variable throughout the study area. ANOSIM analysis results suggest that the upper and lower zones have distinct, statistically different physical environments from each other. Regression analyses relating species quadrat cover to physical environmental variables were performed for the total, upper, and lower riparian zones. Numerous differences were identified between the upper and lower riparian zones that the riparian scale analyses did not represent accurately. There were ten instances in which the zone scale analyses identified a relationship in either the upper or lower zone, while the corresponding riparian scale analysis failed to identify any relationship. The results of this study indicate that vegetation and the physical environment are statistically different between the upper and lower zones on this river, and that relationships between a given plant species and environmental variable can vary between zones. Future research and management efforts should consider and address the potential for such between-zone variation.