Naturalists for many years have known that birds sing most at dawn and at dusk, but to which of many physical variables they are responding has not been completely determined. This problem has taken on importance in the field of wildlife management because calling rate has been used as an index of abundance for numerous game birds, namely the Pheasant, Phasianus colchicus( Kimball, 1949); the Mourning Dove, Zenaidura macroura (McClure, 1939); the Bobwhite Quail, Colinus virginianus (Bennitt, 1951; Rosene, 1957): and the Woodcock, Philohela minor (Pitelka, 1943). Of the possible variables which can influence calling in birds, light intensity has received the most attention. Haecker (1916), using a crude optical light meter, found a definite and high correlation between time of beginning morning song and light intensity. In 1924 he published findings with the same conclusion after measuring light intensity photometrically. Dorno (1924), in repeating part of Haecker's work, came to the conclusion that light was significant, but that the variation in the time of beginning morning song; and sunrise was due to differences in dispersal of light rays rather than actual light intensity. This dispersal was due, in his opinion, to latitude and season of the year. Since that time many workers have been concerned with the problem . The first American to deal with light intensity as the factor that governs time of morning calling was Craig (1926). He found a close correlation between early morning song of the Eastern Wood Pewee ( Contopus virens) and the curve of civil twilight. Walker (1928) found that the length of time before sunrise a bird sings is probably dependent on the total amount of light present, but that weather factors influence song to a great extent. In agreement with these findings were those of Lutz (1931) studying singing in the House Wren (Troglodytes musculus). He found that one individual began to sing at almost the same time on each of 24 mornings. If the morning was bright, song came earlier; if couldy, it came later, but time never varies more than 15 minutes. Wiens (1960) stated that the song of the Cardinal (Richmondena cardinalis) was closely associated with sunrise during April. He recorded calls beginning 10 minutes )plus or minus 3 minutes) before sunrise during this month. Eynon (1960), in compiling field notes of the late Aldo Leopold, showed that the first morning songs of several passerine and game birds are closely associated with very low light intensities that occur during the twilight before sunrise. Studies showing effect of light intensity on animal activity are not limited to birds. Alexander and Moore (1958), studying singing in two species of Cicada (insects), found that both are sensitive to changes in light intensity. Singing increased as the sun came up and decreased as the sun passed behind a cloud. They attempted to stimulate singing with taped song under cloudy conditions, but only one species responded.Steven (1959) showed that schools of fish occurring during daylight hours break up as light falls below 0.1 foot-candle. He stated that there was no sudden change from day to night behavior patters at an level of light; the change was progressive. Other factors which have been shown to affect time and rate of bird calling are temperature, wind, and rain. Groebbels (1925) believed that early morning singing was a direct response to prevailing low temperatures. This study measure the composite and individual effects of time of day, season, light intensity, wind, rain, and temperature on calling. An attempt was made to apply correction factors to make possible more standardized calling counts. Observations were made on penned Chukars to relate sex and social organization to rally calling.

Analisou-se a composição, estrutura e padrão sazonal de formação dos bandos mistos de aves em um remanescente de floresta paludosa na planície costeira do Rio Grande do Sul, sul do Brasil. Efetuaram-se amostragens mensais de fevereiro de 2005 a janeiro de 2006, com observações ocasionais em 2004 e janeiro de 2005. Pontos de contagem foram utilizados para estimar a abundância relativa das espécies integrantes dos bandos, e transecções foram efetuadas para as observações dos bandos mistos. Quarenta e sete espécies foram registradas como participantes em 92 bandos amostrados. O número de indivíduos foi positivamente correlacionado com o número de espécies dentro dos bandos. O pool dos não-passeriformes foi pobremente representado nos bandos, enquanto passeriformes suboscines e oscines foram igualmente representados na riqueza dos bandos. Onívoros e insetívoros foram bem representados, mas os nectarívoros tiveram apenas uma espécie integrante; frugívoros e granívoros foram ausentes nos bandos mistos.Espécies migratórias foram pouco representadas nos bandos, como havia sido verificado para outras áreas de Floresta Atlântica. A regularidade de uma espécie em integrar bandos mistos foi uma função de sua abundância relativa, e as espécies mais conspícuas tenderam a ser os integrantes mais regulares. Detectaram-se quinze co-ocorrências significativas em 153 combinações possíveis (10%), onze delas positivas e quatro negativas. Nenhum tipo estrutural de bandos foi distinguido por meio de análise de agrupamento. Associações positivas e negativas talvez estejam relacionadas a similaridades e dissimilaridades na distribuição vertical de cada par de espécies. Os atributos de freqüência, tamanho e riqueza dos bandos foram os menores encontrados em qualquer outro estudo na Floresta Atlântica. Os bandos formaram-se o ano inteiro no Faxinal, mas a freqüência, o tamanho e a riqueza dos bandos diferiram significativamente entre os meses e entre as estações. Osvalores máximos destes três atributos foram encontrados na estação não-reprodutiva, lembrando os padrões encontrados em outros locais da Floresta Atlântica. A combinação de um pequeno grupo de espécies abundantes e bem distribuídas, junto com uma baixa riqueza local talvez torne a estrutura dos bandos mistos no Faxinal menos variável do que em outros locais da Floresta Atlântica. / The composition, structure and seasonal pattern of formation of mixed-species bird flocks were analyzed in a swamp forest remnant in the coastal plain of Rio Grande do Sul state, southern Brazil. Field work was carried out monthly from February 2005 to January 2006, plus occasional observations in 2004 and January 2005. Point counts estimated the relative abundance of flocking species, and transects were performed to access data relative to the flocks. Forty seven species were recorded as participants in 92 flocks sampled. The number of individuals was positively correlated with the number of species within the flocks. Non-passerines pool is less represented in the flocks, while suboscines and oscines were equally represented in the flock richness. Omnivores and insectivores were well represented, but nectarivores had only one flocking species while frugivores and granivores were absent in flocks. Migrants had low representation on the composition and structure of flocks, as predicted for lowland Atlantic Forest areas. The regularity of a given species in to integrate flocks was a function of its relative abundance, and the species more conspicuous tended to be the more regular flock joiners. Fifteen significant co-occurrences of 153 possible pairs (10%) were detected, eleven positively and four negatively associated. None structural type of flocks was recognized through cluster analyses. Positive and negative associations were better explained by similarities or dissimilarities on vertical distribution of each species pair. The attributes of frequency, flock size and flock richness were the lowest found in any study on Atlantic Forest. Flocks were formed throughout the year at Faxinal, but frequency, flock size and flock richness differed significantly between months and between seasons. Higher values of these three attributes were found in non-breeding season, reaching the pattern found elsewhere in Atlantic Forest. The combination of a little set of abundant and well distributed speciesand low richness perhaps makes the flock structure at Faxinal less variable than that of Atlantic Forest elsewhere.

Analisou-se a composição, estrutura e padrão sazonal de formação dos bandos mistos de aves em um remanescente de floresta paludosa na planície costeira do Rio Grande do Sul, sul do Brasil. Efetuaram-se amostragens mensais de fevereiro de 2005 a janeiro de 2006, com observações ocasionais em 2004 e janeiro de 2005. Pontos de contagem foram utilizados para estimar a abundância relativa das espécies integrantes dos bandos, e transecções foram efetuadas para as observações dos bandos mistos. Quarenta e sete espécies foram registradas como participantes em 92 bandos amostrados. O número de indivíduos foi positivamente correlacionado com o número de espécies dentro dos bandos. O pool dos não-passeriformes foi pobremente representado nos bandos, enquanto passeriformes suboscines e oscines foram igualmente representados na riqueza dos bandos. Onívoros e insetívoros foram bem representados, mas os nectarívoros tiveram apenas uma espécie integrante; frugívoros e granívoros foram ausentes nos bandos mistos.Espécies migratórias foram pouco representadas nos bandos, como havia sido verificado para outras áreas de Floresta Atlântica. A regularidade de uma espécie em integrar bandos mistos foi uma função de sua abundância relativa, e as espécies mais conspícuas tenderam a ser os integrantes mais regulares. Detectaram-se quinze co-ocorrências significativas em 153 combinações possíveis (10%), onze delas positivas e quatro negativas. Nenhum tipo estrutural de bandos foi distinguido por meio de análise de agrupamento. Associações positivas e negativas talvez estejam relacionadas a similaridades e dissimilaridades na distribuição vertical de cada par de espécies. Os atributos de freqüência, tamanho e riqueza dos bandos foram os menores encontrados em qualquer outro estudo na Floresta Atlântica. Os bandos formaram-se o ano inteiro no Faxinal, mas a freqüência, o tamanho e a riqueza dos bandos diferiram significativamente entre os meses e entre as estações. Osvalores máximos destes três atributos foram encontrados na estação não-reprodutiva, lembrando os padrões encontrados em outros locais da Floresta Atlântica. A combinação de um pequeno grupo de espécies abundantes e bem distribuídas, junto com uma baixa riqueza local talvez torne a estrutura dos bandos mistos no Faxinal menos variável do que em outros locais da Floresta Atlântica. / The composition, structure and seasonal pattern of formation of mixed-species bird flocks were analyzed in a swamp forest remnant in the coastal plain of Rio Grande do Sul state, southern Brazil. Field work was carried out monthly from February 2005 to January 2006, plus occasional observations in 2004 and January 2005. Point counts estimated the relative abundance of flocking species, and transects were performed to access data relative to the flocks. Forty seven species were recorded as participants in 92 flocks sampled. The number of individuals was positively correlated with the number of species within the flocks. Non-passerines pool is less represented in the flocks, while suboscines and oscines were equally represented in the flock richness. Omnivores and insectivores were well represented, but nectarivores had only one flocking species while frugivores and granivores were absent in flocks. Migrants had low representation on the composition and structure of flocks, as predicted for lowland Atlantic Forest areas. The regularity of a given species in to integrate flocks was a function of its relative abundance, and the species more conspicuous tended to be the more regular flock joiners. Fifteen significant co-occurrences of 153 possible pairs (10%) were detected, eleven positively and four negatively associated. None structural type of flocks was recognized through cluster analyses. Positive and negative associations were better explained by similarities or dissimilarities on vertical distribution of each species pair. The attributes of frequency, flock size and flock richness were the lowest found in any study on Atlantic Forest. Flocks were formed throughout the year at Faxinal, but frequency, flock size and flock richness differed significantly between months and between seasons. Higher values of these three attributes were found in non-breeding season, reaching the pattern found elsewhere in Atlantic Forest. The combination of a little set of abundant and well distributed speciesand low richness perhaps makes the flock structure at Faxinal less variable than that of Atlantic Forest elsewhere.

Analisou-se a composição, estrutura e padrão sazonal de formação dos bandos mistos de aves em um remanescente de floresta paludosa na planície costeira do Rio Grande do Sul, sul do Brasil. Efetuaram-se amostragens mensais de fevereiro de 2005 a janeiro de 2006, com observações ocasionais em 2004 e janeiro de 2005. Pontos de contagem foram utilizados para estimar a abundância relativa das espécies integrantes dos bandos, e transecções foram efetuadas para as observações dos bandos mistos. Quarenta e sete espécies foram registradas como participantes em 92 bandos amostrados. O número de indivíduos foi positivamente correlacionado com o número de espécies dentro dos bandos. O pool dos não-passeriformes foi pobremente representado nos bandos, enquanto passeriformes suboscines e oscines foram igualmente representados na riqueza dos bandos. Onívoros e insetívoros foram bem representados, mas os nectarívoros tiveram apenas uma espécie integrante; frugívoros e granívoros foram ausentes nos bandos mistos.Espécies migratórias foram pouco representadas nos bandos, como havia sido verificado para outras áreas de Floresta Atlântica. A regularidade de uma espécie em integrar bandos mistos foi uma função de sua abundância relativa, e as espécies mais conspícuas tenderam a ser os integrantes mais regulares. Detectaram-se quinze co-ocorrências significativas em 153 combinações possíveis (10%), onze delas positivas e quatro negativas. Nenhum tipo estrutural de bandos foi distinguido por meio de análise de agrupamento. Associações positivas e negativas talvez estejam relacionadas a similaridades e dissimilaridades na distribuição vertical de cada par de espécies. Os atributos de freqüência, tamanho e riqueza dos bandos foram os menores encontrados em qualquer outro estudo na Floresta Atlântica. Os bandos formaram-se o ano inteiro no Faxinal, mas a freqüência, o tamanho e a riqueza dos bandos diferiram significativamente entre os meses e entre as estações. Osvalores máximos destes três atributos foram encontrados na estação não-reprodutiva, lembrando os padrões encontrados em outros locais da Floresta Atlântica. A combinação de um pequeno grupo de espécies abundantes e bem distribuídas, junto com uma baixa riqueza local talvez torne a estrutura dos bandos mistos no Faxinal menos variável do que em outros locais da Floresta Atlântica. / The composition, structure and seasonal pattern of formation of mixed-species bird flocks were analyzed in a swamp forest remnant in the coastal plain of Rio Grande do Sul state, southern Brazil. Field work was carried out monthly from February 2005 to January 2006, plus occasional observations in 2004 and January 2005. Point counts estimated the relative abundance of flocking species, and transects were performed to access data relative to the flocks. Forty seven species were recorded as participants in 92 flocks sampled. The number of individuals was positively correlated with the number of species within the flocks. Non-passerines pool is less represented in the flocks, while suboscines and oscines were equally represented in the flock richness. Omnivores and insectivores were well represented, but nectarivores had only one flocking species while frugivores and granivores were absent in flocks. Migrants had low representation on the composition and structure of flocks, as predicted for lowland Atlantic Forest areas. The regularity of a given species in to integrate flocks was a function of its relative abundance, and the species more conspicuous tended to be the more regular flock joiners. Fifteen significant co-occurrences of 153 possible pairs (10%) were detected, eleven positively and four negatively associated. None structural type of flocks was recognized through cluster analyses. Positive and negative associations were better explained by similarities or dissimilarities on vertical distribution of each species pair. The attributes of frequency, flock size and flock richness were the lowest found in any study on Atlantic Forest. Flocks were formed throughout the year at Faxinal, but frequency, flock size and flock richness differed significantly between months and between seasons. Higher values of these three attributes were found in non-breeding season, reaching the pattern found elsewhere in Atlantic Forest. The combination of a little set of abundant and well distributed speciesand low richness perhaps makes the flock structure at Faxinal less variable than that of Atlantic Forest elsewhere.