Inaug.-Diss.--Königsberg. / Vita.

The lion-tailed macaque (Macaca silenus) is endangered due to habitat destruction with less than 3500 individuals remaining in isolated fragments of South-West India. Lion-tailed macaques do not reproduce readily in captivity and captive breeding may be relied upon for future conservation. Poor welfare can have negative effects on reproduction so it is important that lion-tailed macaque welfare is examined in captive groups. The aims of this thesis were to understand certain aspects of lion-tailed macaque welfare (behaviour and HPA physiology) in captive populations, with the view to making suggestions for management to promote the species' welfare and reproduction. Behaviour (188 hours), urine (n=133) and faecal samples (n=294) were collected from 38 lion-tailed macaques housed in four groups at the North of England Zoological Society (Chester Zoo), Bristol Zoological Gardens, Assiniboine Park Zoo and San Diego Wild Animal Park. The study successfully developed and validated assays to detect cortisol in lion-tailed macaque urine and faeces. The assays were then subsequently used to explore behaviour and HPA activity in these endangered primates. The institution in which the individuals were housed and basic life history parameters (age and sex) were explored to further understand the interplay between behaviour and physiology. Social relationships were assessed by measuring proximity (inter-individual distances and time spent in "arms-reach"). Finally the effect of visitors on behaviour, HPA activity and enclosure use was explored. There was significant variation between institutions in behaviour and HPA activity but not proximity. The age of lion-tailed macaques modified their behaviour, but not their HPA activity or proximity. The sex of lion-tailed macaques did not modify behaviour, HPA activity or proximity. The effect of visitors on lion-tailed macaques in the current study is not clear and confirms previous research on the visitor effect on captive primates. It can be concluded from this research that lion-tailed macaques are sensitive to the environment in which they are housed, indicating factors which may have negative effects on their captive breeding rates and ability to cope with habitat fragmentation for population's in-situ. The study has highlighted the need for each captive and wild group of lion-tailed macaques to be considered and monitored separately with regard to welfare and breeding.

In the topical abundance or superabundance of Shakespeares King Lear, almost all major thematic patterns, images, and symbols are linked to Lears enigmatic companion, the Fool. The Fool surpasses Shakespeares other fools when he is given a major role, yet he is more than a major figure: he is the pivot for action and interpretation. The presence and the importance of the Fool are emphasized further when almost half of Lears characters are referred to as fools. The stark, barren hinterland of Lear is shot through by the conspicuously forceful presence of folly. The fools propensity for misrule coupled with his centrality to the text results in a dramatic structure that itself breaks the rules. King Lear is the most generically puzzling play in Shakespeares corpus. Lear, a tragedy, draws upon comedy, history, romantic comedy, romance, and morality in indefinable and unparalleled ways. Just as form is juxtaposed in Lear, religious systems or identities are also contrasted. Pagan, Christian, existential, nihilistic, and moralistic interpretations are readily discernable. The fool, a potential nexus for structural questions, may also be at the heart of the question of spiritual identity. To locate the fool at the source of both structural and spiritual problems, and to discern why the fool factors so prominently especially in a play viewed as Shakespeares darkest, are the two endeavours of this thesis.

In the topical abundance or superabundance of Shakespeares King Lear, almost all major thematic patterns, images, and symbols are linked to Lears enigmatic companion, the Fool. The Fool surpasses Shakespeares other fools when he is given a major role, yet he is more than a major figure: he is the pivot for action and interpretation. The presence and the importance of the Fool are emphasized further when almost half of Lears characters are referred to as fools. The stark, barren hinterland of Lear is shot through by the conspicuously forceful presence of folly. The fools propensity for misrule coupled with his centrality to the text results in a dramatic structure that itself breaks the rules. King Lear is the most generically puzzling play in Shakespeares corpus. Lear, a tragedy, draws upon comedy, history, romantic comedy, romance, and morality in indefinable and unparalleled ways. Just as form is juxtaposed in Lear, religious systems or identities are also contrasted. Pagan, Christian, existential, nihilistic, and moralistic interpretations are readily discernable. The fool, a potential nexus for structural questions, may also be at the heart of the question of spiritual identity. To locate the fool at the source of both structural and spiritual problems, and to discern why the fool factors so prominently especially in a play viewed as Shakespeares darkest, are the two endeavours of this thesis.

The aim of the present work was to investigate the impact of the varying environmental conditions on the taxonomic and phenotypic diversification of a geographically widespread and ecologically successful Old World primate genus, the macaques (Cercopithecidae: Macaca). To this end, the relationship between geography, ecology, phylogeny, and phenotypic variation among macaques was investigated. Constraints to phenotypic variation – and thus evolution – were also analysed in the form of observed amounts of phenotypic variation and patterns of phenotypic integration. A total of 72 standard linear measurements of teeth and associated cranial and mandibular structures were taken for a total sample of 744 specimens from 13 species of macaques. Climate and ecological data were collated from the literature. Univariate and multivariate statistics were employed for the analysis. Patterns of variation, covariation, and allometry were analysed in the dentition, both within and between species. The ecogeographical analysis was carried out by means of two-block partial least squares and a type of multivariate regression, both in a phylogenetic framework. Phylogenetic signal was tested for by means of Blomberg’s K. Macaque teeth differ in their variability. All teeth covary with each other, although correlations are strongest within tooth classes. Size was a strong contributing factor to dental integration, as evinced by lower correlations between teeth once allometric effects were removed. Integration patterns also showed modularity between the anterior and the posterior dentition. Between-species variation in overall craniodental size was associated with temperature, latitude, and body size. Species also varied, albeit to a lesser degree, along an antero-posterior contrast in relative tooth size. Larger anterior were found to be associated with frugivory and tropical ecology, whereas a larger posterior dentition was linked to a more folivorous diet and temperate environments. The latter pattern was largely a function of phylogenetic relatedness. Phylogenetic signal was generally strong in the dentition, although it was substantially greater in the anterior teeth (incisors and canines) than in the posterior teeth (premolars and molars). Macaques show adaptive differentiation in body size in response to temperature along a latitudinal cline, corroborating the presence of the Bergmann effect in macaques. There was no conclusive support for further adaptive differentiation, despite an association between relative tooth size and diet. Allometry appears to channel evolutionary divergence of macaques along a line of least evolutionary resistance, and developmental modularity allows for partly uncoupled evolution of the anterior and posterior dentition. Future research should be aimed at broadening the taxonomic scope to include craniodental variation of the African papionins and cercopithecins in order to put the observed macaque patterns in a broader evolutionary context.