Thesis (Ph. D.)--Murdoch University, 1993. / Thesis submitted to the School of Veterinary Studies. Includes bibliographical references (leaves 463-484).

Thesis (Ph. D.)--University of Washington, 2008. / Vita. Includes bibliographical references (leaves 87-95).

Thesis (M. S.)--Ohio State University, 2008. / Title from first page of PDF file. Includes bibliographical references (p. 91-100).

Thesis (Ph. D.)--Duke University, 2008.

Thesis (Ph. D.)--University of Texas at Austin, 2002. / Vita. Includes bibliographical references (leaves 161-182). Available also in an electronic version.

Dissertation (PhD)--University of Stellenbosch, 2007. / Bibliography. Also available via the Internet.

Thesis (Ph. D.)--Miami University, Dept. of Zoology, 2004. / Title from second page of PDF document. Includes bibliographical references.

Thesis (Ph. D.)--Florida State University, 2006. / Advisor: J. Anthony Stallins, Florida State University, College of Social Sciences, Dept. of Geography. Title and description from dissertation home page (viewed June 7, 2006). Document formatted into pages; contains xiii, 284 pages. Includes bibliographical references.

Thesis (PhD)--Stellenbosch University, 2005. / ENGLISH ABSTRACT: Despite more than three decades of research, and the postulation of more than 30 hypotheses of function, the adaptive significance of play remains unknown. This study quantitatively evaluated a selection of hypotheses of function, using data collected from a wild population of small, social carnivore, the meerkat, Suricata suricatta. The study found that although play in meerkats carried an energetic cost, with individuals modulating their frequency of play in response to their energy intake, none of the hypotheses evaluated by the study could identify the adaptive benefits that meerkats derived from play. Play did not increase 'social harmony' by reducing aggression between playmates, nor did it strengthen an individual's bonds to its social group, such that it remained in the group for longer, or contributed more to the group's cooperative activities. There was no evidence that meerkats used play to strengthen alliances between individuals, and young meerkats played no more frequently with their future dispersal partners than with matched controls with which they did not disperse. Play fighting experience did not improve a meerkat's subsequent fighting skills, and individuals that ultimately won the dominant breeding position within a group (through serious fighting) played no more frequently, and no more successfully, as youngsters, than the littermates that they defeated in combat. Although play was inhibited by aggression, meerkats did not use play to contest, assert or establish dominance status, and there was little evidence to suggest that the preference young meerkats showed for play partners that were well matched in age, size and ability arose from their use of play for self-assessment. This study assessed only those hypotheses of function that predicted benefits that were of importance to the inclusive fitness of the study species. For example, the enhancement of social harmony and group cohesion should be invaluable to a species whose survival is dependent upon social cooperation; and the high reproductive skew exhibited by this species places huge value upon fighting skill and the ability to win social dominance. As a consequence, this study's negative findings suggest strongly that play is not capable of providing these benefits, and that play behaviour is unlikely to be used for these purposes in any mammal species. I conclude that the most likely function of play (based on play's ubiquitous characteristics, and the findings of neurological research on rats) is the promotion of growth of the cerebral cortex. / AFRIKAANSE OPSOMMING: Ten spyte van meer as drie dekades van navorsing en die voorstelling van meer as 30 hipoteses oor funksie, bly die aanpassingswaarde van spelonbekend. Hierdie studie is 'n kwantitatiewe evaluasie van verskeie hipoteses oor funksie, en gebruik data versamel vanuit 'n wilde bevolking van 'n klein sosiale karnivoor, die meerkat, Suricata suricatta. Die studie het bevind dat hoewel spel in meerkaaie 'n energetiese koste beloop, met individue wat hul spelfrekwensie aanpas by energie-inname, geen-een van die hipoteses onder beskouing die aanpassingswaarde van spel vir meerkaaie kon verduidelik nie. Spel het nie "sosiale harmonie" bevorder deur die afuame in aggressie tussen speelmaats nie, en het ook nie 'n individu se verbintenis tot sy sosiale groep versterk sodat hy langer in die groep sou bly of meer sou bydra tot samewerkingsaktiwiteite nie. Daar was geen bewyse vir die gebruik van spel in die versterking van bondgenootskappe tussen individue nie, en jong meerkaaie het nie meer gereeld met toekomstige verspreidings-venote gespeel as met gepaarde kontroles saam met wie hulle nie uiteengegaan het nie. Speelse gevegte het nie 'n meerkat se daaropvolgende gevegsvermoëns verbeter nie, en die individue wat uiteindelik die dominante voortplantingsposisie in 'n groep gewen het (deur ernstige stryd) het nie meer gereeld ofmeer suksesvol as jongelinge gespeel in vergelyking met die werpselmaats wat hulle in die stryd oorwin het nie. Hoewel spel deur aggressie onderdruk is, het meerkaaie spel nie gebruik om dominante range te beveg, bevestig of tot stand te bring nie. Daar was min bewyse ter ondersteuning van die voorstel dat jong meerkaaie se voorkeur vir speelmaats wat hul gelyke is in ouderdom, grootte en vermoë, onstaan het in die gebruik van spel vir selfondersoek. Hierdie studie het slegs die hipoteses van funksie beskou wat voorspellings gemaak het oor die voordele wat belangrik is in die inklusiewe fiksheid van die studie-species. Byvoorbeeld, die verbetering van sosiale harmonie en groepsamehang behoort van onskatbare waarde te wees vir 'n species wat afhanklik is van sosiale samewerking vir oorlewing; en die hoë graad van voorkeuraanwas duidelik in hierdie species plaas groot waarde op gevegsvaardighede en die vermoë om sosiale dominansie te wen. Gevolglik dui hierdie ondersoek se negatiewe bevindinge daarop dat spel nie hierdie voordele kan bied nie, en dat speelgedrag heel waarskynlik nie vir hierdie doeleindes in enige soogdier-species gebruik word nie. Ek kom tot die gevolgtrekking dat die heel waarskynlikste funksie van spel (gebaseer op spel se alomteenwoordige kenmerke en die bevindinge van neurologiese navorsing op rotte) die bevordering van groei in die serebrale korteks is.

Captive animals are frequently subject to imposed, uncontrollable stressors to which they respond through behavioural flexibility, or, failing which, exhibit pathology. Chimpanzees provide an intriguing model to examine how captive environments influence the responses of animals to stress. My study investigates the responses of a group of chimpanzees to imposed stressors of captivity at the Johannesburg Zoo, South Africa. My study comprised four components. Firstly, I examined the effects of spatial restriction on chimpanzee behaviour with regard to an enclosure enlargement, testing several existing models of coping with spatial crowding and another model, based on the coping hypothesis of abnormal behaviour. Behavioural observations of the chimpanzees in their indoor and outdoor exhibits before, during, immediately after and 10 weeks after the enclosure reconstruction revealed that the chimpanzees used tension-reduction and conflict-avoidance tactics as a means to cope with spatial crowding. Moreover, abnormal behaviour appears to provide an outlet for stress under crowding. Secondly, I assessed the long-term effects of past spatial environments on the space use and group spacing of the chimpanzees, five years after the enclosure change. Through behavioural observations and mapping the locations of individuals, I found that the chimpanzees exhibit space-use bias and limited group spacing, contingent on the dimensions of the old enclosure that were not explained by factors such as social or thermal conditions and zoo visitor effects. I propose that the spacing patterns may be due to spatial learned helplessness. Thirdly, I examined the effect of two social manipulations, mandated by zoo management, on the behaviour and socio-dynamics of the chimpanzees. The chimpanzees responded to social change through selective social interactions and non-social behavioural responses suggest that removing an individual was less stressful than the merging of two groups. Finally, I investigated the role of shade as a thermoregulatory resource for captive chimpanzees. Individuals used shade frequently despite observations taking place during the austral winter period, suggesting that shade is a valuable thermal resource for chimpanzees. In conclusion, the chimpanzees responded to most imposed stressors (spatial crowding, social change and thermal stress) through behavioural flexibility, implying successful coping, but failed to cope with previous spatial restrictions, resulting in limited space-use behaviour.