Nutrient availability and herbivory can regulate primary production in ecosystems, but little is known about how, or whether, they may interact with one another. Here I investigate how nitrogen availability and insect herbivory interact to alter above- and belowground plant community biomass in an old-field ecosystem. In 2004, 36 experimental plots were established in which soil nitrogen (N) availability (at three levels) was manipulated and insect abundance (at two levels) in a completely randomized plot design. In 2009, after six years of treatment, I measured aboveground biomass and assessed root production at peak growth. Overall, I found a significant effect of soil N availability on both above- and belowground plant biomass while insects affected only aboveground biomass of subdominant plant species and coarse root production; there were no statistical interactions between N availability and insect herbivory for any response variable. Specifically, responses of aboveground and belowground community biomass to nutrients were driven by reductions in soil N, but not additions, indicating that soil N may not be primarily limiting production in this ecosystem. Insect herbivory altered the aboveground biomass of the subdominant plant species and altered allocation patterns to coarse root production belowground. Overall, the results of six years of nutrient amendments and insect removals suggest strong bottom-up influences on total plant community productivity.

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A fast response vertical anemometer and wet and dry bulb thermocouples were used to measure the turbulence within a forest canopy. Five trials of ten minutes duration were run in each of a sixty-five foot high pine forest and a fifteen foot high lodgepole pine forest. [...]

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Sensible heat flux, a key component of the surface energy balance, is difficult to estimate in practice. This study was conducted to see if backpropagation neural networks could estimate sensible heat flux by using horizontal wind speed, air temperature, radiometric surface temperature, net radiation, and time as input. Ground measurements from the First ISLSCP (International Satellite Land Surface Climatology Project) Field Experiment (FIFE), collected in 1987 and 1989 over a prairie grassland in Kansas, were used for network training and validation. Networks trained on part of the data from a narrow range of space-time coordinates performed well over the other part, with error (root mean square error divided by mean of observations) values as low as 0.24. This indicates the potential in neural networks for linking sensible heat flux to routinely measured meteorological variables and variables amenable to remote sensing. When the networks were tested with data from other space-times, performance varied from good to poor, with average error values around 1.26. This was mainly due to lack of input variables parameterizing canopy morphology and soil moisture, indicating that such variables should be incorporated in the design of future networks intended for large scale applications.

Over the next three decades, there are many predicted disturbances to Earth’s oceans, such as El Nino and hurricanes, which will lead to mass coral bleaching effects. Marine protected areas have been utilized worldwide to maintain coral population sizes and remediate external stressors, such as overfishing or mining. Using a series of modeling techniques, this thesis will propose an experiment that will determine the optimal distance and location for a future MPA in New Zealand. It will also be measuring whether one large reserve or a network of smaller MPAs are more effective in the regeneration of stony corals. These models will be based on Solemosmilia variabilis, the most common stony coral in the region. Based on past studies, it is hypothesized that there will be a significant positive increase with the metapopulation growth of corals in both protected areas. It is also predicted that there will be a higher rate of connectivity within a network of smaller marine protected areas if the MPAs are less than 2 km apart. If the distance is greater, one larger MPA will be more effective due to the lower rates of genetic drift.

Increasing introductions of non-native terrestrial slugs (Mollusca: Gastropoda) are a concern to North American regulatory agencies as these generalists impact the yield and reduce the aesthetic value of crop plants. Understanding how the increase in diversification in North American cropping systems affects non-native gastropods and finding effective biological control options are imperative for pest management; however, little research has been done in this area. This study tested the hypothesis that dietary diversification affects the biological control capacity of a generalist predator and allows the slug pest Deroceras reticulatum (Müller) (Stylommatophora: Agriolimacidae) to more effectively fulfill its nutritional requirements. Results showed no significant correlations between dietary diversification and slug development; however, this was likely due to the addition of romaine lettuce to all treatments. The study also showed that dietary diversification had no significant effect on D. reticulatum egg production, with self-fertilizing slugs consistently having significantly higher egg production than outcrossing slugs. Most significantly, this research demonstrated reductions in plant damage by D. reticulatum in treatments containing the North American carabid beetle Scarites quadriceps Chaudoir (Coleoptera: Carabidae) with the presence of alternative prey having no effect, supporting its use in biological control efforts in spite of its generalist feeding habits.

Northern fringe populations generally have low amounts of genetic variation and inhabit habitats where specific adaptations are needed. On the Swedish west coast, the natterjack toad (Bufo calamita) inhabits skerry islands. I have examined: I) adaptation to two environmental stressors in this habitat; II) the genetic population structure within the skerry habitat; III) the effects of neutral genetic variation, selection and genetic drift on trait divergence within the skerry habitat; and IV) the effects of genetic variation on fitness under three thermal conditions of varying stressfulness. V) I have also studied the impact of putative local adaptations on the Scandinavian green toad (Bufo viridis) conservation programme. The results suggest that the skerry natterjack toads are locally adapted to the desiccation risk in their habitat. However, despite inhabiting a more saline habitat, they had a lower salinity tolerance when compared to their conspecifics in the more general habitat. The lowered salinity tolerance is most likely explained by the presence of negative genetic correlations between salinity tolerance and desiccation avoidance and suggests that the occurrence of multiple environmental stressors may constrain adaptation. Within the skerry habitat, the toads exhibited a strong population structure with populations differing in their levels of genetic variation. Moreover, within the skerry habitat, the results suggest uniform selection pressures. However, correlations between trait values and neutral genetic variation suggest that inbreeding depression may affect trait values and thus potentially constrain adaptation. In the natterjack toad, fitness costs associated with lack of genetic variation were only present under benign conditions and not under more natural conditions. This suggests that environmental stress masks inbreeding depression in these traits under natural conditions. In the study regarding the Scandinavian green toads, I found that one population inhabiting a saline habitat had a higher salinity tolerance than other populations in less saline habitats. This suggests the presence of local adaptation, which should be acknowledged in the green toad conservation programme. Several of the northern fringe populations of toads fulfill the criteria of being Evolutionary Significant Units and their conservation thus should be prioritized.

Recent studies within the past decade or so have shown the importance of algae in geomorphic and hydrologic processes of lotic systems. However, the ecohydraulic role of algae in bedrock systems has largely been ignored. In addition, the utility of algae as indicators of channel dynamics have often been assumed by geomorphologists, but relatively few studies have examined this relationship. The purpose of this study was to determine whether algae, specifically diatoms, are useful indicators of channel geomorphological dynamics, and to examine if distinct habitats or biotopes typical in fluviokarst and bedrock systems provide unique habitat space for diatoms, and to address the potential ecohydraulic implications. The investigation was performed in a 100 m reach of Shawnee Run, a limestone, fluviokarst tributary to the Kentucky River in Mercer County, KY. The results of the study showed that periphyton are not useful indicators of channel dynamics, and that biotopes and other distinct habitats, including riffles, bedforms, and fine sediment, do not provide unique habitat in terms of diatom community composition.

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