No description available.

No description available.

No description available.

No description available.

Urban areas are responsible for the majority of global anthropogenic CO2 emissions. Urbanization has altered the structure and function of terrestrial ecosystems and is increasing rapidly, further modifying global carbon cycling. The three research papers in this dissertation explore the role of urban vegetation in the carbon cycle using a combination of atmospheric observation, field measurements, remote sensing, and modeling. First, I characterized the spatiotemporal patterns of observed atmospheric CO2 mixing ratios and compared these data to estimated CO2 fluxes at three sites across Boston's urban-to-rural gradient. Total fossil fuel emissions estimates ranged from 1.5 to 37.3 Mg C ha-1 yr-1 between rural Harvard Forest and urban Boston. Despite large differences in emissions, atmospheric CO2 concentrations only differed by approximately 5%. The growing season length in Boston was approximately 31 days longer than in Harvard Forest, enhancing the period for biological carbon uptake. In Boston, gross primary production was 3.8 Mg C ha-1 yr-1, which was ~75% lower than gross primary production at Harvard Forest and ~10% of total anthropogenic carbon fluxes in Boston. Second, I assessed how forest-to-urban land cover change affected both aboveground biomass and productivity across eastern Massachusetts. I found that urban land covers contained less than half the biomass of adjacent forests, but the mean basal area increment of existing trees nearly doubled with development over time from 17.1 ± 3.0 to 35.8 ± 4.7 cm2 yr-1. Scaling this increase in growth suggests an aboveground biomass growth rate of 1.8 ± 0.4 Mg C ha-1 yr-1, a rate similar to that found in Harvard Forest, despite having only ~1/3 the standing aboveground biomass. Last, I assessed how above- and belowground ecosystem characteristics changed as a function of time since development and development intensity. I found that soil C and aboveground biomass showed significant differences with time since development. My data suggests that soil C, N, and bulk density are dependent on land use history, with previously agricultural sites consistently showing higher rates of soil N and C accumulation than previously forested and grassland sites. Taken as a whole, this dissertation highlights the potential consequences of altered ecological and environmental conditions on tree growth, the legacy effects of land use history, climate, and land management practices on below ground soil C and N, and the importance of vegetation in the C cycle in urban areas.

A stochastic framework to simulate the process of initiation, diameter growth, death and self-pruning of branches in loblolly pine (Pinus taeda L.) trees was developed. A data set was obtained from a destructive sampling of whorl sections from 34 trees growing under different initial spacing. Data from dissected branches were used to develop a model for representing knot shape, which assumed that the live portion of a knot can be modeled by a one-parameter equation and the dead portion by assuming a cylindrical shape. For the developed knot model analytical expressions were derived for estimating the volume of knots (live/dead portions) for three types of branch conditions on simulated trees: (i) live branches, (ii) non-occluded dead branches, and (iii) occluded dead branches. This model was intended to recover information on knots shape and volume during the simulation process of branch dynamics. Three different components were modeled and hierarchically connected: whorl, branches and knots. For each new growing season, whorls and branches are assigned stochastically along and around the stem. Thereafter, branch diameter growth is predicted as function of relative location within the live crown and stem growth. Using a taper equation, the spatial location (X,Y,Z) of both live and dead portion of simulated knots is maintained in order to create a 3D representation of the internal stem structure. At the end of the projection period information on (i) vertical trend of branch diameter and location along and around the stem, (ii) volume of knots, and (iii) spatial location, size and type (live and dead) of knots can be obtained. The proposed branch model was linked to the individual-tree growth and yield model PTAEDA3.1 to evaluate the effect of initial spacing and thinning intensity on branch growth in sawtimber trees. The use of the dynamic branch model permitted generation of additional information on sawlog quality under different management regimes. The arithmetic mean diameter of the largest four branches, one from each radial quadrant of the log (i.e. Branch Index, BI) and the number of whorls per log were considered as indicators of sawlog quality. The developed framework makes it possible to include additional wood properties in the simulation system, allowing linkage with industrial conversion processes (e.g. sawing simulation). This integrated modeling system should promote further research to obtain necessary data on crown and branch dynamics to validate the overall performance of the proposed branch model and to improve its components. / Ph. D.

Sustainable management of mixed-species forests for timber is underpinned by research on forest stand dynamics and quantification of tree recruitment, growth and mortality rates. Different performance among species across light gradients theoretically prevents more shade-tolerant species from excluding shade-intolerant species, driving succession and allowing species coexistence. This research investigates stand dynamics and performance trade-offs for co-occurring tree species: Nothofagus fusca (red beech) and Nothofagus menziesii (silver beech), which together dominate extensive areas of New Zealand’s indigenous forest. Using permanent plot data, measurements of permanently tagged individuals are used to quantify recruitment, growth and mortality rates for each species, across size classes and life-history stages (i.e. seedlings, trees). First, seedling growth and mortality is examined in relation to microhabitats (e.g., light, substrate type) and contrasted with patterns of seedling abundance. Second, spatially explicit permanent plot data are used to examine tree growth in relation to competition, local disturbance and tree size over a 23-year period. Third, the influence of competition and disturbance on tree mortality and spatial patterns of tree mortality are examined. Fourth, a simulation model for tree population dynamics is parameterised for mixed-Nothofagus forest and used to evaluate long term consequences of disturbances (e.g. alternate harvesting regimes) on structure and composition. Small-scale disturbance favoured each species at different life stages and for different measures of performance (i.e. recruitment, growth or mortality). N. fusca seedlings and trees grew fast in high light microhabitats, such as those created by small-scale disturbances, but adult N. fusca mortality was elevated near sites of recent disturbance. By contrast, N. menziesii trees grew faster near sites of recent disturbance, which may help this species persist. Consequently, simulation results showed that small-scale disturbance frequency was a major determinant of forest composition and structure, determining whether N. fusca or N. menziesii is dominant. The simulation model could be developed further and used to inform the sustainable management of mixed-Nothofagus forests.

A alta demanda por madeira de eucalipto para diversas finalidades traz a necessidade da expansão de plantios para áreas em condições de déficit hídrico, o que leva o interesse de entender os processos fisiológicos relacionados ao crescimento da árvore, para possibilitar uma alta produtividade das florestas nessas áreas. Com base nessas informações, o objetivo deste trabalho foi avaliar o efeito da variação de níveis de restrições hídricas em clones de Eucalyptus spp. plantados em diferentes condições edáficas, pelo estudo das respostas ecofisiológicas, para isso foram instalados em janeiro 2012 dois experimentos na Estação Experimental de Ciências Florestais de Itatinga (EECFI) pertencente a Escola Superior de Agricultura \"Luiz de Queiroz\" (ESALQ) - USP, o experimento 1 foi instalado no solo classificado como Latossolo vermelho distroférrico típico A moderado textura argilosa (LVdf) e o experimento 2 no solo Latossolo vermelho distrófico típico A moderado textura argilosa (LVd-1).Cada experimento foi implantado em blocos casualizados em arranjo fatorial de 6 x 3, seis clones de eucalipto (COP 1404, COP 1407, AEC 1528, AEC 0042, AEC 0224, AEC 0144) e três espaçamentos (3m x1m, 3mx2m, e 3mx4m), com seis repetições, e parcelas quadradas constituídas por 25 plantas, sendo as nove plantas centrais consideradas como a área útil de medição. Foram realizadas avaliações das variáveis: DAP (cm), altura total (m), comprimento da copa (m) e índice de área foliar (IAF m2/m2) trimestralmente, no período de 21 até 37 meses de idade para as variáveis altura total e DAP, e o comprimento da copa e IAF a partir do mês 27. Para a mensuração do IAF foram utilizados dois equipamentos LAI-2000, onde a coleta de dados foi fora e sob o dossel. O folhedo foi coletado mensalmente em todas as parcelas. A avaliação da eficiência e uso de água foi aos 29 e 35 meses, analisando área foliar e as caraterísticas estomáticas em quatro blocos, nos quais três árvores foram amostradas sistematicamente. Para avaliação no incremento de diâmetro no tronco, se instalaram faixas dendrométricas em duas árvores por parcela e as leituras foram quinzenais, sendo correlacionadas com variáveis climáticas no período de 15 e 45 dias de defasagem. Os clones que apresentaram melhor desenvolvimento e estratégia do uso da água foram AEC 1528, AEC 0144 e AEC 0042 obtendo como respostas: maior DAP, altura total, comprimento da copa, área foliar, diâmetro estomático, e menor IAF e concentração de estômatos. Em relação ao peso de folhedo depositado verificou-se que a tendência de deposição oscilou no decorrer do tempo em função da fenologia da planta. Na dinâmica de crescimento cambial os incrementos apresentaram correlações com as variáveis climáticas, principalmente com a precipitação pluviométrica, quanto maior a defasagem maior a correlação. Correlacionando todas as variáveis, os clones apresentaram distintos padrões de respostas ecofisiológicas. / Eucalypts plantations are expanding to drought áreas in Brazil in order to attend the high demand of eucalypts wood for various purposes. Therefore, the understanding the physiological process related to tree growth in this condition is one of the research priorities. This study aimed to evaluate the effect of ecophysiological responses to water restrictions levels in growth of Eucalyptus spp clones, planted in different soil conditions. The two field experiments were planted in January 2012 at the Itatinga Experimental Station, College of Agriculture \"Luiz de Queiroz\" (ESALQ), São Paulo State University. Experiment 1 was planted on a site which the soil is classified as \"A moderate typical dystrophic Red Latosol with clay texture (LVdf)\" and experiment 2 on \"A moderate typical dystrophic Red Latosol with clay texture (LVd- 1)\". The experiments were established in a randomized block design with factorial 6 x 3 arrangement [six eucalypts clones (COP 1404, COP 1407, AEC 1528 AEC 0042, AEC 0224, AEC 0144) and three spacings (3m x1m, 3mx2m and 3mx4m)], with six replications, and square plots (5x5 trees). The measurements were carried out on the nine central trees. From the 21 to 37 months of age, the following variables were assessed every three months: DBH (diameter at breast height in cm), height (m) and crown length (m). Leaf area index (LAI m2 / m2) was evaluated every three months after 27 months of age. Two LAI-2000 equipments were used to measure the LAI, one under and another out of the canopy. Litter was collected monthly on all plots. The efficiency of water use was evaluated at 29 and 35 months of age, based on leaf area and stomatal characteristics of samples collected in three trees sampled systematically in four blocks. Dendrometer belts were used to evaluate the diameter growth of two trees per plot. Data collected fortnightly from dendrometers belts were correlated with climatic variables with 15 and 45-da lags. AEC 1528, AEC 0144 and AEC 0042 clones showed higher DBH, total height, crown length, leaf area, stomatal diameter and lower IAF and stomatal concentration, showing better strategy of water use and growth. Deposition of litter fluctuated over time and varied with the tree phenology. Cambium growth dynamics correlated with climate variables, especially with the rainfall. Higher correlations were observed with higher measured lag. Clones showed distinct patterns of ecophysiological responses when the correlations of all variables were considered.

A alta demanda por madeira de eucalipto para diversas finalidades traz a necessidade da expansão de plantios para áreas em condições de déficit hídrico, o que leva o interesse de entender os processos fisiológicos relacionados ao crescimento da árvore, para possibilitar uma alta produtividade das florestas nessas áreas. Com base nessas informações, o objetivo deste trabalho foi avaliar o efeito da variação de níveis de restrições hídricas em clones de Eucalyptus spp. plantados em diferentes condições edáficas, pelo estudo das respostas ecofisiológicas, para isso foram instalados em janeiro 2012 dois experimentos na Estação Experimental de Ciências Florestais de Itatinga (EECFI) pertencente a Escola Superior de Agricultura \"Luiz de Queiroz\" (ESALQ) - USP, o experimento 1 foi instalado no solo classificado como Latossolo vermelho distroférrico típico A moderado textura argilosa (LVdf) e o experimento 2 no solo Latossolo vermelho distrófico típico A moderado textura argilosa (LVd-1).Cada experimento foi implantado em blocos casualizados em arranjo fatorial de 6 x 3, seis clones de eucalipto (COP 1404, COP 1407, AEC 1528, AEC 0042, AEC 0224, AEC 0144) e três espaçamentos (3m x1m, 3mx2m, e 3mx4m), com seis repetições, e parcelas quadradas constituídas por 25 plantas, sendo as nove plantas centrais consideradas como a área útil de medição. Foram realizadas avaliações das variáveis: DAP (cm), altura total (m), comprimento da copa (m) e índice de área foliar (IAF m2/m2) trimestralmente, no período de 21 até 37 meses de idade para as variáveis altura total e DAP, e o comprimento da copa e IAF a partir do mês 27. Para a mensuração do IAF foram utilizados dois equipamentos LAI-2000, onde a coleta de dados foi fora e sob o dossel. O folhedo foi coletado mensalmente em todas as parcelas. A avaliação da eficiência e uso de água foi aos 29 e 35 meses, analisando área foliar e as caraterísticas estomáticas em quatro blocos, nos quais três árvores foram amostradas sistematicamente. Para avaliação no incremento de diâmetro no tronco, se instalaram faixas dendrométricas em duas árvores por parcela e as leituras foram quinzenais, sendo correlacionadas com variáveis climáticas no período de 15 e 45 dias de defasagem. Os clones que apresentaram melhor desenvolvimento e estratégia do uso da água foram AEC 1528, AEC 0144 e AEC 0042 obtendo como respostas: maior DAP, altura total, comprimento da copa, área foliar, diâmetro estomático, e menor IAF e concentração de estômatos. Em relação ao peso de folhedo depositado verificou-se que a tendência de deposição oscilou no decorrer do tempo em função da fenologia da planta. Na dinâmica de crescimento cambial os incrementos apresentaram correlações com as variáveis climáticas, principalmente com a precipitação pluviométrica, quanto maior a defasagem maior a correlação. Correlacionando todas as variáveis, os clones apresentaram distintos padrões de respostas ecofisiológicas. / Eucalypts plantations are expanding to drought áreas in Brazil in order to attend the high demand of eucalypts wood for various purposes. Therefore, the understanding the physiological process related to tree growth in this condition is one of the research priorities. This study aimed to evaluate the effect of ecophysiological responses to water restrictions levels in growth of Eucalyptus spp clones, planted in different soil conditions. The two field experiments were planted in January 2012 at the Itatinga Experimental Station, College of Agriculture \"Luiz de Queiroz\" (ESALQ), São Paulo State University. Experiment 1 was planted on a site which the soil is classified as \"A moderate typical dystrophic Red Latosol with clay texture (LVdf)\" and experiment 2 on \"A moderate typical dystrophic Red Latosol with clay texture (LVd- 1)\". The experiments were established in a randomized block design with factorial 6 x 3 arrangement [six eucalypts clones (COP 1404, COP 1407, AEC 1528 AEC 0042, AEC 0224, AEC 0144) and three spacings (3m x1m, 3mx2m and 3mx4m)], with six replications, and square plots (5x5 trees). The measurements were carried out on the nine central trees. From the 21 to 37 months of age, the following variables were assessed every three months: DBH (diameter at breast height in cm), height (m) and crown length (m). Leaf area index (LAI m2 / m2) was evaluated every three months after 27 months of age. Two LAI-2000 equipments were used to measure the LAI, one under and another out of the canopy. Litter was collected monthly on all plots. The efficiency of water use was evaluated at 29 and 35 months of age, based on leaf area and stomatal characteristics of samples collected in three trees sampled systematically in four blocks. Dendrometer belts were used to evaluate the diameter growth of two trees per plot. Data collected fortnightly from dendrometers belts were correlated with climatic variables with 15 and 45-da lags. AEC 1528, AEC 0144 and AEC 0042 clones showed higher DBH, total height, crown length, leaf area, stomatal diameter and lower IAF and stomatal concentration, showing better strategy of water use and growth. Deposition of litter fluctuated over time and varied with the tree phenology. Cambium growth dynamics correlated with climate variables, especially with the rainfall. Higher correlations were observed with higher measured lag. Clones showed distinct patterns of ecophysiological responses when the correlations of all variables were considered.

Humans have affected the Swedish forests for a long time, mainly through agriculture and forestry. Since the beginning of the 20th century, a lot of changes have taken place in forest policy, which have affected the structures of our forests, such as tree age, size and species distributions. By using information from different types of historical data, we can increase our understanding of the earlier structures of forests and how to manage them in the future. This study investigated changes in annual growth and changes in age, diameter, height and tree species distribution in the forests of Östergötland from year 1927 to the period 1983-2017, using data from the Swedish National Forest Inventory. Six tree species where used, Picea spp, Pinus spp, Quercus robur, Betula spp, Alnus spp and Populus tremula. There was an increase in the proportion of Picea spp, and a decrease in the proportion of Pinus spp since 1927. The proportion of deciduous trees also increased but not as much. Interestingly, the proportion of Betula spp had fallen since 1927, but the now larger proportion of young trees indicate that the proportion of Betula spp is increasing again. Q. robur, P. tremula, Picea spp and Pinus spp also had a larger proportion of younger trees in the more recent period. There was a significantly higher proportion of tall trees during the latter period for all tree species. For growth rate in width, the results varied among tree species. For Betula spp it had decreased and for Q. robur it had increased. For the coniferous trees there was hardly any difference between the periods. Overall, the results show that Östergötland's forests have been influenced by forestry and increased ungulate populations since the first inventory 1927 and that there have been changes in the composition of the forests. / Människan har länge påverkat de svenska skogarna. Sedan 1900-talets början har en hel del förändringar skett inom skogspolitiken, vilket har påverkat bland annat skogarnas ålder- och storleksfördelning, samt trädslagsfördelning. Genom att använda information från olika typer av historiska data kan vi öka vår förståelse om hur skogar såg ut förr och vad som har påverkat dem. Denna studie undersökte skillnader i tillväxt, höjd-och diameterfördelning, samt åldersfördelning i Östergötlands skogar från år 1927 till perioden 1983–2017, med hjälp av data från riksskogstaxeringen. Sex trädslag användes; gran (Picea spp), tall (Pinus spp), ek (Quercus robur), björk (Betula spp), al (Alnus spp) och asp (Populus tremula). En kraftig ökning av andelen gran hade skett sedan 1927, medan andelen tall hade minskat. För lövträden hade det endast skett en liten ökning i andel. Intressant var också att andelen björk hade minskat sedan 1927 och att andelen björkar yngre än 50 år var större den senare perioden, vilket indikerar att björkens andel kan vara på väg att öka igen. Ek, asp, gran och tall hade också en större andel yngre träd under den senare perioden. För alla trädslag fanns det en betydligt mindre andel höga träd år 1927. För årlig tillväxt i bredd varierade resultaten trädslagen emellan. För björk hade årsringsbredden minskat sedan 1927, medan den hade ökat för ek. För barrträden var det knappt någon skillnad mellan perioderna. Resultaten visar att Östergötlands skogar har påverkats mycket av skogsbruket, samt klövviltsbete, sedan 1927 och att det har skett flera strukturella förändringar i skogarnas sammansättning.