Evapotranspiration dominates the water budget in drylands, exerting important controls not only on the dynamics of water, but also on the amount and distribution of vegetation on a landscape. The spatial and temporal variability of vegetation cover imposes constraints on key ecohydrological processes that feedback to the dynamics of evapotranspiration and, most importantly, its partitioning between direct evaporation and transpiration from plants, one of the most significant ecohydrological challenges. Yet, lacking are systematic evaluations of how variations in woody plant cover--a fundamental vegetation attribute of landscapes that can vary spatially with amount of cover and temporally with leaf phenology-- influence the dynamics of soil microclimate and ultimately the partitioning of evapotranspiration into its components. This study presents the results of field experiments that systematically evaluated the effects of amount of canopy cover and its seasonality in both surface microclimate and soil evaporation. These field observations are complemented by controlled experiments that directly evaluate the relationship between amount of canopy cover and the partitioning of evapotranspiration, with an assessment of its larger-scale implications using a regional land surface-atmosphere model. Finally, this study presents a classroom-adaptation of the evapotranspiration partitioning experiment that was used to effectively translate new scientific concepts and information into k-12 classrooms. Overall, the results from this study provide a comprehensive understanding about the interactive ways in which canopy cover, canopy structure attributes and plant phenology influence soil surface microclimate--characterized by near-ground solar radiation and soil temperature--and soil evaporation. More specifically, the results illustrate how the main control of deciduous-woody vegetation on soil evaporation is the addition of litter to the surface. However, in absence of litter, attributes of woody cover influence soil evaporation variably with season and phenology. Further, The results from this study illustrate how the partitioning of evapotranspiration exhibits a non-linear response to amount of woody canopy cover. Notably, when incorporated into a regional surface-atmosphere model, this non-linearity strongly affects water fluxes, highlighting the potential implications for ecological, hydrological, and atmospheric processes associated with the partitioning of evapotranspiration, providing important insights for natural resource management.

Woody plant encroachment, that is, a substantial increase in the abundance of woody plants in a grassland or savanna, occurs in many parts of the world. It often has large effects on plant and animal populations and communities and on ecosystem properties and processes. However, little is known about the dynamics of woody plant encroachment and how these are affected by soils, by topography, and by the spatial pattern of the vegetation. Encroachment in turn can affect the spatial pattern of the vegetation. Using data from historical aerial photographs, I measured changes in woody plant cover and constructed, parameterized and compared a set of dynamic models of woody plant encroachment in central Texas savannas. These models predicted final woody cover from initial woody cover and the initial spatial configuration of woody plants. Then I incorporated soil and topography into these models to determine their effects. Finally, I examined the effects of encroachment on the spatial pattern of the vegetation. Incorporating negative density dependence in our models improved their fit, demonstrating that encroachment is density-dependent. A function that predicted the formation of new woody patches from a density-independent seed supply also improved the models' performance. The improvement in the models that resulted from incorporating the total length of woody-herbaceous edges confirmed that encroachment in this system occurs in part by the outward expansion of woody patches. The spatial pattern of the vegetation changed during woody plant encroachment. Spatial pattern (measured as degree of fragmentation) often had a non-linear relationship with cover. Furthermore, the spatial heterogeneity in fragmentation, that is, plot-to-plot variation in the degree of fragmentation, also changed during encroachment. Topography and soil type had, in general, little effect the dynamics of woody plant encroachment. Therefore, a relatively simple model of woody plant encroachment provided good predictions of woody cover at the end of the time periods. Other systems experiencing woody plant encroachment, forest succession, or invasion by non-native plants could be modeled using the same approach. / text