Amodal completion enables animals (birds, mammals, but also fish) to perceive partly occluded objects as whole. Most of the studies focusing on the occlusion phenomenon were carried out in a laboratory and were based on either operant conditioning or filial imprinting techniques. This study concentrates on behavior of untrained animals in their natural habitat. Pairwise preferential experiments were used to reveal responses of tits (the great tit, Parus major; the blue tit, Cyanistes caeruleus; the marsh tit, Poecile palustris) to two dummies placed near an experimental and an alternative feeder. The dummies used were the complete dummy of a sparrowhawk and a pigeon, and partly occluded (the lower or upper torso hidden in shrubs) and amputated (only lower or upper torso on the perch) models of sparrowhawks - 15 combinations altogether. The tits considered all variants of torsos to be predators. The great tit and the blue tit perceived the model with occluded lower torso as more dangerous than the one with amputated lower torso. Such discrimination between these torsos requires the ability of amodal completion. The great tit also confirms this ability as it regarded the complete sparrowhawk and the model with occluded lower torso as equally dangerous. In the remaining cases, the number of arrivals...

Pair-wise preferential experiments were used to reveal dangerous predator differentiation and ability of amodal completion by four tit species. Firstly was revealed that the tits{\crq} behaviour towards the feeder was in agreement with predictions of the dynamic risk assessment theory. The presence of any predator at the feeder lowered the number of visits to the feeder. Likewise, the tits evaluated the sparrowhawk as more dangerous than the kestrel because its presence lowered the number of visits more than the kestrel. Secondly, we observed tits{\crq} reactions to both partly occluded and amputated dummy of sparrowhawk in two different treatments (torsos vs. complete dummy of pigeon, torsos vs. complete dummy of sparrowhawk). All birds clearly classified both torsos as ``full-featured{\crqq} predators and kept away of them when pigeon on the second feeder. However, when sparrowhawk was presented on the second feeder, number of visits to amputated predator was higher than to occluded one. Birds risked arriving to ``utter{\crqq} amputated torso while the fear of ``full-featured{\crqq} occluded torso stayed without change when second feeder did not provide safe alternative. Such discrimination between torsos needs ability of amodal completion.

Pair-wise preferential experiments were used to reveal dangerous predator differentiation and ability of amodal completion by four tit species. Firstly was revealed that the tits{\crq} behaviour towards the feeder was in agreement with predictions of the dynamic risk assessment theory. The presence of any predator at the feeder lowered the number of visits to the feeder. Likewise, the tits evaluated the sparrowhawk as more dangerous than the kestrel because its presence lowered the number of visits more than the kestrel. Secondly, we observed tits{\crq} reactions to both partly occluded and amputated dummy of sparrowhawk in two different treatments (torsos vs. complete dummy of pigeon, torsos vs. complete dummy of sparrowhawk). All birds clearly classified both torsos as ``full-featured{\crqq} predators and kept away of them when pigeon on the second feeder. However, when sparrowhawk was presented on the second feeder, number of visits to amputated predator was higher than to occluded one. Birds risked arriving to ``utter{\crqq} amputated torso while the fear of ``full-featured{\crqq} occluded torso stayed without change when second feeder did not provide safe alternative. Such discrimination between torsos needs ability of amodal completion.

<p>Three main theories have been proposed to account for the completion of occluded objects, namely local theories (e.g., Kellman & Shipley, 1991), global theories (e.g., Boselie & Leeuwenberg, 1986), and integrative models of completion (e.g., Sekuler, 1994; van Lier et al., 1994). Here, we investigated age-related changes in the completion of a complex partially occluded object using a prime-matching task. Subjects were shown a prime (global, local, occluded, or no prime) that was followed immediately by two shapes that were judged as being the same or different. In Experiment 1, we tested adults (n = 36/group) at various prime durations (150 - 700 msec) to tap into earlier and later representations of the occluded object. Although the occluded object primed both the global and local shapes at 150 and 500 msec (<em>p</em>s<0.05), only the global shapes were primed at 300 and 700 msec (<em>p</em>s<0.05). Overall, our results are most consistent with integrative models of completion. In Experiment 2, we tested 8-year-olds (n = 20) at a prime duration of 700 msec and 11-year-olds (n = 30/group) at a prime duration of 300 or 700 msec. For 11-year-olds, unlike adults, the occluded object did not significantly prime either the global or local shapes at 300 msec (<em>p</em>>0.50). For both 8- and 11-year-olds, the global, local, and occluded primes did not significantly prime either shape at 700 msec (<em>p</em>s>0.50). Based on the current testing conditions, we found that the perceptual completion process may not be adult-like even at 11 years of age.</p> / Master of Science (MSc)