The rise and demise of mountain belts, caused by growth, modification, or removal of the continental lithosphere are fundamental processes that influence almost all Earth systems. Understanding the nature, timing, and significance of active processes in the creation and evolution of modern mountain belts is challenged by a lack of middle crustal and lower crustal exposures. Analogues can be found in ancient orogens, whose deeply eroded roots offer a window into deeper processes, yet this record is complicated by overprinting events and complex deformational histories. Research presented herein constrains the tectonic history of multistage Appalachian Orogen, type locality of the Wilson cycle. Data-driven analysis of newly assembled geochronologic, geochemical, and geothermobarometric databases are synthesized with structural fabrics and geophysical imaging to constrain the timing and nature of crustal thickening and thinning events. Results identify a two-stage crustal thickening history in the dominant Acadian Orogeny and suggest the existence of a high elevation, low relief orogenic plateau. This plateau, the Acadian altiplano, formed in central and southern New England by ca. 380 Ma and exited for at least 50 m.y. until underwent orogen parallel collapse ca. 330-310 Ma. Collapse of the plateau likely formed the geophysically observed 12-15 km offset in Moho depth in western New England, and implies that the step has existed for ca. 300 m.y. These data constrain a four-dimensional record of crustal evolution over a period exceeding 100 m.y. Recognition of the Acadian altiplano may have important implications for the genesis of critical Li deposits, paleoclimate, and evolution of the Appalachian basin. Further, present a region that may provide an analogue for studying mid-crustal processes such as partial melting, ductile flow, and plutonism underneath modern plateaus.

Island biogeography theory predicts that the patterns and distributions of spatially isolated populations are governed by large scale processes. The high elevations forests in the Southern Appalachians represent a series of naturally fragmented islands that harbor many isolated populations of species at the southern limits of their range. Understanding the governing forces of population dynamics in this region will enhance the probability of species persistence in the face of threats such as global warming and human development. We surveyed bird populations across multiple elevations in Virginia and combined this with a multi-scale habitat analysis to determine influences of species presence and species richness. We detected 101 species across the elevation gradient, including 12 species with special conservation status and ten species whose presence increased with increasing elevation. These ten elevation sensitive species responded to habitat variables at both the microhabitat and landscape scale, with species-specific patterns of habitat variable correlation emerging. Habitat type was least effective in predicting species presence for any elevation sensitive species. Species richness declined over the elevation gradient until the highest elevations, where this trend reversed and richness began to increase. This pattern was driven by an increase in short-distance migrants beginning at mid-elevations, which ultimately overpowered a corresponding decrease in long-distance migrants beginning at similar elevations. Habitat analysis linked these patterns to a preference of short-distance migrants for smaller, more isolated non-forested patches, and a historical lack of persistence for long-distance migrants. Conservation and management decisions for the region should focus on a multi-scale approach that preserves all habitat types for continued species presence and high species richness, although the persistence of particular elevation sensitive species is compounded by unique species-habitat relationships and the perception of islands as species-specific. Continued monitoring of these fragmented populations in light of both short- and long-term threats which span multiple scales of influence will maintain high species richness and ensure the persistence of crucial breeding habitat. / Master of Science

Two threats to bats are especially pervasive in the central Appalachian Mountains of the eastern United States: a fungal disease called White-nose Syndrome (WNS) and wind energy development. White-nose Syndrome has caused the death of millions of bats in North America, and multiple hibernating bat species are affected in the central Appalachians. Wind energy is one of the most rapidly-growing energy sources in eastern United States, and bats are often killed when they fly near wind turbines. Fatality rates at wind turbines is highest in bat species that migrate instead of hibernate. There is limited data on bats during the autumn and spring seasons in the central Appalachian Mountains, and the impacts of WNS and wind energy development may be exacerbated during these seasons. Therefore, I sought to determine patterns and drivers of activity for hibernating bat species during autumn and spring around hibernacula. Similarly, I set out to determine patterns and drivers of activity for migratory bat species during autumn and spring along mountain ridgelines in the central Appalachians. Lastly, I searched for evidence of potential WNS-induced changes in the summer ecology of the once common northern long eared bat. This study can help elucidate patterns of bat activity during largely understudied seasons. Furthermore, it can provide useful information needed by land managers to implement actions that could help alleviate and/or avoid potential additive negative impacts on bat species with existing conservation concerns. / MS

Calcite c-axes and e-twin plane orientations were measured in both matrix cements and younger fracture fills from late Cambrian to Middle Ordovician age limestone samples taken from the NW and SE limbs of the Massanutten Syncline, located within the North Mountain thrust sheet. Paleostress magnitude estimates using the Rowe and Rutter (1990) twin density technique indicate a differential stress of 240±31 MPa for samples collected from both limbs of the syncline. Three distinct patterns of paleostress orientations (compression directions) have been detected in the samples; each pattern is observed on both the NW and SE limbs of the syncline. The first pattern, exhibited by calcite grains cementing late fractures, is characterized by a maximum of compression axes oriented sub-perpendicular to bedding possibly indicating either thrust sheet loading or stress refraction associated with folding. Samples in which calcite grains from both fracture fills and earlier matrix cements were measured are characterized by a bimodal distribution of compression axes—the first point maximum being oriented sub-perpendicular to bedding, the second maximum placing compression directions at low to moderate angles to bedding. Restoration of bedding to horizontal results in this second set of compression axes plunging to either the SE or NW, sub-parallel to the regional thrust transport direction. The third pattern, originating from early cements, places compression directions plunging to the NE-SW at angles which are sub-parallel to bedding. These compression directions do not seem to correlate with major tectonic movements or thrust sheet loading and may reflect stresses associated with either movement over lateral ramps or oblique thrusting. / Master of Science

A series of anthropogenic disturbance conditions have altered the vegetation of the southern Appalachians during the past 200-years. The objective of this research was to identify the nature and timing of these vegetation changes in order to better understand the underlying causes. A total of 304 land surveys were collected for a small agricultural watershed from early settlement in 1787 through to the present day. Witness corners recorded tree species, shrubs, stumps, snags and non vegetative markers. Types of witness corners were tallied and tested for shifts in frequency across time periods. Tree species were also classified by silvical characteristics including sprouting capability, shade tolerance, and seed type and these groupings were tested for shifts in frequency across time periods. Landform bias of the witness corners was tested using references contained in the surveys. Results showed significant shifts in white oak (Quercus alba L.), chestnut (Castanea dentate Marsh. Borkh.), chestnut oak (Quercus prinus Wild.), black oak (Quercus velutina Lam.), red oak(Quercus rubra L.), black locust (Robinia pseudoacacia L.), yellow poplar (Liriodendron tulipifera L.), and scarlet oak (Quercus coccinea Muenchh.). The central change was a steady decline in white oak, probably due to the absence of fire and changes in soil properties. Chestnut replaced white oak as the dominant species, but was removed by chestnut blight in the 1930's. Sprouting capability appeared to be the most important silvical characteristic across all species. / Master of Science

Fire occurrence was widespread in the central Appalachians pre-European settlement due to Native American ignition and occasional lightning strikes, and continued through European settlement. During this time, low to mixed severity burns supported a suite of ecological communities that were fire adapted. In the mid-20th century, the frequency and intensity of fire decreased regionally, resulting in profound forest composition shifts. Land managers now are prioritizing prescribed fire as a restoration tool in current and transitioning fire dependent communities. However, it is unclear how the re-introduction of fire will affect bat community assemblages, particularly after the severe White-nose Syndrome related population declines of many cave-hibernating bat species. To address this concern we used acoustic detectors to sample bat activity levels in burned and unburned environments to examine habitat and temporal effects of fire on bat species in a repeatedly burned landscape. We found evidence for weak positive fire effects on the northern long-eared bat, Indiana bat, little brown bat, big brown bat/silver-haired bat group, high frequency phonic group, and total bat activity. Temporal effects of fire were only apparent for the big brown bat, where we observed a negative relationship between activity and time since fire. Additionally, historic wildfires may offer a suitable surrogate to assess long-term burn impacts on bats, which in turn can be used to better inform bat and prescribed fire relationships. To examine effects of historic fire on bats, we assessed bat presence using acoustic detections at 16 paired burned and unburned forest stands in Shenandoah National Park. Overall, we found few or mostly equivocal relationships of bat occupancy across species relative to burn condition or time since fire at SNP, indicating there is little evidence to support the concept that fire has a significant ecological effect on bats in this portion of the central Appalachians. Riparian areas are particularly important for bats, and serve as foraging and drinking areas, roost sites, and travel corridors. Because fire impacts dry upland and mesic riparian areas differently, is possible that fire will impact bats differently in burned and riparian habitats. To examine fire effects on bats in riparian and upland habitats, we used paired sampling to monitor bat activity in burned, unburned, riparian, and non-riparian areas. Burn and riparian variables had empirical support to explain activity of all bat species. However, coefficients for these species were small and confidence intervals overlapped zero indicating that differences between habitat configurations were marginal. Our results suggest bats have somewhat species-specific responses to fire that differ between upland and riparian habitats, but that large landscape level prescribed fire has a slightly positive to neutral impact on all bats species identified in at our study site post-fire suppression. / Master of Science

The high-elevation forests of the Southern Appalachian Mountains have been impacted and rearranged by a tiny introduced pest from Europe, known as the Balsam Woolly Adelgid (Adelges piceaea), creating a concern for conservation. Breeding bird censuses, along with botanical surveys, have been conducted periodically on an established 60-acre plot since 1967 on the virgin forested slopes of Mount Guyot, Great Smoky Mountains National Park, with the last census being completed in 2015. Breeding bird populations are shown to rise and fall in response to the forest’s changes over the last 48 years. Comparisons are made between all studies in order to assess how bird populations are being affected by the changed forest dynamics.

The stratification of bird species along elevational gradients is widely reported, with montane bird communities typically characterized by distinctive species occurring in relatively small and isolated populations; as such, these species are the subject of considerable interest to ecologists and conservationists. The stratification of species along elevation is largely attributed to compressed climatic zonation. Recent evidence that bird species are shifting up in elevation has fueled speculation that these species are tracking their climactic niches in response to climate change. However, there is also evidence plant communities are shifting in elevation, presenting a potential additional mechanism explaining changes observed in the bird community. Uncertainty as to the degree to which climate directly influences bird abundance versus the degree to which climate indirectly influences bird abundance via habitat composition and structure represents a key impediment to understanding the ecology of these species in montane environments. To address this question I measured species abundance, habitat characteristics, and temperature at 150 survey points located along 15 elevational transects in the Presidential Mountains of New Hampshire in the summers of 2014 and 2015. I used N-mixture models to correct for imperfect detection of species and structural equation models, incorporating abundance, habitat, temperature, and precipitation derived from a downscaled regional dataset to assign variation to the direct and indirect effects of climate upon birds. Analysis of 21 species revealed species-specific patterns on how climate exerts direct effects and indirect effects mediated by forest composition and structure on bird abundance. This work represents an important contribution to the ecological understanding of the pathways by which climate influences bird abundance. Finding that 62% of species experience both direct and indirect effects of climate, with 62% experiencing stronger direct than indirect effects, these results underscore the vulnerability of these species to climate change. With 81% of species found to experience indirect effects of climate via forests, these findings indicate great conservation value of maintaining forest habitat amidst climate change. Overall, this information will facilitate the refinement of predictive models of the abundance of montane bird species and represents an approach that will advance future investigations of climate effects in the Northern Appalachians and other systems.

The health of the northern hardwood forest in the Southern Appalachian Mountains of Tennessee, North Carolina, and Virginia has recently gained attention from the media and environmental stakeholders. This project was designed to examine concentrations of metals, including lead, aluminum, and calcium in growth rings of an important northern hardwood species, American beech (Fagus grandifolia Ehrh.) at Mount Rogers and Whitetop Mountain, Virginia. Dominant and codominant trees were sampled from sixteen research plots located at two different elevations. Samples were crossdated, divided into sections of ten-year periods, and analyzed using atomic absorption spectroscopy. Concentrations of metals were negatively correlated with ring width. Elevation and aspect were significantly associated with the concentration of the cations analyzed. Both lead and calcium concentrations increased during the 1860s and again during the mid-1900s. This increase in lead and calcium may suggest the mobilization of these elements in soil.

Ever since the first plate-tectonic model for the Appalachians was proposed, the Laurentian margin has been interpreted as having experienced a collision-related dynamo-thermal event during the Middle Ordovician Taconic orogeny. In the western Newfoundland Appalachians, evidence for this collision is well-preserved in the Dashwoods subzone. Nevertheless, rocks of the neighbouring Corner Brook Lake block (CBLB), which is located in the heart of the Laurentian realm, did not show evidence for such an event. Instead, it was affected by Early Silurian Salinic deformation and associated peak metamorphism. Even though this difference in Early Palaeozoic tectonic history between the Dashwoods and the CBLB is widely known, it has not been satisfactorily explained. To better understand the Early Palaeozoic history of the region, in particular to test and better explain the lack of a Taconic dynamo-thermal event in the CBLB, field mapping, microscopic work, and U-Pb and 40Ar/39Ar geochronological studies were undertaken in the western and northern part of the Dashwoods subzone, and in the southern part of the CBLB. In addition, the kinematic history of the Baie Verte-Brompton Line - Cabot Fault Zone (BCZ), the tectonic zone that separates the two unique tectonic fragments, was studied. The western and northern parts of the Dashwoods subzone contain variably foliated igneous units of Middle Ordovician age (ca. 460 Ma) that are associated with the regionally voluminous Notre Dame continental arc. A ca. 455 Ma conjugate set of late syn-tectonic pegmatite dykes in the BCZ demonstrates a dextral sense of shear along the BCZ (DBCZ-1) during the Late Ordovician to earliest Silurian, and constrains the minimum age of the main phase of ductile deformation in the Dashwoods subzone. The fault-bounded CBLB has been affected by a single west-vergent deformational event, constrained between ca. 434 and ca. 427 Ma. More importantly, no evidence – neither petrographic nor geochronological – is present that would indicate that the CBLB was affected by a significant Taconic dynamo-thermal event. Hence, the CBLB and Dashwoods could not have been juxtaposed until after the late Early Silurian. Furthermore, the basement to the CBLB is devoid of any Grenville (sensu lato; ca. 1.0-1.3 Ga) U-Pb ages, which is in sharp contrast with crystalline basement elsewhere in the region, such as the Long Range Inlier. Therefore, it is highly unlikely that the CBLB represents the para-autochthonous leading edge of the Laurentian craton in the Newfoundland Appalachians, as commonly accepted. The CBLB is interpreted as a suspect terrane that has moved over 500 km parallel to the strike of the orogen. Docking to the external Humber Zone is likely to have occurred during the Early Silurian. Final juxtaposition with the Dashwoods took place after the late Early Silurian (post-Salinic) as a result of protracted dextral movement along the BCZ (DBCZ-2 and DBCZ-5). Current tectonic models for the Newfoundland Appalachians mainly focus on well-documented Early Palaeozoic orthogonal convergence of various terranes with the Laurentian margin, but large-scale orogen-parallel movements have rarely been considered. The possibility of large-scale strike-slip tectonics documented here, in addition to the convergent motions, may have significant implications for the tectonic interpretation of the Early Palaeozoic evolution of the Newfoundland Appalachians.