Livestock grazing impacts more land than any other use. Yet knowledge of grazing impacts on native fauna is scarce. This thesis takes a predictive approach to investigating the effects of livestock grazing on Australian woodland birds, employing some novel methodological approaches and experimental designs. These include methods of analysis to handle zero-inflated data and the application of Bayesian statistics to analyse predictions based on expert opinion. The experimental designs have enabled impacts of grazing to be separated from the frequently confounding effects of other disturbances, and to consider the effect of grazing on habitat condition in the context of different surrounding land uses. A distinguishing feature of many datasets is their tendency to contain a large proportion of zero values. It can be difficult to extract ecological relationships from these datasets if we do not consider how these zeros arose and how to model them. Recent developments in modelling zero-inflated data are tested with the aim of making such methods more accessible to mainstream ecology. Through practical examples, we demonstrate how not accounting for zero-inflation can reduce our ability to detect relationships in ecological data and at worst lead to incorrect inference. The impact of grazing on birds was first examined through the elicitation of a priori predictions from 20 Australian ecologists. This expert knowledge was then used to inform a statistical model using Bayesian methods. The addition of expert data through priors in our model strengthened results under at least one grazing level for all but one bird species examined. This study highlights that in fields where there is extensive expert knowledge, yet little published data, the use of expert information as priors for ecological models is a cost effective way of making more confident predictions about the effect of management on biodiversity. A second set of a priori predictions were formulated using a mechanistic approach. Habitat structure is a major determinant of bird species diversity and livestock grazing is one mechanism by which structure is altered. Using available information on the vegetation strata utilised by each species for foraging and the strata most affected by grazing, predictions of the impact of grazing on each bird species were formulated. We found that foraging height preference was a good predictor of species’ susceptibility to grazing. This approach is a starting point for more complex predictive models, and avoids the circularity of post hoc interpretation of impact data. The confounding of grazing with tree clearing was addressed by examining the impact of pastoral management on birds in sub-tropical grassy eucalypt woodland in Southeast Queensland, where land management practices have made it possible to disentangle these effects. Changes in bird species indices were recorded across woodland and riparian habitats with and without trees across three levels of grazing, replicated over space and time. Tree removal had a dramatic influence on 78% of the bird fauna. 65% of species responded significantly to changes in grazing level and the abundance of 42% of species varied significantly with habitat, level of clearing and grazing. The impact of grazing on birds was most severe in riparian habitat. Finally, the extent to which landscape context and local habitat characteristics influence bird assemblages of riparian habitats in grazed landscapes is addressed. Over 80% of bird species responded significantly to changes in local riparian habitat characteristics regardless of context, while over 50% of species were significantly influenced by landscape context. The influence of landscape context increased as the surrounding landuse became more intensive. These results suggest that it is not enough to conserve riparian habitats alone but conservation and restoration plans must consider landscape context. The ability to predict which bird species will be most affected by grazing will facilitate the transformation of this industry into one that is both profitable and ecologically sustainable. Results from this thesis suggest that any level of commercial grazing is detrimental to some woodland birds. Habitats with high levels of grazing support a species-poor bird assemblage dominated by birds that are increasing nationally. However, provided trees are not cleared and landscape context is not intensively used, a rich and abundant bird fauna can coexist with moderate levels of grazing, including iconic woodland birds which are declining elsewhere in Australia.

Because of the scarcity of information for bird communities at the major mainland rivers of southeastern Alaska, the main objective of this study was to provide baseline information including distribution, status, and habitat associations of breeding birds. I conducted a meta-analysis of all known reports (including the current study) conducted at major mainland rivers during the breeding season. I described bird species composition, distribution, abundance estimates, status, habitat associations, and guild membership for all birds recorded at 11 major mainland rivers. Based on incidental observations, 170 species were recorded by all studies. Of these, 134 species were known or suspected to breed, accounting for 50% of all birds known from Alaska and 80% of all birds known from southeastern Alaska. In addition, I provided information on species of management concern as well as management implications and recommendations. I used point counts to survey birds within deciduous riparian vegetation at 6 major mainland rivers during 2000-2002. I compared bird species composition, abundance, richness, and diversity among four main vegetation types of deciduous riparian vegetation: shrubland, young deciduous forest, mature deciduous forest, and mixed deciduous-coniferous forest. Species richness was similar among all habitat types; however, relative abundance and diversity of birds was highest in mixed forest stands. Mature forests had the greatest number of species associated with the Canadian interior. I also used point counts to compare bird species composition, abundance, richness, and diversity among 6 major mainland rivers consisting of three trans-mountain and three coastal rivers. Latitude, connectivity, and availability of mature and mixed forests were the major factors thought to cause differences in bird communities among rivers. Contrary to our predictions, coastal rivers had higher bird species richness, diversity, point abundance , and point richness than trans-mountain rivers. Of the 10 species associated with the Canadian interior recorded during point counts, 8 occurred at both trans-mountain and coastal rivers.

Macroecology is the study of relationships between organisms and the environment at large spatial and temporal scales. This field of research examines patterns in species abundance, distribution and diversity. Understanding patterns in species distribution and richness can contribute significantly to our knowledge of community assembly and macroecological patterns, as well as to the effective conservation of threatened species and habitats. Although there have been a plethora of studies on birds in India over the years, there is a critical need to accurately delineate species distributions and understand patterns of richness. The focus of this study was to understand the factors (abiotic and biotic) that influence the distribution and composition of bird species in the Western Ghats, as well as to explore patterns in their geographic range sizes. The objectives of this study were addressed at the scale of the entire Western Ghats using a combination of field surveys, secondary data collection and species distribution modeling. The specific approaches to address these questions and the findings are outlined below. Chapter 2: Bird species in the Western Ghats – Patterns in composition and richness Fine-scale data on species presence and abundance are essential for exploring patterns in species distribution and richness. Despite the fact that birds have been extensively studied in the Western Ghats, systematic data collection and compilation of information over the entire mountain range has not been carried out, especially for the purpose of testing macroecological questions. This chapter describes patterns in bird species presence, abundance, composition and richness within the Western Ghats. The study area, site selection protocol and the sampling technique have also been described in detail. This dataset establishes a baseline of information about birds in the Western Ghats and subsets of this larger dataset will be used to address various questions in the following chapters. Chapter 3: Predicting bird species distribution in the Western Ghats Detailed knowledge of species’ ecological and geographical distributions is fundamental for conservation, as well as for understanding ecological and evolutionary determinants of spatial patterns of biodiversity. However, occurrence data for a vast majority of species are sparse, resulting in information about species distributions that is inadequate for many purposes. Species distribution models attempt to provide detailed predictions of distributions by relating presence or abundance of species to environmental predictors. In this chapter, we describe the usage of Maxent, a species distribution modelling technique based on presence-only data, to predict the distributions of bird species within the Western Ghats. For this purpose, we put together primary locations of bird species presence along with a published dataset. Using a number of important environmental layers, predicted species distribution maps were derived for 98 bird species, including 13 endemics, in the Western Ghats. Additionally, we calculated predicted range sizes for each of these species and obtained percentage contributions of important environmental predictors to each species’ distribution. This is the first study to develop species distribution models for bird species within the Western Ghats. Chapter 4: Patterns of range size among bird species Understanding large-scale patterns of variation in species geographic range size is fundamental to questions in macroecology and conservation biology. In general, range is believed to be influenced by a combination of environmental factors, evolutionary history and biotic interactions, mediated by species specific traits. These patterns need to be examined even for well-studied taxa like birds, especially within biodiversity hotspots faced by persistent degradation due to anthropogenic activities such as the Western Ghats. In this chapter, we use a dataset of 98 bird species within the Western Ghats to examine trends in range sizes, measured as latitudinal extent of occurrence and predicted range size from species distribution models. We show a significant relationship between latitude and range size for these bird species, supporting Rapoport’s rule. As far as we know, this relationship has never been tested at such low latitudes for birds. We also find that species traits such as body size, mean abundance and diet do not seem to show any discernable effect on patterns of range size. Additionally, we found that widely-used bird species range maps (in this case, from BirdLife International) are inaccurate representations of species ranges in comparison to the predicted species distribution maps that were derived in the previous chapter. We quantitatively demonstrated that these expert-drawn maps need to re-evaluated, especially since they are used to make conservation decisions. This is the first study to quantify species range sizes of birds within the Western Ghats and assess such range maps that are used to determine conservation status of species. Chapter 5: Environmental predictors of bird species distribution One of the major goals in ecology is to understand patterns and processes that determine species diversity. The drivers of global species richness gradients have been studied, especially in the case of birds, in terms of contemporary and historical factors. Such broad scale processes may not always reflect the processes affecting richness and distribution at smaller scales. Therefore, understanding the factors that influence individual species distributions is the first step towards this larger goal. In this chapter, we examined the environmental predictors that contributed to the predicted distribution of bird species observed in the Western Ghats, using the variable importance contribution values derived in Chapter 3. We found that a large proportion of the 98 bird species studied were influenced by normalized differential vegetation index, annual precipitation and elevation. The predictors did not differ among birds of different diet guilds and body size classes. Using Prinicipal components analysis, we observed that all 98 bird species are spread out across the environmental ordination space depicted by the PC axes 1 and 2. These axes are governed by measures of habitat heterogeneity and water-energy related variables, consistent with other tropical studies. The insectivorous guild seemed to occupy a variety of environmental niches across this space and other guilds seemed to be nested within the insectivorous guild. Similarly, larger sized birds were spread across the entire environmental ordination space, with species of smaller sizes nested within. This is the first step in trying to understand environmental predictors acting on birds in the Western Ghats. Further detailed studies need to be carried out to come to definite conclusions. Chapter 6: Relative roles of floristics and vegetation structure on bird species composition On the basis of the hierarchical model of habitat selection, it is known that birds select suitable habitats based on vegetation structure (physiognomy) at coarse biogeographic scales, and plant species composition (floristics) at more local scales. This chapter examines the relative influence of tree species composition and vegetation structure on bird species composition in the Western Ghats. These relationships were specifically assessed across the entire Western Ghats, within regions of the Western Ghats as well as within specific forest types. We found that floristics had a strong association with bird species composition across the Western Ghats and within evergreen and mixed deciduous habitat types. This association seems to be independent of the structural variation in the region. There was a decrease in association strength from the southern to the northern Western Ghats, in terms of both floristics and structure. We did not find an association between vegetation structure and insectivore composition, whereas phytophage composition did show a stronger association with floristics than structure. This is the first study at the scale of the entire Western Ghats to test the relative roles of floristics and vegetation structure. Taken as a whole, this dissertation examines large-scale macroecological questions regarding species distribution, range size and patterns of composition using primary data at the scale of the Western Ghats. The findings of this study have established a foundation that will help further our understanding of species distribution and richness in the Western Ghats, and aid in the decision making for conservation strategies in the future.