To show how predaceous fish and dragonflies affect benthic community structure, we conducted enclosure experiments in the littoral zone of Bays Mountain Lake, Tennessee. A 'natural' benthic assemblage was subjected to all combinations of 2 densities of 3 predator treatments - small sunfish, 0 or 4/m2; large dragonfly larvae, 0 or 15/m2; and dragonfly eggs at 2 densities, 90 or 900/m2. Treatments were assigned randomly in each of 6 spatiotemporal blocks. Net effects of predation over 4 mo show that small sunfish had 'strong' effects (>50% reduction of densities) on triclads, large daphnid cladocerans, and snails, and 'moderate' effects (50% > reduction > 25%) on small dragonflies and ostracods; all these effects were statistically significant, except the one on ostracods. Large dragonflies had moderate non-significant effects on triclads and ostracods. There were no significant increases in prey density associated with fish predation; but chydorid cladocerans and midge larvae showed 'weak' non-significant increases that might be caused by 'indirect effects' of fish predation on invertebrate predators or grazers. There were only 2 significant interaction terms indicative of 'higher-order interactions': Fish X Dragonfly on the large daphnid Simocephalus, and Fish X Egg Density on snails, which were associated with relatively low probabilities. In both cases, the net effect of dragonflies was to reduce prey densities more when fish were present.