This study aims to investigate the preservice teachers&rsquo / perceptions about online communication in blended learning. The study tries to understand strengths and weaknesses of online communication from the learners&rsquo / point of view. To reach this aim, four research questions are asked: (1) What are the learners&rsquo / perceptions about the online synchronous communication tool? (2) What are the learners&rsquo / perceptions about web-based support? (3) What are the learners&rsquo / perceptions about collaboration with online communication? (4) What are the learners&rsquo / perceptions about the roles of the instructor at blended learning as (a) administrator, (b) facilitator, (c) technician, and (d) evaluator? To answer the research questions, a case study in line with action research design was conducted. An undergraduate course in blended learning form was selected as a case and both synchronous and asynchronous communication tools were utilized throughout the semester. At the end of the semester, data about perceptions was collected via four questionnaires and interviews with the students. Both qualitative and quantitative results showed that the online communication facilities in the case were perceived to be adequate by the students. Yet, the students reflected diverse thoughts about preference of communication modalities in synchronous communication. Moreover, the value of asynchronous communication was pointed out. It is concluded that communication needs, communication partner and other contextual factor have impact on selection of communication modalities.