Competition between native perennial grasses and Bromus tectorum has been studied for many years. Recently, soil nutrients have been immobilized in an effort to preferentially inhibit growth of B. tectorum relative to native species. Most of this work has focused on soil N, but interactions with soil P are less studied and may be important as well. Additionally, although competitive effects and nutrient responses of several Centaurea spp. are well documented, data are lacking on the competitive effects that Centaurea virgata exerts on the establishment of native and exotic communities. A field experiment and a greenhouse experiment were conducted to evaluate influence of N and P on the native perennial bunchgrass Pseudoroegneria spicata in competition with B. tectorum and C. virgata. Chapter 2 describes the effects of nutrient immobilization and subsequent 2 x 2 factorial additions of N and P. Treatments were applied to transplanted individuals of the study species in monocultures and mixtures. Analysis of variance (ANOVA) was performed on shoot dry mass, soil water content, and physiological responses including photosynthetic rate, stomatal conductance, and transpiration. Results suggest that B. tectorum may be P-limited in certain circumstances, and that physiological activity of all species is subject to interactions between available N and P. In Chapter 3, I assess response of the study species to 2 x 2 factorial additions of N and P on a very low-nutrient soil in a greenhouse. Relative effects of different species mixtures on P. spicata are reported. I report ANOVA results of tiller number, leaf number, specific leaf area, shoot dry mass, root dry mass, and water addition. Results indicate that N and P co-limit P. spicata and B. tectorum, and to a lesser extent, C. virgata. B. tectorum showed the highest percentage response to fertilization. Land managers may be more successful establishing native grasses if they can minimize soil fertility.

The objectives of this study were to determine the effects of two forms of polyacrylamide (PAM) conditioners (Cross-linked and Non-cross-linked PAM) on evaporation, saturated hydraulic conductivity, water retention, crust and crack formation of soils, seed germination, and seedling and tubeling growth. The two PAM conditioners, 0.2% concentration by weight, were mixed with seven soils of different textures (sandy loam, silt, silty clay loam, silt loam, fine sand, medium sand, and coarse sand) to investigate the effects on evaporation, saturated hydraulic conductivity, and water retention. Soil samples of different textures were brought to field capacity and placed in a growth chamber for two weeks to measure evaporation under a controlled environment. A second experiment was carried out in the field to determine the effects of the two PAM conditioners on seedling emergence of crested wheatgrass, Agropyron desertorum, as well as on soil cracking, penetrometer resistance, and soil moisture. The two PAMs were mixed with a silt loam Xerollic Calciorthid at 0.2% concentration by weight. Seedling emergence was monitored directly for two weeks. Soil moisture was measured by TDR. Cracking was described by photographic means. Penetrometer resistance was measured by a hand-held. penetrometer. The third experiment was also carried out in the field, using the same soil texture as in experiment 2, to investigate the effects of the two PAMs on soil moisture at depths between 25 to 45 cm and on sagebrush (Artemisia tridentata) growth. Evaporation was found to be significantly lower in the fine-textured controls than under the two PAM treatments. The sandy loam and sandy soils experienced significantly higher evaporation from the controls. The two PAM conditioners significantly reduced saturated hydraulic conductivity on all soil textures. Water retention increased in the PAM-treated textures at the matric potential range used (0.0, 0.05, 0.1, 1.5 MPa) • The PAM application also did not improve grass seedling emergence or improve soil moisture, and did not have any significant affects on sagebrush growth. Larger cracks were found in the two plots treated with PAM than the controls. Lower penetrometer resistance occurred in the two PAM treatments compared to the untreated control. From this study, it can be concluded that the application of PAM conditioners, at relatively high concentrations used, could be more viable on sandy textures. Other researchers are advised to try lower application rates than used here, particularly with finer textured soils.