Due to the discrepancy among series-connected cells in a lead-acid battery pack, the restored capacities may not be the same during the charging/discharging processes. Through repeated charging and discharging, one can find that the unbalance of individual series-connected battery cell of lead-acid battery is getting worse, because those cells of poor state of charge may have negative voltage, causing reverse charging from the other cells. This abnormal operation results in not only more energy consumption, but also the acceleration of the battery-cell aging and the remarkable reduction of its lifetime. For the purpose of investigating this problem, the operating characteristics of each cell is under study. The battery cells are purposely charged by different floating voltages and discharged by different cutoff voltages. The investigation results show that the lifecycles of battery cells are affected more significantly by floating charging voltage than by the discharging cutoff voltage. It is also found that a long time reverse charging has a destructive effect on the battery cells.