published_or_final_version / Geography and Geology / Doctoral / Doctor of Philosophy

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In dryland environments, the availability of soil moisture is the primary control on plant species’ distributions. In the sandhill regions of the northern Great Plains, vegetation establishment has transformed highly mobile, desert-like dune fields into stabilized landscapes covered by mixed-grassland prairie. This study examines how dune stabilization has modified the spatiotemporal distribution of soil moisture resources. An ergodic (space-for-time) approach was used, comparing soil moisture dynamics on active and vegetation-stabilized dunes in the Bigstick Sand Hills of southwestern Saskatchewan. Results indicate that while dune stabilization has enhanced near-surface soil moisture availability, deeper profile soil moisture recharge is reduced. Through better understanding how vegetation has modified soil moisture dynamics in stabilizing sandhill regions, better management practices may be implemented to maintain water resource availability and ecosystem health. / xii, 97 leaves : ill., maps ; 29 cm

Every species has a limit to its geographic distribution. The problem is that we don’t really know why. Classical explanations propose that range limits occur where the ecological conditions overwhelm adaptation and populations are no longer self-sustaining. It is generally expected that population abundance and fitness decline towards range margins across a gradient of declining habitat quality. Current evolutionary and theoretical explanations of range limits predict that this geographic pattern of demography will result in genetic constraints in marginal populations, such that range expansion is thwarted by reduced evolutionary potential. In this thesis, I tested the key assumptions and predictions of range limit theory through an empirical evaluation of two coastal dune, endemic plants; Camissonia cheiranthifolia (Spreng.) Raim. (Onagraceae) and Abronia umbellata L. (Nyctaginaceae). In geographic wide surveys of a large proportion of populations across both species’ ranges, neither species exhibited declining abundance or performance towards its range limits. Central populations of C. cheiranthifolia tended to have a higher production of seeds per unit area than marginal populations. Although this pattern demonstrated the potential for gene flow from central sites to swamp selection in marginal sites, results from a transplant experiment suggested that this was unlikely. Experimental populations of C. cheiranthifolia originating from ≥ 675 km south of the northern range limit exhibited similar levels of fitness when planted at the range margin. Along a 200 km transect across the limit, and in contrast to expectations, fitness of all populations increased towards the limit and generally remained high beyond the limit. Individuals from all populations reproduced and matured fruit beyond the limit, suggesting that if individuals dispersed beyond the limit that populations would establish. The species’ abrupt distributional limit also did not correspond to an abrupt shift in ecological conditions, despite the association of fitness with plant community and microhabitat variables. Overall, ecological tolerances to fitness or niche-based explanations to range limits do not adequately describe the distributions of either species. Constraints on dispersal rate, the influence of anthropogenic factors on habitat dispersion and limited genetic variability for fitness related traits are addressed. / Thesis (Ph.D, Biology) -- Queen's University, 2007-05-13 16:45:44.456

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The erosion of headwater wetlands in the Sand River catchment, in the lowveld of north-eastern South Africa has led to a focus on their rehabilitation, both for livelihood security for those that use them for subsistence agriculture, as well as for provision of streamflow regulation services for the Sand River itself. One such wetland, the Craigieburn-Manalana itself undergoing severe erosion was subject to technical rehabilitation using concrete weirs and gabion dams to stabilize the erosion gullies during 2007. Through a series of papers the research discussed in this thesis examined the response of the wetland?s hydrodynamics to the implementation of these measures. Through the installation of a network of hydrometric apparatus the research has shown that the wetlands hydrology is largely controlled by the presence of both horizontal and vertical clay aquicludes within a hydraulically conductive sandy matrix. The sequence of these aquicludes had allowed for artesian phreatic surface phenomena identified in a relatively hydrologically intact region of the wetland. The gully erosion had initiated hydraulic drawdown of the wetland?s water table leading to the desiccation of the system. The construction of a buttress weir within the erosion gully had restored the wetlands hydrodynamics to that typical of conditions upstream of a clay-plug. The research also explored the role that clay plays in terms of controlling the wetland?s hydro-geomorphic setting through geophysical analysis. A conceptual model was then derived that states that these wetlands are held in place by clay-plugs that form through clay illuviation from the hillslopes at regions of valley confinement. This has important implications for the connectivity of wetland process domains. The research also determined the inputs of surface and subsurface flows to the wetland and it was found through detailed examination of soil moisture responses and variably saturated soil physics modelling using the HYDRUS model, that the wetland is hydrologically connected to its contributing hillslope by threshold induced preferential flow pathways, via macropores, that only respond after specific antecedent soil moisture conditions are met. In addition, the thesis describes novel approaches to use information provided by soil scientists for the development of catchment hydrological models. It was shown that the use of this hydropedology information improved the low flow response function of the catchment model, ACRU. This development has important implications for up-scaling of catchment process domains, or hydrological response units by being able to generalize on hillslope hydrological responses based on configuration of their soil type elements. The research also undertook to examine the role that the wetlands play in catchment processes. It was found through water budgeting, supported by hydrological time-series, stable isotope analysis and the quantification of vegetation water use within the wetland and contributing catchment, that these wetlands do not augment baseflows during the dry season. Furthermore, it is only early on during the wet season that these systems may attenuate peak flows, thereafter they act as conduits for high storm flows. Similarities emanated from this research with previous hydrological studies of headwater wetland systems in southern Africa and these are discussed. / Thesis (Ph.D.)-University of KwaZulu-Natal, Pietermaritzburg, 2011.

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