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Tempo de digest?o e caracteriza??o do trato digest?rio de larvas de pacam? (Lophiosilurus alexandri) / Food and initial development of pacam? larvae (Lophiosilurus alexandri)Pereira, Daiane Kelly Alves 29 August 2016 (has links)
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Previous issue date: 2016 / Funda??o de Amparo ? Pesquisa do Estado de Minas Gerais (FAPEMIG) / Coordena??o de Aperfei?oamento de Pessoal de N?vel Superior (CAPES) / Conselho Nacional de Desenvolvimento Cient?fico e Tecnol?gico (CNPq) / Banco do Nordeste do Brasil S.A. (BNB) / A aquicultura ? uma pr?tica brasileira que cresceu expressivamente nas ?ltimas d?cadas, contudo, a produ??o de pescados ainda ? insuficiente para atender a demanda interna. Na ?sia, maior produtor mundial de pescados, 95% das esp?cies produzidas s?o nativas, enquanto no Brasil, esse percentual est? abaixo de 20%. Este estudo teve como objetivo verificar o tempo de digest?o e avaliar o trato digest?rio de larvas de Lophiosilurus alexandri, uma esp?cie nativa do Rio S?o Francisco. As larvas foram avaliadas com 12 e 19 dias de vida, alimentadas com n?uplios de art?mia salina. Para avalia??o do tempo de digest?o nos diferentes dias de amostragem foi utilizada uma regress?o linear. Ao t?rmino do experimento foram aferidas as medidas de peso (g), e comprimento total (CT), comprimento de boca (CBO) e largura de boca (LBO) (mm). Para caracteriza??o do trato, al?m das j? citadas, foram tomadas medidas de comprimento total, comprimento boca/es?fago, comprimento do est?mago, comprimento do intestino, comprimento da boca, largura de boca e quociente intestinal para 12 e 19 dias de vida. As larvas apresentaram nos diferentes dias de amostragem peso, CT, CBO, LBO e QI de 41,18 mg e 76,88 mg, 17,78 mm e 20,98 mm, 0,42 mm e 0.73 mm, 2,71 mm e 3,57 mm, 0,29 mm e 0,32 mm aos 12 e 19 dias respectivamente. Foi verificado para os par?metros observados um maior desenvolvimento das larvas com 19 dias em rela??o as de 12 dias. Os par?metros de qualidade de ?gua mantiveram-se est?veis durante todo o per?odo experimental, permanecendo dentro dos valores aceit?veis para a larvicultura da esp?cie, assim como o crescimento em peso e comprimento. A avalia??o do desenvolvimento do sistema digest?rio das larvas foi realizada atrav?s de an?lises histol?gicas. O tempo de digest?o em larvas com 12 dias de vida foi menor (2 h 39 min 18 s) do que as de 19 dias (3 h 5 min 50 s). Por outro lado, larvas com 19 dias de vida apresentam trato digest?rio mais diferenciado em rela??o a 12 dias, permitindo assimilar melhor o alimento, aumentando assim a probabilidade de sobreviv?ncia dos indiv?duos. / Disserta??o (Mestrado) ? Programa de P?s-Gradua??o em Zootecnia, Universidade Federal dos Vales do Jequitinhonha e Mucuri, 2016. / Aquaculture is a Brazilian practice that has grown significantly in the last decades, however, fish production is still insufficient to meet domestic demand. In Asia, the world's largest fish producer, 95% of the species produced are native, while in Brazil, this percentage is below 20%. This study aimed to verify the digestion time and to evaluate the digestive tract of larvae of Lophiosilurus alexandri, a species native to the S?o Francisco River. The larvae were evaluated at 12 and 19 days of age, fed with Artemia salina nauplii. To evaluate the digestion time on the different sampling days a linear regression was used. At the end of the experiment, the measurements of weight (g), total length (CT), length of mouth (BOD) and mouth width (LBO) (mm) were measured. For characterization of the tract, in addition to those already mentioned, measurements were taken of total length, mouth length / esophagus, length of the stomach, length of the intestine, length of mouth, mouth width and intestinal quotient for 12 and 19 days of life. The larvae presented, on the different days of sampling, weight, CT, CBO, LBO and IQ of 41.18 mg and 76.88 mg, 17.78 mm and 20.98 mm, 0.42 mm and 0.73 mm, 2.71 mm And 3.57 mm, 0.29 mm and 0.32 mm at 12 and 19 days respectively. It was verified for the observed parameters a greater development of the larvae with 19 days in relation to the 12 days. The water quality parameters remained stable throughout the experimental period, remaining within acceptable values for the species larviculture, as well as growth in weight and length. The evaluation of the development of the digestive system of the larvae was performed through histological analysis. The digestion time in larvae with 12 days of life was lower (2 h 39 min 18 s) than those of 19 days (3 h 5 min 50 s). On the other hand, larvae with 19 days of life present digestive tract more differentiated in relation to 12 days, allowing assimilating the food better, thus increasing the probability of survival of the individuals.
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Adapta??es morfol?gicas do trato digest?rio do peixe neotropical Steindachnerina notonota (Characiformes, Curimatidae) ao h?bito alimentar detrit?voro / Morphological adaptations of the digestory tract of neotropical fish Steindachnerina notonota (Characiformes, Curimatidae) to the detritivore feeding habitSilva, Lenilda Teixeira da 21 June 2016 (has links)
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Previous issue date: 2016-06-21 / A Detritivoria ? comum em peixes dulc?colas, sobretudo na regi?o neotropical, e para muitas esp?cies s?o escassas as informa??es sobre o trato digest?rio, como ? o caso da esp?cie Steindachnerina notonota, na qual, tem import?ncia ecol?gica como elo da cadeia alimentar, servindo de alimento para os peixes carn?voros, e atuando na depura??o dos ecossistemas aqu?ticos sujeitos ? polui??o org?nica. O objetivo do presente estudo consiste em analisar as adapta??es morfol?gicas do trato digest?rio, desta esp?cie end?mica da caatinga, regi?o do semi?rido brasileiro. Assim, foram utilizados 10 esp?cimes adultos (machos e f?meas), necropsiados, para an?lises macrosc?picas e microsc?picas do trato digest?rio. Todas as estruturas foram observadas no estereomicrosc?pio. Os ?rg?os foram fixados em formol 10% e submetidos ? colora??o de Hematoxilina e Eosina, e PAS - ?cido Peri?dico Schiff (rastros branquiais, ?rg?o epibranquial, es?fago, est?mago, intestino m?dio e intestino posterior) e analisados em microscopia de luz. Os rastros branquiais tamb?m foram fixados em paraformalde?do 10% para an?lise em microscopia eletr?nica de varredura. Os esp?cimes analisados possu?am comprimento total entre 6,4 a 13 cm, comprimento padr?o entre 4,8 a 10,3 cm, e peso de 3,23 a 35,92 g. As adapta??es estruturais no trato digest?rio foram: boca do tipo terminal; o particular complexo bucofar?ngeo; aus?ncia de dentes e da l?ngua; tem tr?s tipos de formatos de rastros branquiais com c?lulas mucosas e bot?es gustativos; possui o ?rg?o epibranquial com suas peculiaridades; um est?mago mec?nico com musculatura muito desenvolvida na regi?o pil?rica; e um intestino muito longo e enovelado, que tem internamente uma exclusiva prega helicoidal. Os aspectos morfol?gicos do trato digest?rio de S. notonota t?m rela??o ao h?bito alimentar especializado detrit?voro-ili?fago. / Detritivory is the common freshwater fishes, especially in the Neotropical region, and for many species are scarce information about the digestory tract, as is the case of the species Steindachnerina notonota, in which, it has ecological importance as links in the food chain, serving of feed for carnivorous fishes, and acting in the depuration of aquatic ecosystems subject to organic pollution. The objective of present study consists of analyzing the morphological adaptations of the digestory tract, of this endemic species of the caatinga, Brazilian semiarid region. Thus, 10 adult specimens were used (males and females), necropsied for macroscopic and microscopic analysis of the digestory tract. All structures were observed in the stereomicroscope. The organs were fixed in 10% formalin and submitted to hematoxylin and eosin staining, and PAS - periodic acid Schiff (gill rakers, epibranchial organ, esophagus, stomach, midgut and hindgut) and analyzed by light microscopy. The gill rakers were also fixed in 10% paraformaldehyde for analysis by scanning electron microscopy. The specimens analyzed had total length between 6,4 to 13 cm, standard length between 4,8 to 10,3 cm, and weight of 3,23 to 35,92 g. The structural adaptations in the digestory tract were: mouth of the terminal type; the particular complex bucopharyngeal; absent from tooth and tongue; it has three types of formats of gill rakers with mucous cells and taste buds; possess the epibranchial organ with its peculiarities; a mechanical stomach with highly developed musculature in the pylorus; and a very long and coiled intestine, which internally has exclusive helical fold. The morphological aspects of digestory tract of S. notonota are related to detritivore-ilyophago specialized feeding habit.
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Estrat?gias tr?ficas do peixe tra?ra, Hoplias malabaricus e do peixe cascudo, Hypostomus pusarum do a?ude Marechal Dutra, Rio Grande do Norte, BrasilPessoa, Emilly Kataline Rodrigues 04 July 2014 (has links)
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Previous issue date: 2014-07-04 / Conselho Nacional de Desenvolvimento Cient?fico e Tecnol?gico (CNPq) / O peixe tra?ra, Hoplias malabaricus (Erythrinidae) e o peixe cascudo, Hypostomus pusarum (Loricariidae) s?o peixes de ?gua doce, da regi?o neotropical de consider?vel import?ncia ecol?gica e econ?mica. O trabalho teve como objetivo investigar as estrat?gias tr?ficas e a morfo-histologia do trato digest?rio desses peixes no a?ude Marechal Dutra, Acari, do semi?rido do Rio Grande do Norte, Brasil. Foram analisados 133 exemplares de tra?ra, H. malabaricus e 118 esp?cimes de cascudo, H. pusarum. As duas esp?cies estudadas apresentam diferentes n?veis de cadeia alimentar e ocupa??o espacial na coluna de ?gua do a?ude. Os resultados do trabalho est?o apresentados sob a forma de quatro artigos cient?ficos. O primeiro artigo descreve a anatomia e a histologia do trato digestivo e os h?bitos alimentares da tra?ra, H. malabaricus. O intestino da tra?ra ? curtocom um coeficiente intestinal de 0,72 ? 0,09. O ?ndice de import?ncia alimentar demonstrou que H. malabaricus se alimenta preferencialmente de material animal, sendo 72,8% de peixes e 27,2% de camar?es. A histologia do seu tubo digest?rio confirma seu h?bito alimentar carn?voro. O segundo artigo aborda um estudo pioneiro sobre os h?bitos alimentares e a caracteriza??o anat?mica e histol?gica do trato digest?rio de H. pusarum. O intestino do cascudo ? longo com um coeficiente intestinal de 10.8?0.7. O ?ndice de import?ncia alimentar demonstrou que H. pusarum se alimenta preferencialmente de material org?nico em decomposi??o (88.7%) e microalgas filamentosas e diatom?ceas (11.3%). O terceiro artigo compara os aspectos morfol?gicos do trato digest?rio de H. malabaricus e H. pusarum, relacionando-os com seu h?bito alimentar. A disposi??o dos ?rg?os digest?rios em ambas as esp?cies est? diretamente relacionada com a forma da cavidade peritoneal e o formato do corpo. O intestino curto em H. malabaricus e muito longo em H. pusarum est? relacionado com seus h?bitos alimentares. A morfologia dos tubos digest?rios de H. malabaricus e H. pusarum confirmam seus h?bitos alimentares, carn?voro e detrit?voro/herb?voro, respectivamente. O quarto artigo aborda os aspectos alimentares e reprodutivos (estrutura de comprimento e peso, rela??o peso-comprimento, tipo de crescimento, propor??o sexual) do cascudo, H. pusarum. A esp?cie apresenta crescimento alom?trico negativo, com a predomin?ncia de f?meas na popula??o amostrada.Hoplias malabaricus, que habita o ambiente pel?gico, ? considerado carn?voro, enquanto que H. pusarum, que habita o ambiente bent?nico, ? caracterizado como detrit?voro/herb?voro. Cada esp?cie estudada mostra um regime alimentar bem diferente, sem que haja competi??o tr?fica entre elas. As estruturas morfo-anat?micas do trato digest?rio refletem sua estrat?gia alimentar. / The fish, Hoplias malabaricus (Osteichthyes: Erythrinidae) and armored catfish,
Hypostomus pusarum (Loricariidae) are of freshwater origin from the neotropical region, and
are considered ecologically and economically of important. This work investigated the trophic
strategies, the morphology and histology of the digestive tract of these fish captured from the
Marechal Dutra reservoir, Acari, the semiarid region of Rio Grande do Norte, Brazil. A total
of 133 individuals of H. malabaricus and 118 specimens of H. pusarum were analyzed. The
two study species occupy different levels in the food chain and spatial distribution in the
water column of the reservoir. The results of this study are presented in the form of four
scientific papers. The first article describes the morphology and histology of the digestive
tract and the feeding habits of H. malabaricus (Bloch, 1794). This fish has a short intestine,
with an intestinal coefficient of 0.72 ? 0.09. The dietary importance index indicates that H.
malabaricus feeds preferentially on animal matter, especially on fish (72.8%) and prawns
(27.2%). The histology of its digestive tract confirms the carnivorous feeding habit. The
second article discusses about the pioneering work on the feeding strategy and the
characterization of anatomy and histology of the digestive tract of H. pusarum. The intestine
of this fish is long, with an intestinal coefficient of 10.8?0.7. The dietary importance index
indicates that H. pusarum feeds preferentially on organic matter in decomposition (88.7%)
and on filamentous microalgae and diatoms (11.3%). The third article compares the
morphological aspects of the digestive tract of H. malabaricus and H. pusarum, in relation to
their food habits. The arrangement of the digestive organs in both species is directly related to
the shape of the peritoneal cavity and the form of the body. The short intestine of H.
malabaricus and the long intestine of H. pusarum are associated with their feeding habits. The
morphology of the digestive tracts of H. malabaricus and H. pusarum confirm their food
habits, carnivorous and detritivorous / herbivorous, respectively. The fourth article discusses
the food and reproductive aspects (length and weight, length-weight relationship, type of
growth and sex ratio) of H. pusarum. This species has a negatively allometric growth, with
the predominance of females in the sampled population. H. malabaricus inhabits the pelagic
environment and is a carnivore, while H. pusarum, lives in the benthic environment and is
characterized as a detritivore/herbivore. Each species studied shows a very different diet,
without trophic competition between them. The morphological and anatomical structures of
the digestive tract reflect their feeding strategy.
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Din?mica populacional e alimentar de Stegastes fuscus, (Osteichthyes:pomacentridae) em arrecifes da praia de B?zius, no Rio Grande do NorteCanan, Bhaskara 14 December 2007 (has links)
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Previous issue date: 2007-12-14 / The damselfish, Stegastes fuscus Cuvier, 1830 (Osteichthyes: Perciformes: Pomacentridae), is abundant in the coastal reefs of B?zios Beach, Rio Grande do Norte, Brazil and they play an important role in the reef community ecology. The
present study investigated the feeding strategy of this species considering the food habits and morfo-histology of the digestive tract. Influence of the environmental
correlates such as temperature, rainfall and luminosity in the tidal rock pools were studied. The fish were captured on a monthly basis from September 2004 to August 2005, during which period 842 individuals of S. fuscus were captured, 125 males, 437 females and 280 individuals without sex identification. The sex ratio observed was 1 M : 3,5 F. The total body length of males varied from 3.6 to 11.3 cm, with a mean of 7.77 cm; that of females varied from 2.9 to 11.4 cm, with a mean of 7.85 cm, and that of sex grouped individuals varied from 2.9 to 11.4 cm, with a mean of 7.83. However,
there was no difference between males and females in total body length. This species presented a positive alometric growth and the equations obtained for the relation
between body mass and total body length were: Wt =0,0174Lt
3,1123 for males; Wt =0,0137Lt 3,2294 for females and Wt = 0,0148Lt 3.1928 for sex grouped individuals. The
relation between total body length and standard length was L t = 1 ,3223Ls + 0,1527 for sex grouped individual s. February to August was associated to a long period of gonadal resting. The fish spawned during January and in September -October. The frequency of fish with empty stomachs occurred during August to December, whereas frequency of fish with food contents i n stomachs occurred during January to July. This
species is considered as a preferential herbivore based on the volume of macroalgae in its diet composition. The morfo -histological aspects of S. fuscus confirm herbivory.
Among the environmental factors con sidered only rainfall showed a correlation with the feeding habits of this species / O peixe-donzela, Stegastes fuscus Cuvier, 1830 (Osteichthyes: Perciformes: Pomacentridae), ? abundante nos arrecifes costeiros da praia de B?zios, Rio Grande do Norte e desempenha um important e papel ecol?gico em comunidades recifais. O presente estudo investigou a din?mica populacional e alimentar desta esp?cie levando em considera??o o regime alimentar e a morfo -histologia do trato digest?rio. Avaliou -
se a influ?ncia das vari?veis ambientais, tais como, temperatura, pluviosidade e luminosidade em po?as de mar?. Foram realizadas coletas mensais no per?odo de setembro de 2004 a agosto de 2005, durante as quais foram capturados 842 indiv?duos de S. fuscus, sendo 125 machos, 437 f?meas al?m de 280 indiv?duos com sexo n?o identificado. A propor??o sexual observada foi de 1 M : 3,5 F. Em rela??o ao
comprimento total os valores para machos machos variaram de 3,6 a 11,3 cm, com m?dia de 7,77 cm; para as f?meas de 2,9 a 11,4 cm, com m?dia de 7,85 cm e p ara os
sexos agrupados de 2,9 a 11,4 cm, com m?dia de 7,83 cm, n?o havendo diferen?as entre os sexos quanto ao comprimento total. A esp?cie estudada apresentou um
crescimento do tipo alom?trico positivo e as equa??es obtidas para a rela??o entre peso total e comprimento total foram: Wt = 0,0174Lt 3,1123 para machos; Wt =0,0137Lt 3,2294 para f?meas e Wt = 0,0148Lt 3,1928 para os sexos agrupados. A rela??o entre o comprimento total e o comprimento padr?o foi L t = 1,3223Ls + 0,1527 para os
sexos agrupados. Nesses peixes foi observado um per?odo de repouso gonadal que se prolongou de fevereiro a agosto com a desova sendo realizada em janeiro e em setembro e outubro. Em rela??o ao regime alimentar, as maiores freq??ncias de est?magos sem alimento ocorreram entre os meses de agosto a dezembro enquanto que as maiores freq??ncias de est?magos com alimento ocorreram nos meses de janeiro a julho. Em fun??o do volume de macroalgas em sua composi??o de dieta, a esp?cie Stegastes fuscus ? considerada como preferencialmente herb?vora e os
aspectos morfo-histol?gicos confirmam essa prefer?ncia. Dentre os fatores abi?ticos considerados por ocasi?o do estudo utilizando Stegastes fuscus, apenas a pluviosidade
influenciou significativamente no regime alimentar desta esp?cie
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Estrat?gia de vida do peixe anual Hypsolebias antenori (Cyprinodontiformes: Rivulidae) do semi?rido brasileiroNascimento, Wallace Silva do 04 June 2014 (has links)
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Previous issue date: 2014-06-04 / Conselho Nacional de Desenvolvimento Cient?fico e Tecnol?gico / This work deals with the life strategy of an endangered annual fish, Hypsolebias
antenori (Cyprinodontiformes: Rivulidae), of the Brazilian semiarid region. The eggs of these
fish hatch at the onset of the rainy season, grows rapidly and they reproduce during the rainy
season. When the water puddles dry out, the entire population dies. The resistant eggs which
are buried in the bottom of the dried pools go through diapause stages, during which time the
embryonic development becomes temporarily arrested. With the onset of the next rainy
season, the eggs hatch and a new generation is formed. Specimens of H. antenori were
captured during 2011 and 2013, in temporary water pools located in the hydrographic basin of
river Jaguaribe in Cear?, Brazil. Sex ratio, the length-weight relationship, the growth type,
first sexual maturity, anatomy and histology of the digestive tract, development of gonads,
reproductive strategy, karyotypic pattern of the species, and the conservation status of H.
antenori were investigated. The results of this study are presented in the form of eight articles.
The first article is about the fish faunal composition of the hydrographic basins of Rio Grande
do Norte, Brazil, wherein the record of H. antenori is included. The second article deals with
the sex ratio, secondary sexual characteristics of males, the length-weight relationship and the
type of growth. Males show a pattern of intense coloration with well developed fins. The sex
ratio showed a significant predominance of females (1M:1.7 F). Males were larger in length
and weight. The equation of weight and total length relationship was Wt=0.0271Lt3,8937,
showing a positively allometric growth, indicating greater increase in weight than in length.
The third article discusses the anatomy and histology of the digestive tract of H. antenori. It is
considered as a generalist feeder with characteristics of omnivore, which utilizes different
food sources. The fourth article discusses the stages and phases of gonad development and
type of spawning of H. antenori. The fifth article is about the r reproductive strategy adopted
by H. antenori which helps in successful reproduction over a short period of life. The sixth
article deals karyotypic pattern of the species, constituting the first cytogenetic contribution to
the genus. The seventh article discusses about the risk of extinction of this species which
suffers a series of threats, such as, habitat loss through land use, deforestation, construction of
reservoirs, pollution due to domestic and industrial sewage, besides pesticides and
agrochemicals. Furthermore, decreasing rainfall and intensification of aridity due to global
climate changes, interferes with the reproductive cycle. The eighth article deals with
aggressive behavior adopted between males and among females during reproduction. All
temporary water pools sampled during this study were in high degree of degradation, mainly
due to human action. There is a great need for conservation measures to protect the
populations of annual fish, including the creation of protected areas in the semiarid ephemeral
aquatic environments of Brazil / Este trabalho trata sobre a estrat?gia da vida de um peixe anual em perigo Hypsolebias
antenori (Cyprinodontiformes: Rivulidae), do semi?rido brasileiro. Os ovos dessa esp?cie
nascem em po?as tempor?rias, crescem rapidamente, reproduzem e no per?odo de estiagem
quando a po?a seca, toda a popula??o morre. Os ovos resistentes depositados no substrato da
po?a s?o capazes de passar por um est?gio de diapausa, quando seu est?gio de
desenvolvimento embrion?rio ? estacionado. Com a chegada do per?odo de chuvas, os ovos
eclodem e uma nova gera??o ? formada. Os exemplares de H. antenori foram capturados
durante 2011 e 2013, nas po?as tempor?rias localizadas na bacia do rio Jaguaribe no Cear?,
Brasil. Foram investigadas a propor??o sexual, a rela??o peso-comprimento, o tipo de
crescimento, a primeira matura??o sexual, anatomia e histologia do trato digest?rio,
desenvolvimento das g?nadas, estrat?gia reprodutiva, dados cariot?picos da esp?cie, e o estado
de conserva??o de H. antenori. Os resultados desse trabalho est?o apresentados sob a forma
de oito artigos. O primeiro artigo ? sobre a composi??o da ictiofauna das bacias hidrogr?ficas
do Rio Grande do Norte, Brasil, incluindo o registro de H. antenori. O segundo artigo trata
sobre a propor??o sexual, caracter?sticas secund?rios sexuais dos machos, a rela??o pesocomprimento
e o tipo de crescimento. Os machos apresentam um padr?o de colora??o intensa
com nadadeiras mais desenvolvidas. A propor??o sexual apresentou uma predomin?ncia
significativa de f?meas (1M: 1,7F). Os machos foram maiores em comprimento e em peso. A
equa??o originada da rela??o peso total e comprimento total foi Wt = 0,0271Lt3,8937,
apresentando um crescimento alom?trico positivo, indicando maior incremento no peso do
que em rela??o ao comprimento. O terceiro artigo trata sobre a anatomia e histologia do trato
digest?rio de H. antenori. ? considerado como um generalista com as caracter?sticas de
on?voria, que aproveita fontes alimentares diferentes. O quarto artigo trata sobre os est?dios e
fases de desenvolvimento das g?nadas e o tipo de desova de H. antenori. O quinto artigo trata
sobre a estrat?gia reprodutiva r adotada pela H. antenori que auxilia na reprodu??o bem
sucedida durante um curto per?odo de vida. O sexto artigo trata sobre os dados cariot?picos da
esp?cie, constituindo a primeira contribui??o citogen?tica para o g?nero. O s?timo artigo trata
sobre o risco de extin??o dessa esp?cie, que v?m sofrendo uma s?rie de amea?as, tais como, a
perda de habitats pela ocupa??o do solo, desmatamento, constru??o dos reservat?rios,
polui??o devido aos efluentes dom?sticos, industriais, pesticidas e agrot?xicos. Al?m disso, a
diminui??o da pluviosidade e intensifica??o da aridez em decorr?ncia das mudan?as
clim?ticas globais vem interferindo em seu ciclo reprodutivo. O oitavo trabalho trata sobre o
comportamento agressivo adotado entre os machos e entre as f?meas durante a reprodu??o.
Todas as po?as tempor?rias amostradas durante este trabalho estavam em um alto grau de
degrada??o, principalmente devido ? a??o antr?pica. H? uma grande necessidade de medidas
de conserva??o para proteger as popula??es de peixes anuais, entre elas especialmente a
cria??o de ?reas de prote??o nos ambientes aqu?ticos ef?meros do semi?rido brasileiro
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