Thesis (M.S.)--Naval Postgraduate School, 1984. / Includes bibliographical references (p. 143-145).

Thesis (Ph. D.)--University of Wisconsin--Madison, 1976. / Typescript. Vita. eContent provider-neutral record in process. Description based on print version record. Includes bibliographical references (leaves [259]-264).

Wind-driven up welling events near Brooks Peninsula off the west coast of Vancouver Island have been identified during the summers of 1988 and 1989 from sea surface NOAA AVHRR thermal imagery obtained at the UBC Satellite Oceanography and Meteorology Laboratory. Software has been developed to characterize the strength and extent of the surface cooling associated with the upwelling. A two-dimensional, 2-layer finite difference model with 1 km resolution has been formulated to examine the small-scale dynamics of the upwelling events. The model uses local wind and includes realistic coastline and bathymetry. The wind-stress and coastline configuration appear to be the main factors contributing to favourable upwelling regions. The results of the model compare favourably to the observed starting location of the upwelling. / Science, Faculty of / Earth, Ocean and Atmospheric Sciences, Department of / Graduate

The role of heterotrophic bacteria in the carbon and nitrogen flux of the pelagic food web was investigated during laboratory and field-based studies of the temporal development of the planktonic community after upwelling. Bacterial community structure, activity and production were closely coupled to the upwelling cycle and to the dynamics of the phytoplankton community. The initial bacterial population (<l x 10⁶ cells ml⁻¹, 20 to 40 μg C l⁻¹) was metabolically dormant. Increased availability of phytosynthetically produced dissolved organic carbon (PDOC) stimulated bacterial growth (0.016 h⁻¹) and abundance (8 to 10 x 10⁶ cells ml⁻¹, 140 to 200 μg C l⁻¹). Rapid successions in the dominant plateable strains were attributed to substrate preferences and substrate availability. Significant correlations of bacterial biomass with total standing stocks of phytoplankton and particulate carbon provided evidence of close coupling between bacteria and PDOC, and between bacteria and recalcitrant substrates available during phytoplankton decay. These relationships were best described by power functions, suggesting that bacterial biomass was relatively reduced at high levels by predation. A microcosm study indicated that zooflagellate predation could control bacterial biomass. Low net growth yields (34 to 36%) of flagellates suggested inefficient transfer of carbon to higher trophic levels, but considerable nitrogen regeneration (ca 6 to 7 μg N mg dry weight⁻¹ h⁻¹). Thymidine-measured bacterial production (TTI, <0.1 to 1.25 mg C m⁻³ h⁻¹) was linearly related to phytoplankton growth. Non-uniform response of bacteria to added tracer substrates may result in underestimates of bacterial production by 2 to 34 times by TTI, particularly in deep or oligotrophic waters, or during phytoplankton decay. Close coupling of copepod (Calanoides carinatus) production to the upwelling cycle suggested co-existence of the microbial food web and the classical diatom-copepod food chain. Recently upwelled water was dominated by phytoplankton. Assuming that all phytoplankton carbon was available for utilisation, copepods and bacteria were calculated to consume approximately 12 and 22% of primary production respectively. As the bloom declined the planktonic community was increasingly dominated by bacteria, detritus and mesozooplankton. On average, copepods consumed 60% of primary production, while bacteria consumed 49%. Carbon consumption requirements of both bacteria and copepods were satisfied by resource partitioning and carbon cycling. Under food-limiting conditions herbivorous copepods may switch to omnivory, ingesting microzooplankton of the microbial food web, and stimulating enhanced remineralisation to further sustain primary production. A generic size-based simulation model of the dynamics of the plankton community indicated that bacteria and the microbial food web increase the overall productivity of the planktonic food web, and that heterotroph predation in the smaller size classes (<200 μm) is an important mechanism in nutrient recycling.

Thesis (M.S.)--Oregon State University, 2006. / Printout. Includes bibliographical references. Also available on the World Wide Web.

The aim of this thesis was to assess the influence of upwelling on alga-grazer interactions in rocky shore communities along the south coast of South Africa using grazer exclusion treatments with controls and procedural controls set out in a block design and monitored for algal cover roughly monthly for one year. In the first experiment grazers were excluded from treatment plots at two upwelling and two non-upwelling sites and the rates of algal biomass accumulation were then compared. The upwelling sites showed significantly faster algal colonisation rates, with Ulva rigida being the first species to colonise the rocks. Final algal cover and biomass did not differ significantly between upwelling and non-upwelling sites in control plots open to grazers, but were significantly higher in grazer exclusion plots at upwelling sites indicating stronger grazing effects. This was confirmed by estimating the intensity of grazing using the log-response ratio (LRR), which was calculated from treatment and control plots. Upwelling sites had significantly lower LLR values indicating stronger grazing effects, than at non-upwelling sites, despite no difference in grazer abundances. The second experiment examined the effects of nutrient addition on algal growth and community composition by comparing high nutrient enrichment plots with low enrichment plots at one upwelling and one non-upwelling site. ANOVA indicated faster growth rates and significantly higher final algal biomass in high enrichment plots compared to low enrichment and control plots at both upwelling and non-upwelling sites. A two-way ANOVA indicated significantly higher algal cover in high enrichment plots compared to the data from the grazer exclusion plots in experiment 1 at both sites, suggesting that nutrient addition plays a major role in algal growth and community composition. The findings of these studies have shown significant differences between treatments, sites and seasons, with significant differences not only occurring in algal cover but also accumulation of algal biomass and recruitment patterns between treatments. The small scale local processes acting within a few centimetres (plots) or tens of meters (among blocks) can also be reflected over larger scales such as sites (upwelling/non-upwelling shores). Further, these studies have demonstrated that various factors such as the effects from increased nutrients at upwelling cells and the change in grazing effects due to enhanced nutrients can determine the abundance and diversity of the community structure, including an increase in the abundance of the fast growing algae Ulva rigida, and a slow recovery of the brown and red algae.

Thesis (M.A.)--Stellenbosch University, 1988. / ENGLISH ABSTRACT: The Benguela upwelling system off southern and south western Africa is a zone of strong and extensive upwelling. Owing to the greater fIsh numbers found along the front between the upwelling and South East Atlantic Ocean the frontal zone is a key element in the ecology of the upwelling area. This discourse focuses on the perturbations of this front. The major data source used was the daily Meteosat satellite imagery for the years 1978, 1979 and 1982 to 1985 in the visible and infrared wavebands. These data enable the investigation of cross frontal activity for the entire Benguela Upwelling System at various spatial and temporal scales. NOAA satellite infrared imagery was also used. At the macroscale two upwelling zones are described. The fIrst is the upwelling core along the coast which exhibits well-developed and persistent upwelling. Offshore of this is an area in where the more transient fllament activity predominates. This outer zone is one of constant change and presents highly variable frontal boundary locations. As far as the persistent upwelling is concerned, the northern boundary closely approximates that suggested by Shannon (1985) of l7oS. Cape Agulhas was the effective southern boundary of any regular upwelling. The mean offshore extent of the outer upwelling is 270 kilometres off Liideritz and 45 kilometres off Cape Town. An analysis of the seasonal location of the front shows that the greatest upwelling extent at both Liideritz and Cape Town is observed in winter. Although the winter upwelling extent is the greater of the two seasons, the summer frontal location at Cape Town, in turn, exhibits remarkable stability. The upwelling off Liideritz, on the other hand, is prone to almost constant frontal location fluctuations. . Most of the variation occurs as a result of the growth and decay of fllaments. Filaments were seen along the entire upwelling zone from Cape Agulhas to Cape Frio. On average, the fllament sector was 270 kilometres wide off Liideritz. Between Cape Point and Cape' Agulhas the southwards extent of the upwelling rarely exceeded 40 kilometres, while the maximum fllament off Cape Point was about 200 kilometres. A fllament at Cape Point generally extends about 40 kilometres westwards and the upwelling zone off Walker Bay stretches about 20 kilometres southwards, onto the Agulhas Bank. Along the coast between Cape Agulhas and Cape Point the fllaments rapidly react to changes in the wind speed and direction. Mesoscale weather systems are therefore important factors in determining fllament activity there. This also true to some extent at Liideritz. The berg wind can induce rapid fllament growth off Liideritz. Despite an upwelling positive wind direction it is the wind speed that determines whether upwelling will develop at Cape Town at all. At Liideritz the wind speeds appear to determine the cross frontal temperature gradient. The greater the wind speed, the steeper the temperature gradient. Cross frontal temperature gradient of between 0.020C and .006oC per kilometre were calculated for Liideritz, which compares well with readings in the Californian upwelling zone (Koblinsky et al. 1984). In the way fllaments extend the length of the frontal zone greatly and the manner in which these cold water streams react to the changing winds, they are complex upwelling frontal features of great variability and importance in the Benguela upwelling system. / AFRIKAANSE OPSOMMING: geen opsomming

Understanding how large-scale processes (>100 kms) influence ecological communities is currently a major focus in ecology. In marine systems, coastal upwelling, a large-scale oceanographic process in which surface water pushed offshore by winds is replaced by cold, nutrient-rich water from depth, appears to cause variation in rocky intertidal communities. Along the central Oregon coast upwelling occurs intermittently during the summer while on the southern coast it begins earlier in the spring and is less variable throughout the summer. Coastal upwelling can affect rocky intertidal communities by altering the delivery of nutrients, larvae, and phytoplankton. I conducted three studies on both the southern and central Oregon coast to understand how differences in upwelling affect rocky intertidal community structure and dynamics. In the first study, I examined the recruitment and growth rates of sessile invertebrates (mussels and barnacles). Recruitment of both mussels and barnacles, and growth of mussels were consistently higher on the central Oregon coast than the southern coast. Upwelled water is nutrient-rich, so differences in upwelling are likely to affect growth rates of macroalgae. In the second study, I tested this hypothesis by monitoring the growth of two species of intertidal kelp at both central and southern coast sites. During El Ni��o years, when upwelling is sharply reduced on the central Oregon coast, algae may fare better at sites on the southern coast where upwelling is less affected. However, during years when upwelling is strong all along the coast, nutrient limitation does not appear to differentially affect macroalgal growth rates. Finally, in the third study, I examined the influence of upwelling on the interactions between microalgal primary producers and herbivorous limpets. I conclude that this interaction is complex and varies both within and between upwelling regions. My research suggests that a transition in upwelling from weak and sporadic on the central Oregon coast to stronger and more persistent on the southern Oregon coast drives the striking differences in rocky intertidal community structure and dynamics between these areas. / Graduation date: 2003

Thesis (Ph. D.)--Oregon State University, 2003. / Typescript (photocopy). Includes bibliographical references (leaves 143-152). Also available online.

Thesis (Ph. D.)--Florida State University, 2005. / Advisor: Dr. James J. O'Brien, Florida State University, College of Arts and Sciences, Dept. of Oceanography. Title and description from dissertation home page (viewed June 8, 2005). Document formatted into pages; contains xiii, 87 pages. Includes bibliographical references.