Following wildfires, plant materials are direct-seeded to limit erosion and annual weed invasion. Seedlings often fail to establish because selected plant materials are not always well adapted to local soil moisture and temperature conditions. In an effort to help improve plant materials selection and to evaluate sites potential revegetation, we have worked toward developing methodology to predict germination and root growth based on site specific soil moisture and temperature conditions. First, we characterized the seedbed environment of 24 sagebrush (Artemisia spp.) steppe sites throughout the Intermountain West to determine the wet-thermal time of five temperature ranges relevant to germination response and thermal-time model accuracy (Chapter 1). Second, we predicted potential germination for 31 plant materials at those same sites (Chapter 2). Third, in preparation to predict root growth at multiple sites, we characterized the drying patterns and the associated plant-available water for in the seedling root zone across nine woodland (Juniperus spp. and Piñus spp.) sites (Chapter 3). For all of these studies, we determined the effects of tree reduction and tree infilling phase at time of tree reduction. Our key findings are that seedbeds generally sum most wet-thermal time at temperature ranges where the germination rates fit thermal accumulation models quite well (R2 ≥ 0.7). The majority of plant materials summed enough wet-thermal time for a potential germination at most sites during the fall, early spring, and late spring. Soil drying primarily occurs from the soil surface downward. Drying rates and Plant available water associated with the first drying event increased with increasing soil depth. Root zone (1-30 cm) plant-available water increased before and decreased after the first spring drying event with increasing soil depth. Tree removal with increasing pretreatment tree infilling phase generally added progress toward germination, plant available water, and wet-thermal time in the seedbed and root zones of the sagebrush steppe in the Great Basin. Because soil moisture and temperature does not appear to be limiting for potential germination, combining germination and root growth models to create a more comprehensive model may allow for a more robust prediction for seedling survival. For either root growth or combined germination and root growth models, plant available water and wet-thermal time before the first spring drying period hold the most potential for successfully predicting seedling survival.

Juniper (Juniperus spp.) encroachment into sagebrush (Artemisia spp.) and bunchgrass communities has reduced understory plant cover and allowed juniper trees to dominate millions of hectares of semiarid rangelands. Trees are mechanically masticated or shredded to decrease wildfire potential and increase desirable understory plant cover. When trees are masticated after a major increase in tree population density and associated decrease in perennial understory cover, there is a risk that invasive annual grasses will dominate because they are highly responsive to the increased resource availability that commonly follows removal of the main resource user. To determine if tree mastication increases resource availability and subsequently favors invasive annual or perennial grasses, we compared soil temperature, water, and nutrient microenvironmental conditions and seedling establishment and growth. We used the major rangeland weed, cheatgrass (Bromus tectorum L.), to represent invasive annual grasses and Anatone bluebunch wheatgrass (Pseudoroegneria spicata (Pursh) A. Löve), a natural accession of native bluebunch wheatgrass, to represent the perennial grasses of the sagebrush-bunchgrass plant community. These comparisons were made between and within paired-adjacent masticated and untreated areas at three locations in Utah dominated by Utah juniper (Juniperus osteosperma (Torr.) Little). Juniper tree mastication generally increased resource availability with masticated areas having greater soil temperature, soil water availability, and soil N supply rates than untreated areas. Prior to juniper tree mastication litter mounds were not found to be resource islands probably because juniper trees themselves were using subcanopy soil water and nutrients. After juniper tree mastication and elimination of these predominant resource users, litter mounds served as resource islands with greater soil water availability and N supply rates than bare interspaces during the critical time for seedling establishment in spring. Plant growth followed in line with greater resource availability after tree mastication with masticated areas having more productive although fewer invasive-annual and perennial grass seedlings than untreated areas. These results suggest that increases in resource availability and warmer spring temperatures associated with mastication will not necessarily favor invasive annual over perennial grass seedling establishment. Resilience of the sagebrush-bunchgrass community to return to dominance after juniper control will likely be greatly influenced by how much of the sagebrush-bunchgrass community remains following tree control and the intensity of propagule pressure by invasive species. If only invasive annuals remain when the trees are treated then invasive annuals would be expected to dominate the post-treatment plant community especially with their ability to establish inside litter mounds unless they were also controlled and perennial grasses planted at the time of treatment.