Abies is the second largest genus of family Pinaceae (after Pinus), consisting of about 51 species, all native to the Northern Hemisphere. There are six native taxa belonging to this genus growing in pure and mixed stands in Turkey. Abies cilicica subsp. isaurica, Abies nordmanniana subsp. bornm&uuml / lleriana, Abies nordmanniana subsp. equi-trojani, Abies x olcayana are endemic and considered as lower risk (LR) species. To determine the genetic relationships in Turkish firs, 18 populations of different subspecies of Abies were collected from different regions of Turkey and non-coding trn regions of chloroplast DNA were sequenced to assess the genetic structure of the studied species. trnL, trnF and trnV region were examined. All the trn regions of Abies species in the world (aproximately 300 species that found in the IPNI (The International Plant Names Index) were investigated in the database of NCBI. The available trn sequences of 23 Abies species worldwide included into the analyses. All analyses to estimate molecular diversity parameters were carried out with the MEGA software. The constructed phylogenetic tree with the trn sequences revealed that Turkish firs formed a monophyletic group with almost no sequence divergence. v Since sequence data for all three sectors of trn were not available from the NCBI data base, the phylogentic analysis with the sequence data of trnL regions were compartively analyzed in all firs. The results showed that Turkish- European species formed a single clade, which clearly differentiated them from the others, such as Japanese species, A. veitchii. Similarly, according to the sequence data of trnF, Turkish fir species were grouped together and distinctly separated from Asian-American Fir species. The results suggest that all Turkish firs may have evolved from single ancestral fir species, most likely from Abies nordmanniana.

Pineacea is the largest family of conifers that includes 51 species of Abies which is the second largest genus after Pinus. There are six native taxa in Turkey belonging to this genus. Four of these taxa (Abies cilicica subsp. isaurica, Abies nordmanniana subsp. bornm&uuml / lleriana, Abies nordmanniana subsp. equi-trojani, Abies x olcayana) are endemic and considered as low risk (LR) species according to the IUCN criteria. To determine the phylogenetic relationship in Abies spp. in Turkey, 18 populations of different taxa were collected from their natural distribution areas in Turkey. The matK gene regions of chloroplast DNA (cpDNA) were studied comparatively to reveal the genetic relationship among Turkish fir species. The available sequences from the NCBI database for the matK region of the other Abies species in the world were also investigated v comparatively with the sequences from Turkish firs. With the matK sequence data, a phylogenetic tree was constructed for the fir species and the molecular diversity parameters such as conserved and variable sites, nucleotide diversity, and evolutionary divergence were estimated using the MEGA software. The results indicated that there are no variable sites among Turkish firs with regard to matK regions of cpDNA. It appears that the matK region of cpDNA for Turkish firs is highly conserved. Since sequence data for all matK regions were not available from the NCBI data base, the phylogentic analysis with the sequence data of matK were comparatively analyzed in all firs including Turkish firs. According to matK1 region, the results showed that there were three major clades. One of the clades included all Turkish fir taxa and one species from European firs, A.numidica / however, A.holopylla, A.firma, A.veitchii, A.sachalinensis, A.nephrolepis, A.lasiocarpa, A.koreana, A.homolephis, A.fraseri, A.fargesii, A.sibirica and A.fabri formed in another clade. In addition to this, A.mariesii, A.hidalgensis, A.bracteata, A.alba formed in different major clade. According to matK1 region the results showed that Turkish firs and European firs are closer to each other. Furthermore, based on matK2 region, the results indicated that Turkish firs formed a monophyletic group.The other fir species, with respect to matK2 regions, formed different clades from Turkish firs. The results based on matK region suggest that all Turkish firs may have evolved from single ancestral fir species and the matK gene region appears to be highly conserved region in fir species.

La présente thèse constitue une étude typologique contrastive des allomorphies pronominales dans deux langues finno-ougriennes : en hongrois et en mordve erzya. On entend ici par typologie contrastive une approche typologique fondée sur la mise en contraste des structures de deux ou plusieurs langues, y compris des langues de la même famille linguistique, afin d’explorer des propriétés à la fois spécifiques et universelles. De ce point de vue, le hongrois et le mordve s’avèrent particulièrement pertinents en termes de structuration des systèmes de marques pronominales, en raison de propriétés morphologiques caractéristiques de l’ouralien central et oriental, tels que l’existence d’une double conjugaison (subjective et objective, voire « objective définie », en mordve), qui induit des séries allomorphiques complexes, tout en suivant des principes réducteurs universels (syncrétisme, sous-spécification et surspécification de certaines marques ou conditions de marquage morphonologique). Cette thèse comprend neuf chapitres, distribués sur trois volets. Le premier volet décrit les structures et les étapes de la modélisation des systèmes pronominaux dans les deux langues. Dans le premier chapitre, nous présentons des généralités historiques et structurales du hongrois et du mordve erzya, ainsi que la place que ces langues occupent parmi les langues finno-ougriennes, du point de vue de la classification et de la typologie. Une série de particularités importantes pour la compréhension des deux systèmes, en termes d’organisation structurale, concerne les propriétés allomorphiques des unités fonctionnelles et relationnelles de type pronominal, telles que l’harmonie vocalique, les suffixes casuels, le système verbal, et l’ordre des mots. Le deuxième chapitre concerne le lien entre les pronoms personnels et des catégories grammaticales fondamentales telles qu’animacité, nombre, personne, définitude, et aboutit à la conclusion que c’est le pronom personnel qui est particulièrement marqué par ces catégories grammaticales – les mêmes qui peuvent avoir, dans les langues du monde, une incidence sur la construction ou l’organisation des systèmes de classes flexionnelles. Le troisième chapitre présente une approche historiographique du hongrois et du mordve erzya; le quatrième chapitre propose une réanalyse de la flexion pronominale erzya, en suivant les mêmes principes que ceux jadis préconisés par András Kornai dans son analyse du système de la flexion nominale du hongrois (Kornai 1994), dans la mesure où ce modèle morphologique traite l’affixation comme une opération sur des traits combinés. Le deuxième volet de cette recherche développe des études de cas exploratoires dans une perspective de TAL : un corpus d’erzya littéraire et un corpus d’erzya biblique sont analysés contrastivement en suivant les démarches et le paramétrage requis par le logiciel Trameur. Le troisième volet sort de l’analyse des registres stylistiques au sein d’une langue donnée pour revenir à une typologie contrastive structurale hongrois-mordve. Dans le dernier chapitre, nous proposons une synthèse de ces deux aspects de la typologie contrastive : contrastes de registres intralangue, contraste de structures interlangues, en fonction d’un ensemble de paramètres partagés. La synergie entre la méthode lexicométrique et la typologie générale constitue l’un des principaux apports heuristiques de cette thèse, dont le but est de développer une typologie des langues finno-ougriennes qui tienne davantage compte de la contrastivité des structures et de leur relativisme que des grands traits catégoriels interlangues, davantage sujets aux biais empiriques et méthodologiques que peuvent recéler les grands corpus. / This dissertation provides a contrastive and typological study of pronoun allomorphy in two Finno-Ugric languages: Hungarian and Erzya Mordvin. Contrastive typology is a typological approach aiming at contrasting the structures of two or more languages, including from the same language family, to explore specific and universal properties. From this standpoint, Hungarian and Mordvin are particularly relevant as to the structure of pronoun markers, due to some morphological characteristics of the central and eastern languages of the Uralic language family, such as double conjugation paradigms (subjective and objective, moreover the "objective definite inflectional paradigm" in Mordvin). This results in complex allomorphic patterns, while following universal principles (syncretism, sub-specification and over-specification of certain markers, or the conditions of morphonologic exponence). The first part describes the structures and modelling stages of the pronominal system in both languages. In the first chapter, we present historical and structural generalities about the Hungarian and Mordvin Erzya languages, and the place they occupy within the Finno-Ugric group, from the point of view of classification and typology. A series of important features to understand the two systems in terms of structural organization, concerns the allomorphic properties of functional and relational units of pronominal type such as vowel harmony, the casual suffixes, the verbal system, and word order. The second chapter deals with the relationship between personal pronouns and basic grammatical categories such as animacity, number, person, definiteness, and concludes that it is the personal pronoun that is most marked by these grammatical categories - the same that may affect, in the languages of the world, the construction or organisation of inflectional classes. The third chapter is a historiographical approach of Hungarian and Erzya to show the outline of the research on the evolutionary periods of both systems. The fourth chapter provides a reanalysis of pronominal inflection in Erzya, following the same principles as those previously recommended by András Kornai's analysis of the nominal inflection system of Hungarian (Kornai, 1994), as it deals with the morphological model considering affixation as an operation on the combined features. The second part of this research develops exploratory case studies from the perspective of NLP (French: TAL) a literary corpus and a biblical corpus of Erzya are analysed following the steps and the settings required by the Trameur software. The third part departs from the contrastive analysis of stylistic registers within a given language to return to a Hungarian-Mordvin contrastive structural typology. In the last chapter, we propose a synthesis of these two aspects of contrastive typology: contrasting registers of intralanguage, contrast-linguistic structures, based on a set of shared parameters. The synergy between the lexicometric method and the general typology is one of the main contributions of this thesis’s heuristics to develop a typology of Finno-Ugric languages that takes greater account of the contrastivity of structures and their relativism as major categorical traits of interlanguage, resulting more sensitive to empirical and methodological biases that may conceal a large corpus.