Thesis (Ph.D.)--York University, 2009. Graduate Programme in Higher Education. / Typescript. Includes bibliographical references (leaves 367-411). Also available on the Internet. MODE OF ACCESS via web browser by entering the following URL: http://gateway.proquest.com/openurl?url_ver=Z39.88-2004&res_dat=xri:pqdiss&rft_val_fmt=info:ofi/fmt:kev:mtx:dissertation&rft_dat=xri:pqdiss:NR51672

<p> The state and federally endangered California clapper rail (<i> Rallus longirostris obsoletus</i>) has declined in abundance and been reduced in range and now occupies fragmented intertidal saltmarsh only within San Francisco Bay. Historically extensive salt marsh habitats existed in San Francisco Bay and today, remnants are largely restricted to the water's edge with dikes and levees separating marshland from modified habitats unsuitable for clapper rails. Clapper rail population abundance has roughly tracked a series of positive and negative impacts including market hunting at the turn of the 20th century, widespread habitat reduction and fragmentation, and invasive species introduction and eradication programs. Despite these changes, rail populations have been subject to the ebb and flow of the tides, which regularly inundate salt marsh habitats. The influence that tides have on vertebrate species living in intertidal saltmarsh should be substantial, but the relationship between tide and California clapper rails is poorly understood. This research identified important ways in which tides influenced demographic processes, space use, and resource selection in California clapper rails. Tidal inundation in San Francisco Bay saltmarshes creates zonation in plant communities, typically with tall monocots in the low marsh (<i>Spartina </i> sp.), short pickleweed (<i>Sarcocornia pacifica</i>) in mid-elevation ranges, and gumplant (<i>Grindela humilis</i>) in the high marsh. Invasive Spartina (<i>Spartina foliosa x alterniflora </i>) grows taller and thicker than native Pacific cordgrass (<i> Spartina foliosa</i>). Invasive Spartina also grows lower onto mudflats, further up into pickleweed areas, and provided both nesting habitat and tidal refuge for clapper rails. In Chapter 1, I examined survival rates of California clapper rails. Specifically, I investigated whether seasonal patterns observed in the early 1990s were still evident and assessed the influence that Invasive Spartina and the degree of tidal inundation on weekly survival rates in four South San Francisco Bay salt marshes. Between January 2007 and March 2010, California clapper rail annual survival was 73% greater in Spartina-dominated marshes (&Scirc; = 0.482) than in a control marsh dominated by native vegetation (&Scirc; = 0.278). Lower survival also occurred during periods when tide height was greatest and during the winter. Survival patterns were consistent with Invasive Spartina providing increased refuge cover from predators during tidal extremes which flood native vegetation, particularly during the winter when the vegetation senesces. Tide heights also strongly influenced selection for artificial habitats provided adjacent to one marsh during the winters of 2010-2011 and 2011-2012. Ten floating islands equipped with canopies providing cover were monitored using time-lapse cameras for evidence of clapper rail use. Clapper rails regularly used artificial islands once tides reached heights equal to the average surface elevation of the marsh. When tides had inundated the marsh plan, observed use of the artificial islands was more than 300 times expected use based on the surface area provided. Probability of use varied among the islands and low levels of use were observed at night. Endemic saltmarsh species are increasingly at risk from habitat change resulting from sea-level rise and development of adjacent uplands. Escape cover during tidal inundation may therefore need to be supplemented if species are to survive. I developed a new method to estimate space use accounting for individual movement phases within non-stationary relocation datasets using simulated radio-telemetry data. To define movement phases I used a nonparametric, multivariate test to detect change points in the mean or variance of a sequence of x,y coordinates. Once all phases (change points) were identified, Gaussian kernel density analysis was used to estimate space use during each phase, which I termed change-point utilization distributions (CPUDs). One advantage of this technique is that the location of change points can subsequently be tested for relationships with conditions that might trigger a change in how individuals use space. Change points in clapper rail movement were associated with a variety of environmental and biotic variables including high tides, nesting activity, intrusion by neighboring clapper rails, and transient movements outside the home range. Change points occurred more than twice as frequently during the highest observed tides relative to all other tide heights. Another use of CPUDs is that space use patterns of adjacent individuals can be evaluated for joint overlap only during specific time periods when overlap occurs. I used CPUDs developed for California clapper rails and identified the point within overlapping space-use estimates where each individual had priority access to the resources within its utilization distribution (i.e. the lowest kernel density isopleth that was common to two overlapping individuals). This provided an estimate of the spatial region at which individuals exhibited territoriality. During the breeding season, space use distributions overlapped less and average territory size increased relative to the non-breeding seasons. Population density implied by these territory sizes (1.38 birds/ha) is comparable to density estimates during the 1970s and 1980s. Together these findings show the great degree to which clapper rail behavior and demography can be influenced by the tides that populations experience. It is my hope that conservation efforts for this species, particularly in the arena of habitat restoration may benefit from this research.</p>

<p> Geophagy, the consumption of mineral soil, is believed to have several benefits for herbivores. Soils high in clay are often implicated in the detoxification of plant secondary metabolites. High mineral concentrations in soils may also provide nutrients that are poorly available from plants. Local observers report that snowshoe hares (Lepus americanus) use a lick in the foothills of the Brooks Range, Alaska. Using soil from this lick and other mineral supplements, I conducted a series of feeding trials on captive snowshoe hares fed felt-leaf willow (Salix alaxensis) or a formulated ration to determine whether geophagy resulted in a physiological benefit and, if so, which soil constituents are therapeutic. When fed willow leaves, hares ate more and lost less weight when they had access to soil. Access to soil increased sodium intake and dietary ratios of sodium to potassium in hares fed willow. Soil consumption resulted in higher calcium to phosphorous ratios for both diets. Across diets, higher sodium to potassium and lower calcium to phosphorus ratios corresponded to reduced weight loss. Access to pure calcium carbonate resulted in reduced weight loss in hares fed winter dormant willow twigs, suggesting that carbonates may also be an important component of this lick. </p>

<p> After nearly being extirpated from the state, black bears in Maryland have rebounded to a point where recreational harvest has now become an important management tool. Having a better understanding of bear population parameters, movements, and harvest vulnerability allows managers to implement hunting more effectively and responsibly. To estimate demographics of the Maryland bear population, we implemented noninvasive genetic sampling of bear hair during summer 2011. We used a model-based sampling design that allowed us to collect samples more efficiently. We used presence-only maximum entropy (Maxent) modeling to classify the study area based on predicted probability of bear occurrence, and allocated the majority of our hair snares to areas with high or medium probabilities. Using microsatellite analysis and mark-recapture methods, we estimated the bear population at 701 individuals. This represents a nearly doubling of the population since the previous estimate in 2005. Our density estimate (0.25 bears/km²) is comparable to other estimates from southeastern and mid-Atlantic states. Our sampling approach did lead to more efficient sample collection, with more hair samples collected at snares located in areas with predicted high or medium probability of bear occurrence than those in low probability areas. However, in the eastern portion of our study area, where bear occurrence is presumed to be much lower, our sampling effort seemed insufficient to collect enough samples for reliable abundance estimation. As a first step toward quantifying harvest vulnerability, we used Global Positioning System (GPS) units to record movements and spatial behaviors of 108 bear hunters during the 2005&ndash;2007 Maryland bear hunting seasons. Median values showed that hunters traveled 2.9 km per hunting event, but only 0.6 km from their starting point. Hunters did not seem to show any preferential use of areas based on the landscape metrics we examined (e.g., elevation, distance from nearest road) except cover type, where 81% of locations were in deciduous forests. We found few differences between spatial behaviors of groups of hunters based on harvest success, residency, and previous bear hunting experience, as classified using post-hunt mail surveys. One notable difference is that successful hunters used steeper slopes than unsuccessful hunters. We also found that hunter perceptions of total distance traveled and distance from nearest roads were often highly inaccurate, showing that hunter surveys are not a useful tool for collecting those data. For Garrett County, Maryland, we used the hunter locations to create a Maxent model of the spatial distribution of harvest pressure. We also created a model using fall telemetry locations of female bears and compared the models to identify areas of high (i.e., high hunter and high bear occurrence) and low (i.e., low hunter and high bear occurrence) harvest vulnerability. Both models showed higher probability of occurrence on public lands. Both high and low vulnerability areas comprised small portions of the county. The low vulnerability areas included 9 larger blocks (>1 km²), which were 2.3 times steeper, 2.0 times farther from roads, and 1.5 times farther from streams than the medians for the study area. Those characteristics may limit hunter access to and use of the areas. Our predicted high vulnerability areas did not correspond to most previous bear harvest locations, indicating that our definition of harvest vulnerability often does not translate to actual harvest. Finally, we used GPS collars to track female bear locations in Garrett County and examine home range dynamics. Fixed kernel estimates for annual, spring, summer, and fall home ranges were 10.40 km², 8.93 km², 16.08 km², and 19.35 km², respectively. Fall and summer home ranges were larger than spring home ranges, but summer and fall ranges were similar. Solitary females had mean spring home ranges 6.9 times larger than females with cubs-of-the-year, but ranges did not differ during other seasons. Bears exhibited high levels of home range fidelity, with home range centroids shifting little among seasons or years. Intraspecific overlap of home ranges occurred during all 3 seasons, but was most common in summer. The results of this study provide Maryland bear biologists and managers with essential information about the state&rsquo;s bear population. Home range estimates represent important baseline information to determine appropriate spatial scales of management. The abundance estimates will be used to set proper harvest quotas with the goal of slowing the bear population growth. The hunter movement analysis and harvest vulnerability modeling may be used by managers to adjust harvest regulations to increase the efficacy of the hunting seasons.</p>

<p> The Golden-winged Warbler (<i>Vermivora chrysoptera</i>) is an imperiled migratory songbird that nests in young forest habitats of eastern North America. As such, this species has recently been the focus of an intensive multi-year, range-wide, breeding ecology study. A major focus of this research involved spot-mapping color banded males to examine relationships between nesting success and territory-scale habitat variables. I compared differences in space and habitat use of individual male Golden-winged Warblers that were monitored using both spot mapping and radio telemetry. An individual's telemetry delineated use area was on average 3.6 times larger than its spot-mapped territory. Almost half (46%) of all telemetry locations were located outside their respective male's spot-mapped territory. Number of saplings was higher in telemetry use areas (22.49 &plusmn; 2.14) than spot-mapped territories (11.80 &plusmn; 1.86). Although the exact motive for extra-territorial movements is unknown, foraging and/or suggestive observations of extra-pair copulation are likely motivating factors. The results of my study suggest Golden-winged Warblers are seeking resources outside their spot-mapped delineated territories. Furthermore, Golden-winged Warblers were found to have more telemetry locations in mature forest than found through spot-mapping. Ultimately, spot mapping alone does not accurately reflect Golden-winged Warbler space use and habitat needs.</p>

<p> My research was focused on greater sage-grouse (<i> Centrocercus urophasianus</i>; hereafter "sage-grouse") nest-site selection, nest success, and hen survival in relation to avian predators. The trade-off between using habitat and avoiding predators is a common decision for prey species including sage-grouse. In Chapter 2, I compared avian predator densities at sage-grouse nest and brood locations to random locations. Sage-grouse were located where densities of small, medium, and large avian predators were 65-68% less than random locations. </p><p> The effects of anthropogenic and landscape features on habitat use of sage-grouse hens have not been evaluated relative to avian predator densities. In Chapter 3, I compared anthropogenic and landscape features and avian predator densities among sage-grouse locations (nest, early-brood, late-brood) and random locations. I found sage-grouse hens chose locations with lower avian predator densities compared to random locations, and selected locations farther away from anthropogenic and landscape features. </p><p> Depredation of sage-grouse nests can be an influential factor limiting their productivity. Predator removal has been simultaneously proposed and criticized as a potential mitigation measure for low reproductive rates of sage-grouse. In Chapter 4, I hypothesized that sage-grouse nest success would be greater in areas where Wildlife Services lowered common raven (<i> Corvus corax</i>: hereafter "raven") density. I found that Wildlife Services decreased raven density by 61% during 2008&ndash;2011 but I did not detect a direct improvement to sage-grouse nest success. However, sage-grouse nest success was 22% when ravens were detected within 550 m of a sage-grouse nest and 41% when no raven was detected within 550 m. In Chapter 5, I assessed interactive effects of corvid densities relative to anthropogenic and landscape features on sage-grouse nest success. I found that sage-grouse nest success was positively correlated with rugged habitat. </p><p> Survival of breeding-age birds is the most important demographic parameter driving sage-grouse abundance. In Chapter 6, I evaluated the effect of raptor densities, proximity to anthropogenic and landscape features, and hen behavior on survival of sage-grouse hens. I found that sage-grouse hen survival was negatively correlated with golden eagle (<i>Aquila chrysaeto</i>s) density, proximity to anthropogenic and landscape features, and hen parental investment (nesting and brood-rearing).</p>

No description available.

<p>Rice not collected by harvesters and natural seeds are important foods for waterfowl. Estimation of abundance of these seeds is necessary for calculating waterfowl habitat conservation needs in the Louisiana Chenier Plain (LCP) and Texas Mid-Coast (TMC). My objectives were to quantify dry mass of rice and other seeds from August-November 2010, and estimate waterbird abundances on farmed and idle ricelands in these regions from December 2010-March 2011. Rice abundance in farmed ricelands ranged from 159.7 kg/ha (CV = 66.6%) to 1,014.0 kg/ha (CV = 8.3%). Natural seed abundance in idle ricelands ranged from 99.7 kg/ha (CV = 32.9%) to 957.4 kg/ha (CV = 17.2%). Greatest waterbird densities occurred in shallowly flooded disked ricelands (mean = 7.35 waterbirds/ha, 90%; CI = 2.37-19.70). Ratoon, disked, and shallowly flooded ricelands are important habitat for non-breeding waterbirds but variable estimates of seed and waterbird abundances warrant continuation of this study.

<p> West Texas, especially the Trans-Pecos region, mainly consists of desert shrubs and grasslands with patches of higher elevation (1,500 &ndash; 2,000 m) mountain ranges. Black bears (<i>Ursus americanus</i>) were extirpated from this area by the 1940s because of predator control and over hunting. In the 1980s, black bears returned to west Texas in a natural recolonization movement from Mexico, where they had survived. The black bear populations of the Trans-Pecos region and northern Mexico fit a mainland-island metapopulation model. Based on previously published research on this recolonization event, I identified several likely habitat recolonization sites and corridor routes for use in predicting possible black bear dispersal throughout the area. Then, using these corridor and recolonization scenarios, I produced a black bear metapopulation model for the Trans-Pecos region.</p><p> The possible habitat recolonization site map was created by combining 2 habitat suitability index (HSI) maps and using these HSI maps to define 'core' and 'useable' black bear habitat within the Trans-Pecos region. Using these locations, along with dispersal probabilities and black bear demographic parameters, I created a corridor dispersal map of the area using the program Circuitscape.</p><p> The metapopulation model was created using STELLA modeling software. Each recolonization location in the Trans-Pecos region (Big Bend National Park, Black Gap Wildlife Management Area, and the Davis Mountains) has its own black bear subpopulation. The metapopulation model is a stochastic compartment model based on a yearly time step (&Delta;<i>t</i> = 1 yr). This model was tested for the effects of: carrying capacity per site, immigration rates from Mexico, rates of dispersal from Black Gap Wildlife Management Area to the Davis Mountains, and the recovery time for the area after complete extirpation from the Trans-Pecos. This information will help local biologists conserve and manage these returning black bears in the Trans-Pecos region. </p>

<p>I investigated percent coverage, plant height, species richness, and woody stem density in plant communities in ten study plots during spring and fall (2010&ndash;2012) within 3 different treatments (continual mowings, one fall mowing, and one fall mowing with native wildflower seeds) on Highway 25 right-of-way in Oktibbeha and Winston counties, Mississippi. I recorded 277 plant species including native and non-native forbs, legumes, grasses, rushes/sedges, and woody plants. Non-native agronomic grasses exhibited greatest coverage greater than 90 percent occurring in all treatments. Percent coverage of plants less than 0.46m height category exceeded 100 while, greater than 0.46m plant height categories averaged 55 percent. Woody stem density ranged from 7,772 year 1 to 10,025 stems/hectare year 2. I detected no significant differences in plant height or woody stems among treatments. One mowing per year retained agronomic plant cover for erosion control and annual cost savings up to 75 percent for roadside maintenance. </p>