No description available.

No description available.

No description available.

No description available.

This research seeks to pinpoint the telencephalic structures most closely correlated with feeding flexibility, which is operationalised as feeding innovation rate per taxon. By calculating a weighted average per taxon of 1030 feeding innovations collated from five zones of the world (western Europe, North America, Australia, New Zealand and India), the study shows that relative size of the hyperstriatum ventrale and, to a lesser extent, the neostriatum , best predicts weighted innovation rate; these two structures are thought to be functionally equivalent to the mammalian neocortex. The worst telencephalic predictors of innovation rate are two structures respectively thought to be involved in primary visual projection and the control of stereotyped, species-specific behaviour, the wulst area and the paleostriatum. / A second presumed correlate of behavioural flexibility, taxonomic variation in the use of urbanised and other anthropogenically-modified habitats, shows a consistent pattern in four geographical zones (Great Britain, North America, Australia and New Guinea), as well as a qualitative association with the relative size of the forebrain. There is no linear correlation, however, between urbanisation rate per taxon and either forebrain size or innovation rate, suggesting that other variables like diet, tameness and neophilia may have to be quantified at finer taxonomic levels in future studies of opportunism in habitat use.

This thesis seeks to determine if juvenile birds preferentially select foraging information from their parents, by examining the acquisition of novel food-finding skills in juveniles of a flock-feeding species, the ringdove (Streptopelia risoria). / The first experimental examined the flock feeding behaviour of juvenile ringdoves foraging in a small aviary flock composed of kin and non-kin; frequencies of local enhancement, food begging and aggression were recorded, as well as the individuals that the juvenile associated with while searching for food. Juvenile ringdoves foraged significantly more often with their kin and were aggressed more by non-kin. The last three experiments tested juveniles on three components of foraging: novel food type, environmental colour cues associated with food and novel food-searching techniques. All three involved a choice-test where the juveniles had their father and an unrelated flock member as demonstrators. All three choice-tests showed there was no preference for selecting either demonstrator's solution. Juvenile doves in the field may appear to learn from their parents simply because they associate more with them. (Abstract shortened by UMI.)

This thesis investigated the use of song repertoires during territorial defense of the song sparrow, Melospiza melodia. The functional and adaptive significances of repertoires in the order Passeriformes were reviewed. Within the family Fringillidae, an examination of repertoire size and life history features of different species revealed no significant correlations. / Observations of six neighboring song sparrows yielded information about the relative uses of repertoire-dependent and repertoire-independent mechanisms in coordinated song interactions, as well as in signalling agonistic probabilities. The behavior of song switching, independent of the identities of song types, was found to most effectively perform both types of functions. / A series of playback experiments tested and supported the hypothesis that the song sparrow's rate of switching song types increases with agonistic stimulation. Song versatility, rather than switching rate per se, was found to be an appropriate agonistic stimulus.

This study investigates the colonization of Barbados, the seasonality of breeding, and the function of breeding plumes in the cattle egret Bubulcus ibis ibis. Colonization occurred around 1956, and the island now has 4 colonies. Numbers of birds at each colony are still increasing. Colonies are evenly spaced around the island. Plumage scores varied seasonally, and males had higher scores than females. Breeding activity peaked in July-August and November-March. Conspecific interference may force poorly plumed birds to breed when environmental conditions are poor. Chick mortality was primarily from starvation during periods of low breeding activity, and primarily from conspecific attack during high breeding activity. Fledging success did not increase with breeding activity. Plumage scores correlated positively in breeding pairs. Well-plumed males fed chicks more often than poorly plumed males and had the higher fledging success. Males contributed more to nest and nestling attendance when their mates were well plumed. Plumes appear to indicate the potential paternal care and the genetic quality of a mate.

The variable plumage characteristics of the sub-adult male cohort of the red-winged blackbird (Agelaius phoeniceus) are described. At one extreme of the plumage variation, there are a group of sub-adult males that are indistinguishable from the adult males. These birds can only be correctly aged by cloacal examination for the bursa of Fabricius in the autumn. At the other extreme, approximately 4% of the population are near perfect female-mimics. A simple scoring system based on the interspersion of dark feathers in the epaulet is presented, which permits classification of the sub-adult males into 1 of 6 plumage classes. These epaulet classes are significantly correlated with other traits of the plumage. / Age when entering the prebasic molt, and the physical condition of the sub-adult male may influence the development of the varied plumage characteristics. / The distribution of the plumage characteristics of a population of sub-adult males collected at a major blackbird roost in the province of Quebec is described for birds captured in the fall and spring. The spring plumage characteristics are more variable than the fall, a phenomenon which is not consistent with plumage wear as has been previously reported, but which may be related to a prealternate molt which the birds undergo. The prealternate molt is prevalent in some but not all of the contour feather tracts, and is restricted to females and the sub-adult male cohorts. / An age- and sex-specific spring migration of red-winged blackbirds is examined. Adult males arrive to the spring roosts first, followed by yearling males and then females. A similar pattern of dispersal to the breeding territories is described. Within the subadult male cohort, the birds with the most adult-male like plumage traits arrive at the vernal roosts first. / The characteristics of the prealternate molt and differential spring migration are discussed in relation to the pressures of sexual selection on the respective age and sex cohorts. / The adaptive significance of variable sub-adult male plumages and delayed plumage maturation in passerines is evaluated. Several competing hypotheses have been advanced to describe the phenomenon of delayed plumage maturation. These hypotheses are reviewed, and a test is proposed which unequivocally differentiates between the various alternatives. (Abstract shortened with permission of author.)

1. Cost constraints in models of territory size are based on time/activity/laboratory estimates that predict birds using larger territories will incur higher energy costs. The predicted form of the cost constraint may be linear, accelerating or decelerating depending on assumptions inherent in the models. The aim of this study was to assess the reality and form of the cost constraint by making direct measurements of the energy costs of territory use in birds that occupy territories of different size and shape; polygonal territories represented by the robin Erithacus rubecula, and linear by the dipper Cinclus cinclus. Free-living energy expenditure was measured using the doubly-labelled water technique, whilst simultaneously recording patterns of territory use by radio-tracking. 2. Territorial robins concentrated their activity in one or more foraging patches located in bushes. Range polygons containing all the foraging patches used by an individual provided estimates of territory area, and were generally of high eccentricity. A small proportion of robins was classified as non-territorial based on range polygon areas. Furthermore, while territorial robins showed high fidelity to ranges over the short term (days), non-territorial individuals were nomadic. Over the longer term (months), however, some territorial robins showed range drift. Dippers similarly used preferred core regions within ranges, although there was no selection for particular habitat features. 3. Because robins occupied territory polygons which varied from polygonal to highly linear, work was focused on this species to allow intra-specific comparison. Robins tended to commute between foraging patches by flying. It was appropriate, therefore, to describe territories in terms of a number of patches linked by a network of flight paths. This generated two further measures of territory size; the number of patches used and the total flight distance between patches. 4. The robins exploited a renewing food supply. Predictions were tested concerning the temporal scheduling of visits to foraging patches within territories. Patches tended to be separated by flight paths of similar lengths, and were visited in a regular sequence. Although the number of foraging patches used varied, all territories had similar total core areas. Robins using many small foraging patches commuted between patches more often and covered a larger total flight distance during each foraging circuit of the territory. The configurations of foraging patches were used in a highly linear manner. This was true even if the territory containing them was of low eccentricity. 5. Changes in structure and pattern of use varied predictably with territory size, and could be described mathematically. Based on this and published time/activity budgets, a suite of models was developed to predict how energy costs would vary with number of patches used and total flight distance between patches. Models were tested by directly measuring the energy expenditure of robins using different territories. The number of patches used and total flight distance between patches were both significantly correlated with energy expenditure, while territory area was not. One of the models showed a significant fit to the observed data, and suggested that the form of the energy cost constraint on territory size was linear. The effect of territory shape on energy costs was minimal. The implications of these results for models of territory size are discussed. 6. The slope and elevation of the energy cost constraint varied with the morphology of territory occupants. Based on this, an association of morphology with territory size was predicted; robins of lower mass and wing-loading using larger territories. The observed data supported these predictions, and suggested a possible genetic predisposition to particular patterns of territory occupancy in the robin.