Dissertation (PhD)--University of Stellenbosch, 2006. / ENGLISH ABSTRACT: The spatial distribution of animals and plants at different scales is a central theme in ecology. Knowledge of biodiversity distribution is essential, especially with the current threat of climate change and invasion by alien species. Since the impact of climate change and alien species will be, and has already been pronounced in polar regions, information on the current spatial distributions of biota in these regions is critical to predict the consequences of climate change and alien species on the future survival, distribution and abundance of indigenous biota. This study was conducted on the sub- Antarctic Prince Edward Islands (consisting of Marion Island and Prince Edward Island), which have experienced rapid climate changes over the past 50 years. Additionally, a number of alien plant, vertebrate and invertebrate (also microarthropod) species have been introduced to these islands. Since microarthropods (mites and springtails) play an essential role in decomposition and mineralization of plant material on the islands, the loss of microarthropods from decomposer communities might have disastrous results for ecosystem processes. Therefore it is essential to know the current distributions of microarthropods in order to predict future distributions patterns in reaction to climate change and invasive species. In this study, the spatial distribution of mites and springtails inhabiting the cushion plant Azorella selago Hook, were examined at different scales of observation. Firstly, the microarthropod community was examined at a fine scale. The relationship between microarthropod species richness and abundance and plant size, isolation, within-plant variability, grass cover and microclimate variables were investigated. Thereafter, the spatial variability of microarthropod abundances was examined within stands of plants, with statistical methods using varying degrees of locational information to determine if microarthropod abundance is random, regular or aggregated at this scale. Further, the spatial variability of microarthropod communities in A. selago at different altitudes and on different sides of Marion Island, i.e. island-wide scale, was examined. The last scale of observation was the island-wide scale, in which microarthropod assemblages were compared between Marion Island and Prince Edward Island, as well as in different plant species. Microarthropods showed spatial heterogeneity within A. selago plants (higher abundances in eastern and southern sides of plants), island-wide (lower springtail abundance on the eastern side of the island and at high altitudes) and between islands (more alien species on Marion Island). In contrast, microarthropod abundance was not spatially heterogeneous within a stand of plants, i.e. mostly randomly distributed. The possible mechanisms responsible for these patterns differ between scales, and range from temperature and nutrient availability at the within plant and island wide scale to alien species between islands. Climate change is likely to alter these distribution patterns of microarthropods, either directly (unfavourable climate for indigenous species growth, development and dispersal) or indirectly (favouring alien microarthropod species; increase in grass cover on A. selago plants may alter community structure). The information presented here, is essential for predicting the possible impacts of climate change on microarthropods in fellfield ecosystems on sub-Antarctic islands. / AFRIKAANSE OPSOMMING: Die ruimtelike verspreiding van diere en plante op verskillende skale is ‘n sentrale tema in ekologie. Kennis van die verspreiding van biodiversiteit is belangrik, veral met die huidige bedreiging van klimaatsverandering en indringerspesies. Omdat die impak van klimaatsverandering en indringerspesies in die poolstreke alreeds duidelik is en nog meer so sal word, is inligting oor die huidige ruimtelike verspreiding van biota in hierdie streke noodsaaklik. Hierdie studie is op die sub-Antarktiese Prins Edward-eilande (bestaande uit Marion Eiland en Prins Edward Eiland) uitgevoer, waar die klimaat vinnig verander het oor die afeglope 50 jaar. Bykomend is ‘n aantal indringer plant, vertebraat en invertebraat (ook mikroartropood) spesies op die eilande ingebring. Omdat mikroartropoda (myte en springsterte) ‘n baie belangrike rol in ontbinding en mineralisering van plantmateriaal op hierdie eilande speel, sal die verlies van mikroartropoda uit die ontbindingsgemeenskap rampspoedige gevolge vir die prosesse in die ekosisteem hê. Kennis van die huidige verpreiding van mikroartropoda is dus noodsaaklik om die toekomstige verspreidingspatrone, asook oorlewing en volopheid van inheemse spesies in reaksie op klimaatsveranding en indringerspesies, te voorspel. In hierdie studie is die ruimtelike verspreiding van myte en springsterte wat in die kussingplant Azorella selago Hook voorkom, op verskillende ruimtelike skale ondersoek. Eerstens is die mikroartropood gemeenskap op ‘n klein skaal ondersoek. Die verwantskap tussen mikroartropood spesies rykheid en volopheid en plantgrootte, isolasie, binne-plant variasie, gras bedekking en mikroklimaat faktore is ondersoek. Daarna is die ruimtelike variasie van mikroartropood volopheid binne in ‘n plantstand ondersoek deur gebruik te maak van metodes wat verskillende grade van ligging data gebruik. Dit is gedoen om uit te vind of mikroartropood volopheid lukraak, reëlmatig of versameld verspreid is. Verder is die ruimtelike verspreiding van mikroartropood gemeenskappe in A. selago op verskillende hoogtes bo seespieël en op verskillende kante van Marion Eiland ondersoek. Laastens is die mikroartropood gemeenskap op Marion Eiland en Prins Edward Eiland, asook die gemeenskap in verskillende plant spesies vergelyk. Mikroartropoda is ruimtelik heterogeen verspreid binne-in A. selago (meer individue in die oostelike en suidelike kante van plante), asook regoor Marion Eiland (minder springstert individue aan die ooste kant van die eiland asook minder op hoë hoogtes bo seespieël) en tussen eilande (meer indringerspesies op Marion Eiland). In teenstelling hiermee is mikroartropood volopheid ruimtelik homogeen oor ‘n plantstand, dus meestal lukraak verspreid. Die meganismes wat moontlik vir hierdie patrone verantwoordelik is wissel van temperatuur en voedselbeskikbaarheid op die binne-plant en oor die eiland skaal tot indringerspesies op die tussen-eiland skaal. Klimaatsverandering mag waarskynlik mikroartropood verspreidingspatrone direk (ongunstige klimaat vir inheemse spesies se groei, ontwikkeling en verspreiding), of indirek verander (indringer spesies word begunstig; toename in grasbedekking bo-op A. selago mag die gemeenskapstruktuur van mikroartropoda verander). Hierdie inligting is belangrik vir die voorspelling van die moontlike impak van klimaatsverandering op mikroartropoda in klipveld ekosisteme op sub-Antarktiese eilande.

Thesis (PhD(Agric) (Conservation Ecology and Entomology))--University of Stellenbosch, 2010. / ENGLISH ABSTRACT: See full text document for abstract / AFRIKAANSE OPSOMMING: sien volteks dokument vir die opsomming

Thesis (PhD (Conservation Ecology and Entomology))--University of Stellenbosch, 2010. / ENGLISH ABSTRACT: Invasive alien trees (IATs) and agriculture are rapidly altering the capacity of ecosystems to provide a variety of essential services, with decreasing habitat quality having an adverse effect on arthropod biodiversity. Although both agricultural intensification and IATs have an impact on ecosystems, it is unclear which of these two is currently the most severe. Here I compare the influences of vineyards and IATs on arthropod diversity in the Cape Floristic Region (CFR) from two perspectives. Firstly, insect diversity from four different vegetation types (native fynbos, IATs, cleared invasive alien trees (CIATs) and vineyards), and their ecotones were assessed. Secondly, soil compaction in the different vegetation types was assessed and compared to determine how soil compaction relates to arthropod species richness and abundance. Surface-active arthropods were used for these evaluations, as they were considered to be the most abundant, yet sensitive groups for these comparative assessments. Pitfall trapping, which has been widely used for such comparative studies, was the method of choice. Additionally, as a result of incidental wildfires at some of the study sites (IATs and fynbos vegetation), it was decided opportunistically to assess recovery of arthropod diversity following the impact of fire. The results categorically indicate that IATs had by far the most significant impact, in comparison with vineyards, in reducing arthropod diversity and changing assemblage composition. Both IATs and vineyards, not surprisingly, supported lower arthropod diversity compared to fynbos, while CIATs supported arthropod species richness comparable to that of fynbos. However, different invertebrate species responded differently to IAT invasion, with some species even adapted to conditions in IAT patches. Environmental factors such as leaf litter and soil compaction were strongly correlated with arthropod species richness. However, other factors, such as soil moisture content, were not correlated with species richness, despite varying significantly between different vegetation types. IATs were found to reduce soil compaction, while vineyards increased soil compaction. The ecotone between adjacent land-use types was important for sustaining high arthropod species richness, particularly for the CIATs/fynbos ecotone, followed by the interface between CIATs/vineyard and vineyard/fynbos. Some species even preferred ecotones. Surface-active invertebrate species richness declined substantially immediately after fire in both IATs and fynbos. Nevertheless, recovery was rapid, with some species recolonizing or re-establishing within three months. Indeed, both species richness and abundance were high following fire. Fire had no significant impact on arthropod assemblage within IATs, despite having a serious impact in the fynbos where a different arthropod assemblage was observed after fire. The recovery of the historic assemblages may be dependent on both the pre-fire composition of the habitat and its adjacent vegetation. Effective co-operation among conservation biologists, farmers, and landowners to actively plan the future of CFR landscape biodiversity is required. / AFRIKAANSE OPSOMMING: Indringer uitheemse bome (IUB) en landbou bedrywighede is vinnig besig om die kapasiteit van ekosisteme vir die voorsiening van „n verskeidenheid van noodsaaklike dienste, met „n afname in habitatkwaliteit en „n nadelige uitwerking op artropood biodiversiteit, te verander. Alhoewel beide, landbou intensifikasie en IUB‟s, ‟n impak op ekosisteme uitoefen, is dit onduidelik watter van hierdie twee huidig die mees ernstige een is. In hierdie studie vergelyk ek die invloed van wingerde en IUB‟s op artropood diversiteit in die Kaapse Floristiese Streek (KFS) vanuit twee perspektiewe. Eerstens, insekdiversiteit van vier verskillende vegetasietipes (natuurlike fynbos, IUB‟s, verwyderde indringer uitheemse bome (VIUB‟s) en wingerde) en hulle ekotone is bepaal. Tweedens, is grondverdigting in en van die verskillende vegetasietipes ondersoek en vergelyk om te bepaal hoe grondverdigting verband hou met artropood spesies rykheid en volopheid. Oppervlak-aktiewe artropode is vir hierdie bepalings gebruik aangesien hulle beskou word as die mees volopste, maar ook mees sensitiewe groepe vir hierdie vergelykende bepalings. Pitvalvangste, wat alom gebruik word vir sulke vergelykende studies, was die verkose werkswyse. Bykomend, as gevolg van onbeplande veldbrande in sommige studie gebiede (IUB‟s en fynbos plantegroei), is oppertunisties van die geleentheid gebruik gemaak om die herstel van artropood diversiteit na die impak van brand te bepaal. Die resultate toon duidelik aan dat IUB‟s, in vergelyking met wingerde, by verre die mees beduidende impak gehad het deur ‟n verlaging van artropood diversiteit en veranderende groeperingssamestellings. Beide, IUB‟s en wingerde, het, nie onverwags nie, in vergelyking met fynbos, laer artropood diversiteit ondersteun, terwyl VIUB‟s artropood spesies verskeidenheid, vergelykbaar met dié van fynbos, ondersteun het. Verskillende invertebrate spesies het egter verskillend gereageer op IUB indringing, en sommige spesies het selfs by toestande in IUB kolle aangepas. Omgewingsfaktore soos blaarafval en grondverdigting was sterk gekorrelleer met artropood spesies rykheid. Ander faktore, soos grondvoginhoud, was egter nie in ooreenstemming met spesies rykheid nie, ten spyte van ‟n beduidende wisseling tussen verskillende vegetasietipes. IUB‟s het grondverdigting verminder, terwyl by wingerde grondverdigting toegeneem het. Die ekotone tussen aanliggende grondgebruike was belangrik om hoë artropood spesies rykheid, veral by die VIUB‟s/fynbos ekotoon, gevolg deur die kontaksones tussen VIUB‟s/wingerd en wingerd/fynbos, te onderhou. Sommige spesies het selfs ekotone verkies. Oppervlak-aktiewe invertebraat spesies rykheid het onmiddelik na brand in beide, IUB‟s en fynbos, wesenlik afgeneem. Tog was herstel vinnig, met sekere spesies wat binne drie maande teruggekeer of hulle hervestig het. Inderdaad was beide, spesies rykheid en volopheid, hoog na ‟n brand. Brand het geen noemenswaardige invloed op artropood groeperings binne IUB‟s gehad nie, ten spyte van ‟n ernstige impak in fynbos waar ‟n verskillende artropood groepering na ‟n brand waargeneem is. Die herstel van die historiese groeperings van beide, die voor-brand samestelling van die habitat en sy aangrensende plantegroei, mag afhanklik wees. Effektiewe en aktiewe samewerking tussen bewaringsbioloë, boere, en grondeienaars om die toekoms van KFS landskap biodiversiteit te beplan, word benodig.

Thesis (PhD (Conservation Ecology and Entomology))--University of Stellenbosch, 2011. / ENGLISH ABSTRACT: Codling moth, Cydia pomonella, is a worldwide pest and of major economic importance to the South African pome fruit industry. Sterile insect release is applied as a component of area-wide integrated pest management and includes the mass-rearing, sterilization and the release of the sterile insects. For sterile insect release, the improvements of rearing methods in terms of the quality of the diet ingredients and the economical aspect of the rearing method are examined. The effect of genetically modified maize meal, containing the Bacillus thuringiensis gene, in an artificial medium for codling moth rearing, is determined. The use of even a small amount of Bacillus thuringiensis resulted in larval mortality and prolonged development. These results are detrimental to a mass-rearing facility and must be considered by any rearing facility that uses genetically modified maize meal if the insect is sensitive to the gene. An alternative to maize meal in the artificial medium was tested and whole wheat flour was considered to be a suitable replacement. Agar agar is an expensive gelling agent used in the artificial medium. An alternative for agar agar (Kelcogel, Elastigel and carrageenen) is tested and the biological effect on codling moth is determined. Factors such as mortality, pupal and moth weight, longevity, fecundity and development time were used as quality parameters. Results showed that Elastigel was a suitable replacement for agar agar, with bigger pupae and moths, higher fecundity and increased longevity. The economical advantage of the replacement is a 40.91% reduction of the diet cost. The other gelling agents tested also gave acceptable results and can be considered if shortages of agar agar or Elastigel occur. A new method of mass-rearing codling moth larvae in a closed rearing system using large trays placed in a ventilated box is designed. This method is more cost and space effective as a smaller area is needed to rear a large number of moths. The risk of diet contamination is less because of the closed environment and more economical and effective air handling. This is the first report of its kind to describe the mass-rearing of codling moth in a closed environment and the risks involved in using genetically modified maize meal in an artificial diet for the codling moth. These results should be incorporated into existing mass-rearing facilities or taking into consideration when designing new mass-rearing facilities. / AFRIKAANSE OPSOMMING: Die kodlingmot, Cydia pomonella, is van ekonomiese belang vir die Suid-Afrikaanse kernvrugte bedryf. Die steriele insek tegniek word gebruik as ‘n komponent in area-wye geïntegreerde plaagbeheer en sluit in die massa-aanteel, sterilisering en vrylaat van steriele insekte. Vir die steriele insek tegniek is die verbetering van die massa-aanteel van die kodlingmot in terme van kwaliteit van die dieet en die ekonomiese aspek van die aanteel metode ondersoek. Die effek van genetiese gemanipuleerde mieliemeel wat die Bacillus thuringiensis geen bevat, in ‘n kunsmatige voedselmedium vir die aanteel van kodlingmot, is bepaal. Daar is gevind dat die gebruik van selfs ‘n klein persentasie Bacillus thuringiensis in die mieliemeel, mortaliteit en ‘n verlengde lewenssiklus in kodlingmot veroorsaak. Die gevolge is nadelig vir ‘n massa-aanteel fasiliteit en behoort in ag geneem te word vir enige insek wat op ‘n kunsmatige medium, wat mieliemeel bevat, geteel word, mits die insek sensitief is vir Bacillus thuringiensis. ‘n Alternatiewe bestanddeel vir mieliemeel, volkoringmeel, word aanbeveel. Agar agar is ‘n duur verdikkingsagent wat in kunsmatige mediums gebruik word. ‘n Alternatief vir agar agar (Kelcogel, Elastigel en carrageenen) is getoets en die biologiese effek op die kodlingmot is bepaal. Faktore soos mortaliteit, papie en mot gewig, langlewendheid, vrugbaarheid en lengte van lewenssiklus was gebruik as kwaliteit parameters. Resultate het getoon dat Elastigel ‘n geskikte plaasvevanger is van agar agar, met groter papies en motte, groter vrugbaarheid en langlewendheid. Die ekonomiese gevolg van die plaasvervanger, is ‘n vermindering van 40.91% van die dieetkoste. Die ander verdikkingagente wat is getoets is, het aanvaarbare resultate gelewer wat noodsaaklik is indien daar ‘n tekort van Elastigel of agar agar ontwikkel. ‘n Nuwe metode van massa-aanteel van kodlingmot larwes is bepaal. Die metode behels ‘n geslote sisteem, waar groter aanteel bakke in ‘n geslote, geventileerde boks geplaas word. Die metode is koste en spasie effektief en ‘n kleiner area word benodig om ‘n groter aantal motte te lewer. Die risiko van kontaminasie van die dieet word verminder as gevolg van die geslote sisteem wat gebruik word en meer ekonomiese en effektiewe lugversorging word gebruik. Hierdie is die eerste verslag van sy soort wat die massa-aanteel van kodlingmot in ‘n geslote sisteem beskryf en wat die risiko aandui van geneties gemanipuleerde mieliemeel in ‘n kunsmatige medium vir die kodlingmot. Hierdie resultate behoort in ag geneem te word vir reeds bestaande massa-aanteel fasiliteite of met die ontwerp van nuwe massa-aanteel fasiliteite.

Thesis (PhD (Conservation Ecology and Entomology))--University of Stellenbosch, 2011. / ENGLISH ABSTRACT: Water storage in reservoirs forms an integral part of the agricultural landscape in the Western Cape Province, South Africa. A few large reservoirs serve primarily as a drinking and industrial water supply, while on private farms, small reservoirs provide irrigation water for the dry summer period. Protection of water quality to secure irrigation and drinking water quality, and the extension of water use efficiency are priority issues in the Western Cape. In the current study, the suitability of rainbow trout (Oncorhynchus mykiss) cage farming as a non-abstractive water use was investigated. The current study concentrated on the identification, and where possible quantification of aquaculture impacts, the identification of successful sites and a description of requirements in which net-cage aquaculture has none or a very low negative impact on water quality (e.g. <15 % change from previous water quality conditions for phosphorus concentrations). In order to study the effects of 5 t trout cage production units in reservoirs <15 ha in area, the general ecology of the irrigation reservoirs was assessed. Sixteen reservoirs without aquaculture production (reference reservoirs) were compared to 26 reservoirs with aquaculture production (production sites with varying production histories). Catchment characteristics were also monitored. Water from different depths (0 m, 2 m, 6 m and near bottom) was tested for physical and chemical qualities as well as nutrient concentrations. Phytoplankton and zooplankton biomass and species composition was monitored. In addition to the general phytoplankton findings, cyanophytes were evaluated for their contribution to algal taint problems that emerged at a number of production sites. Sediments were tested for total phosphorus content and phosphorus release capacity. Indicators and minimum conditions to avoid the most common production problems were formulated. In order to determine long-term production success, which prevents trophic level changes of reservoirs, a mass balance approach (nutrient budget) was employed to indicate the limits for nutrients that can be added. The phosphorus balance indicated long term trends for reservoirs with and without aquaculture. The detailed mass balance approach was compared to a “ready to use” carrying capacity model that estimated the maximum fish load each reservoir could support. The anthropological input of phosphorus into the reservoirs causes a decreasing water quality in the studied reservoirs and this development was also reported for lowland rivers. Twenty percent of the studied reservoirs are in a condition that could be an immediate threat to fish or water bird health (e.g. free ammonia concentrations and pH). Harmful algal blooms were not observed. Aquaculture production evoked changes in water chemistry and ecology in most of the studied reservoirs. Adverse effects of aquaculture sites versus non-aquaculture sites were: increased phytoplankton biomass and species shifts towards sizes >80 μm. The increased phytoplankton abundance influenced pH maxima to values >9 at mid-day. The high pH fluctuations were greatly influenced by the naturally low alkalinity and hence low calcium buffering capacity of Western Cape waters. The deoxygenation of the hypolimnion during stagnation (summer) occurred faster in reservoirs of certain character, greatly dependent on elevation and surface area, with consequent acidification of the hypolimnion, as well as ammonia and total phosphorus (TP) accumulation. In this context, a diversity of each reservoir with and without aquaculture production, with a similar ratio of undisturbed reservoirs to reservoirs with influence of e.g. agriculture, were compared to each other. When grouping the respective differences from the average reference reservoir (i.e. no trout production), a low impact on water quality was manifested at four sites (15 %) with <15 % increase of bottom TP and ammonia, while eight sites (31 %) showed medium effects (59 % mean increase), and a high impact was found at 54 % of the sites (312 % mean increase). In reservoirs without aquaculture, the extent of incoming phosphorus (which could represent an influence by runoff from agricultural land) was very high. However, in small reservoirs (<5 ha), these values were exceeded by the incoming phosphorus from aquaculture practices. In the case of small reservoirs where the carrying capacity was clearly surpassed, effects caused by aquaculture were severe and the assimilation of waste by the system was not possible (in extreme cases aquaculture waste delivered 60 to 90 % of all incoming phosphorus – two to nine times the phosphorus brought in by rivers and runoff). With regards to sediment, only indirect conclusions could be drawn. Aquaculture production increased hypolimnetic anoxia and the latter was shown to increase potential phosphorus release from sediments. This implies that not only will aquaculture increase the phosphorus concentration of surface waters directly, but it will also increase internal loading. The sedimentation rate was increased with cage aquaculture which affects a hypothesized area of approximately 0.2 to 1.0 ha depending on reservoir hydrology. The composition of the sediment increased organic components which can impact on sediment processes. It can be postulated that increased sedimentation of aquaculture waste and extended anoxic conditions impact on macrozoobenthos. Hydrological and nutrient mass exchange of the reservoirs indicated that no annual increase of phosphorus was achieved with low nutrient input (good inflowing water quality) or good water exchange (>5 times per year), and sometimes with extraction of hypolimnetic water during the stagnation period (summer). A model developed by Beveridge (1984) showed similar results to the mass balance approach and can therefore serve as a more ready model to determine suitable stocking rates. The small (man-made) reservoirs in the Western Cape are in a eutrophication process which far exceeds the speed of natural eutrophication (trophic states indicating highly eutrophic or hypertrophic conditions after approximately 10 to 20 years following construction of the reservoirs) and this process is triggered by agricultural practice (indicated by literature – not a subject of this study). However, it is concluded from the data of this study that trout-cage aquaculture duplicated the total phosphorus already present (independent of continuation of the projects, the phosphorus introduced was trapped in the closed systems the reservoirs represent) in only 1 to 2 years of production - which means a significant acceleration of the eutrophication process already in place. There are positive exceptions where trout-cage production is possible without negative effects. Careful site selection is the most important step in successful and sustainable trout production. No impact of aquaculture was recorded at four reservoirs (15 % of the investigated reservoirs) which shared the characteristics of good water exchange (>3 times per year) and a minimum surface area of 5 ha. Additionally, criteria that reduced the risk of algal taint included a minimum water depth of 6 to 7 m in a reservoir at its lowest water point (to avoid intermediate mixing during the stagnation period) as well as cold hypolimnetic conditions (<17 °C) to minimize cyanophyte cyst remobilization. Further improvement of food conversion ratio (feeding management) and feed quality are the next (after site selection) two most important components that determine if a reservoir can be utilised for cage production without any long-term changes. There is potential in advances in feed quality, feed management and waste collection systems. These measures (e.g. the cage size could be decreased to efficate feeding management) can increase the number of sustainable sites and achieve multiplication of water use without water quality deterioration. / AFRIKAANSE OPSOMMING: Die stoor van water in reservoirs vorm ‘n integrale deel van die landboulandskap in die Westelike Kaap gedeelte van Suid-Afrika. ‘n Aantal groter reservoirs voorsien primêr water primer vir drinkdoeleindes en vir aanwending deur industrieë, terwyl kleiner reservoirs op private plase gebruik word vir besproeiing gedurende die droë somerperiode. Die beskerming van die waterkwaliteit en die uitbreiding van watergebruik se doeltreffendheid, word bestempel as ‘n prioriteit vir die Wes-Kaap. In die huidige studie is die geskiktheid van forel hokkultuur as ‘n bykomende watergebruik, maar sonder waterverbruik, ondersoek. Die huidige studie fokus op die identifisering en waar moontlik die kwantifisering van die impak van akwakultuur op besproeiingsreservoirs; die identifisering van potensiële reservoirs vir akwakultuur; en die beskrywing van toestande waarin akwakultuur geen of ‘n baie lae negatiewe uitwerking op die waterkwaliteit (bv. < 15 % verandering vanaf oorspronklike waterkwaliteitstoestande vir fosforkonsentrasies) sal hê. Om die impak van 5 t forel hokproduksie eenhede in <15 ha reservoirs te bestudeer, is die algemene ekologie van die besproeiingsreservoirs bepaal. Watermonsters, geneem op verskillende dieptes (0, 2, 6 en naby die bodem), is getoets vir fisiese- en chemiese kwaliteit sowel as vir die nutriëntkonsentrasies. Die biomassa en spesiesamestelling van fitoplankton sowel as zooplankton is gemonitor. Sestien verwysingsreservoirs is vergelyk met 26 produksiereservoirs (met wisselende vlakke van produksiegeskiedenis). Die karakeristieke eienskappe van die opvangsgebied is ook gemonitor. Tesame met die algemene fitoplankton bevindinge, is blougroen alge ook geëvalueer vir hul bydrae tot algae besmettingsprobleme wat voorgekom het by ‘n aantal produksie persele. Sedimente is getoets vir die totale fosforinhoud en fosforvrystellingskapasiteit. Perseelseleksie is geidentifiseer as een van die mees belangrikste faktore vir suksesvolle hokproduksie. Indikatore en minimum toetstande wat vereis word om die mees algemene produksieprobleme te verhoed, is geformuleer. Om die langtermyn produksie sukses te bepaal, wat trophiese vlakke se verandering in die reservoirs verhoed, is ‘n massa balans benadering (nutriënt balans) gebruik. Die fosfor balans het langtermyn tendense aangetoon en in die geval van die produksie reservoirs, is die akwakultuurimpak gekwantifiseer. Die gedetaileerde massa balans benadering is vergelyk met ‘n “gereed om te gebruik” drakapasiteitsmodel wat die maksimum vis wat die reservoir kan onderhou, geskat het. Die antropogeniese toevoer van fosfor na die reservoirs veroorsaak ‘n afname in die waterkwaliteit van die reservoirs wat bestudeer is. Die verskynsel van ‘n afname in waterkwaliteit is ook vir verskeie laagliggende riviere geraporteer. Sowat 20 % van die besproeiingsreservoirs wat bestudeer is, is tans in ‘n toestand wat die onmiddelike gesondheid van vis en watervoëls kan bedreig. Skadelike algebbloei is nie waargeneem nie. Akwakultuurproduksie het negatiewe effekte getoon in ‘n aantal reservoirs. Die nadelige uitwerking van akwakultuur waar digthede te hoog was, is: toename in fitoplankton biomassa en spesiesverskuiwinge. Die toename in fitoplankton teenwoordigheid het die pH maxima beïnvloed tot waardes >9 teen 12 uur middag. Die hoë pH fluktuasies is grootliks beïnvloed deur die natuurlike lae alkaliniteit en gevolglike kalsium bufferkapasitiet van die Wes-Kaap se waterbronne. Die deoksiginasie van die hipolimnion gedurende stagnasie (somer) het vinniger plaasgevind in oorbelaaide reservoirs, met gevolglik ‘n versuring van die hipolymnion, sowel as die akkumelering van ammoniak en totale fosfor. In hierdie konteks word reservoirs met en sonder landbouproduksie, met ‘n gelyke verhouding van onversteurde reservoirs tot reservoirs wat deur landboubedrywighede beinvloed word, met mekaar vergelyk. By vier persele (15 %) is ‘n lae impak vasgestel (<15 % toename in bodem TF en ammoniak), terwyl agt persele (31 %) ‘n medium impak getoon het (59 % gemiddelde toename in bodem TF en ammoniak) en ‘n hoë impak is opgemerk by 54 % van die persele (31 % gemiddelde toename in bodem TF en ammoniak). In reservoirs sonder akwakultuur, was die omvang van fosforinvloeiing baie hoog en is moontlik veroorsaak deur die invloei van afloopwater uit omliggende landbougebiede. Alhoewel in klein reservoirs (<5 ha) was hierdie waardes oorskadu deur die invloeiing van fosfor deur akwakultuur praktyke.In die geval van klein reservoirs, waar die drakrag duidelik oorskry is, was die gevolge, soos veroorsaak deur akwakultuur ernstig, en die assimilasie van afval deur die sisteem nie moontlik nie. In die uiterste gevalle het akwakultuurafval 60 % - 90 % van alle inkomende fosfor gelewer - twee tot nege maal die fosfor wat deur riviere en afloopwater ingevloei het. Wat die sediment aanbetref, kon slegs indirekte gevolgtrekkings gevorm word. Akwakultuurproduksie het hipolimnetiese anoksia laat toeneem en die laasgenoemde verskynsel, het die potensiaal aangedui vir die toename in die vrystelling van fosfor vanaf die sediment. Dit dui daarop dat akwakultuur nie alleen die fosforkonsentrasie in die oppervlaktwater laat toeneem nie, maar sou ook die interne belading laat kon toeneem. Die sedimentasie tempo het toegeneem met die teenwoordigkeid hokkultuur en het ‘n hipotetiese area van ongeveer 0.05 tot 1.00 ha, afhangende van reservoir hidrologie, beïnvloed. Die samestelling van die sediment het toegeneem in teme van die organiese komponente wat die sedimentasie prosesse kon beïnvloed. Dit kan gepostuleer word dat die toename in sedimentasie van akwakultuurafvalprodukte tesame met verlengde anoksiese toestande, ‘n invloed op die makrosoöbentiese organismes het. Hidrologiese en nutriënt massa uitruiling van die reservoirs het aangetoon dat geen jaarlikse toename in fosfor verkry kan word met lae nutrient toelae (kwaliteit van invloeiende water) of met goeie waterverplasing nie, en soms met die ekstraksie van hipoliminetiese water gedurende die stagnasie periode (somer). Die Beveridge model het soortgelyke resultate getoon tot die massabalans benadering en kan daarom dien as ‘n meer aanvaarbare model om gepaste beladingstempo vas te stel. Kleiner mensgemaakte reservoirs in die Wes-Kaap is onderhewig aan ‘n eutrofikasie proses wat die spoed van natuurlike eutrofikasie (trofies verwys na ‘n hoogs eutrofiese of hipertrofiese toestand ongeveer 10 tot 20 jaar na reservoir konstruksie) oorskry. Literatuur (nie ondersoek in die huidige studie) dui aan dat hierdie versnelde eutrofikasie proses meegebring word deur landbouaktiwiteite in die opvangsarea van die reservoirs. Resultate van die huidige studie het getoon dat forelproduksie in hokstelsels, die konsentrasie van total fosfor wat reeds beskikbaar was, verdubbel het. Die toename in fosforkonsentrasie het binne 1 tot 2 jaar na die aanvang van forelproduksie in die betrokke reservoirs, plaasgevind. Daar is egter uitsonderings waar forelproduksie in hokstelsels moontlik was, sonder die gepaardgaande afname in die waterkwaliteit. Die belangrikste stap vir suksesvolle en volhoubare forelproduksie is deeglike perseelseleksie. Daar is geen impak van akwakultuur waargeneem by vier persele (15 %) wat die eienskappe van goeie waterverplasing (>3 keer per jaar) en ‘n minimum oppervlakarea van 5 ha gehad het nie. Bykomend, sluit kriteria wat die risiko van algbesmetting laat afneem, ‘n minimum waterdiepte van 6 tot 7 m in ‘n reservoir by die laagste punt in (om te verhoed dat intermediêre vermenging plaasvind gedurende die stagnasieperiode) sowel as koue hipolimnetiese toestande (<17 °C) om sianobakterieë sist remobilisasie te minimaliseer. Verdere verbetering van die VOH (voeromsettingsverhouding onder voedingsbestuur) en voerkwaliteit is na perseeleleksie, die volgende komponente wat kan aandui of ‘n perseel gebruik kan word vir hokkultuur sonder enige impak. Vordering met voerkwaliteit en voedingsbestuur kan die aantal volhoubare persele laat toeneem en daardeur meer effektiewe watergebruik teweeg bring, sonder die verwante waterkwaliteit verswakking.

Thesis (MSc)--Stellenbosch University, 2011. / ENGLISH ABSTRACT: The Exxaro Namakwa Sands heavy mineral sands mine at Brand-se-Baai, on the west coast of South Africa, is an important source of income, development and job-creation in the region. However, this comes at a great environmental cost, as strip mining causes large scale destruction of ecosystems through the complete removal of vegetation and topsoil. This is particularly problematic in an environment, such as Namaqualand, where the arid and windy climate, as well as saline and nutrient-poor soils, hamper rehabilitation. These environmental constraints create the need to develop a site-specific rehabilitation program. At Namakwa Sands the objective of rehabilitation is to “rehabilitate and re-vegetate disturbed areas and establish a self-sustaining Strandveld vegetation cover in order to control dust generation, control wind and water erosion, as well as restore land capability. In general, vegetation will be rehabilitated to a minimum grazing standard capable of supporting small stock (sheep) grazing.” In order to achieve this Namakwa Sands conducted rehabilitation experiments with topsoil replacement, seeding of indigenous species and translocation of mature plants. Monitoring is an important part of the rehabilitation process as it allows rehabilitation practitioners to evaluate success and to adapt their management strategies and rehabilitation methods, as well as to evaluate and, if necessary, change their rehabilitation objectives. This study forms part of the monitoring process at Namakwa Sands. It assesses the success of sites that were experimentally rehabilitated in 2001 and a site that was rehabilitated in 2008, using current practice, in order to identify possible management requirements on rehabilitated sites as well as improvements on rehabilitation objectives, methods and monitoring. This study also tests the Landscape Function Analysis (LFA) as rehabilitation monitoring tool by correlating LFA indices with traditional measurements of biophysical variables or their surrogates. Results showed that experimental sites were not successful in returning vegetation cover and plant species richness to the required levels, but did achieve the grazing capacity objective. These sites will need adaptive management to achieve the vegetation cover and plant species richness objectives. The recently rehabilitated site achieved the three-year vegetation cover and plant species richness objectives, as well as the grazing capacity objective, within two years after rehabilitation. Namakwa Sands should therefore continue using the current rehabilitation method. However, rehabilitation should be done in multiple stages in future to decrease the mortality of nursery cuttings and to facilitate the return of late successional species to rehabilitated sites. The sustainability of small stock farming on rangeland with the grazing capacity that is identified as the minimum objective is questionable and this merits further investigation. LFA can be a useful tool to monitor nutrient cycling and soil stability at Namakwa Sands, provided that enough replicates are used. However, LFA cannot be used as is to assess water infiltration at Namakwa Sands, due to assumptions in the calculation of this index that do not hold for the Namaqualand environment. Landscape functioning should be monitored annually to complement vegetation surveys. / AFRIKAANSE OPSOMMING: Die Exxaro Namakwa Sands swaarminerale-sandmyn by Brand-se-Baai, aan die weskus van Suid-Afrika, is ‘n belangrike bron van inkomste, ontwikkeling en werkskepping in die streek. Daar is egter negatiewe omgewingsimpakte aan verbonde, aangesien die strookmyntegniek grootskaalse vernietiging van ekosisteme veroorsaak deur die algehele verwydering van die plantegroei en bogrond. Dit is veral problematies in ‘n omgewing, soos Namakwaland, waar die droë en winderige klimaat, asook die souterige en voedingstof-arme grond, rehabilitasie belemmer. Hierdie beperkings wat deur die omgewing veroorsaak word skep die behoefte om ‘n rehabilitasieprogram te ontwikkel wat spesifiek is tot die terrein. Die doel van rehabilitasie by Namakwa Sands is om te rehabiliteer en herplant op versteurde gebiede en om selfonderhoudende Strandveld plantbedekking te vestig om sodoende stofgenerering te beheer, om wind- en watererosie te beheer, en om grondgebruik-vermoë te herstel. In die algemeen sal plantbedekking gerehabiliteer word tot ‘n minimum weidingskapasiteit wat kleinveeweiding (skaapweiding) kan onderhou. Om dit te bereik het Namakwa Sands rehabilitasie-eksperimente uitgevoer met terugplasing van bogrond, saai van inheemse spesies en oorplanting van volwasse inheemse plante. Monitering is ‘n belangrike deel van die rehabilitasieproses, aangesien dit rehabilitasie-praktisyns in staat stel om sukses te evalueer en om bestuurstrategieë en rehabilitasiemetodes aan te pas, sowel as om rehabilitasiedoelwitte te evalueer en, indien nodig, aan te pas. Hierdie studie vorm deel van die moniteringsproses by Namakwa Sands. Dit assesseer die sukses op persele wat eksperimenteel gerehabiliteer is in 2001 en ‘n perseel wat in 2008 gerehabiliteer is, volgens die huidige praktyk, om moontlike bestuursbehoeftes op gerehabiliteerde persele en verbeteringe aan rehabilitasiedoelwitte, -metodes en –monitering te identifiseer. Hierdie studie toets ook die geskiktheid van die Landscape Function Analysis (LFA) as ‘n rehabilitasie-moniteringsinstrument deur LFA-indekse met tradisionele metings van biofisiese veranderlikes of hul surrogate te korreleer. Resultate dui daarop dat eksperimentele persele nie suksesvol was om plantbedekking en plantspesies-rykdom tot die vereiste vlakke te herstel nie, maar wel die weidingskapasiteit-doelwit bereik het. Hierdie persele benodig aanpassingsbestuur om plantbedekking- en plantspesiesrykdom-doelwitte te bereik. Die perseel wat onlangs gerehabiliteer is, het binne twee jaar na rehabilitasie die drie-jaar plantbedekking- en plantspesiesrykdom-doelwitte, sowel as die weidingskapasiteitdoelwit bereik. Daarom moet Namakwa Sands voortgaan om die huidige rehabilitasiemetode te gebruik. Rehabilitasie moet egter in die toekoms in veelvoudige stadiums gedoen word om die mortaliteit van kwekery-steggies te verminder en om die terugkeer van laatsuksessionele spesies na gerehabiliteerde persele te fasiliteer. Die volhoubaarheid van kleinveeboerdery op weiveld met die minimum vereiste weidingskapasiteit word betwyfel en vereis verdere ondersoek. LFA kan ‘n bruikbare instrument wees om siklering van voedingstowwe en grondstabiliteit te monitor by Namakwa Sands indien genoeg repliserings gebruik word. LFA kan egter nie in die huidige vorm gebruik word om waterinfiltrasie by Namakwa Sands te assesseer nie, aangesien daar aannames in die berekening van die indeks is wat nie juis is in die Namakwaland omgewing nie. Landskapfunksionering behoort jaarliks gemoniteer te word om plantopnames aan te vul.

Thesis (MScConsEcol) -- Stellenbosch University, 2003. / ENGLISH ABSTRACT: This thesis describes the findings of a survey to investigate the attitudes and behaviour of private landholders towards the conservation of Overberg Coastal Renosterveld, a highly endangered grassy-shrubland of the Cape Floral Region (CFR) lowlands, now 94% transformed. Personal interviews were conducted with 36 farmers in the Overberg region of the Western Cape, by administering a questionnaire. The following aspects were quantitatively described using the Statistical Package for Social Scientists (SPSS): Renosterveld management and utilisation behaviour, perceived value of Overberg Coastal Renosterveld; landholder knowledge of its conservation importance and willingness to conserve it. Attitudes towards incentives for conservation; conservancies; and the provincial conservation authorities were also investigated. Willingness to conserve was explored further using SPSS cross-tabulation analyses. It was found that farmers who were more willing to conserve were younger, did not necessarily have a better education, and owned larger farms (> 500 ha) with a greater amount of remnant renosterveld (> 300 ha) than farmers less willing to conserve. Attitudes towards Overberg Coastal Renosterveld were largely negative, due to certain associated plants and animals which are problems for farmers, and because it is not economically advantageous to retain it. However, provision of incentives (of which assistance with fencing and land management was most attractive) and increased extension support will provide practical positive inducements for conservation. There is also a need for more landholder education regarding the scarcity of coastal renosterveld to prevent transformation of remaining fragments. The second component of the study aimed to construct a user-friendly index to measure the conservation attitudes of landholders towards conserving Overberg Coastal Renosterveld. An interative item analysis was executed on the data, using Spearman Rank Order correlations and Cronbach's Alpha. Results yielded an index with two dimensions and a Cronbach Alpha co-efficient of 0.67. The dimensions or components of conservation attitude towards renosterveld included: (i) a landholders' perception of the perceived benefit of Overberg Coastal Renosterveld; and (ii) their willingness to conserve it. The mean conservation attitude score was 0.6 (± 0.03), indicating that landholder attitudes were generally sympathetic towards renosterveld conservation. This suggests that while many landholders do not place a high value on Overberg Coastal Renosterveld due to its low agricultural value, their willingness to conserve it is in some cases higher than expected, which possibly explains the above-average mean conservation attitude score. An analysis of variance showed that the following variables were significantly, positively correlated with conservation attitude: (i) area of Overberg Coastal Renosterveld; (ii) landholder environmental group membership status; (iii) presence of ecotourism activities on the property; and (iv) how long the property had been in the owner's family. The intended application of the index is that index scores (amenable for use in a Geographical Information System database) can assist conservation practitioners in deciding where resources should be allocated, on the assumption that high-scoring individuals are more likely to want to take part in conservation initiatives. / AFRIKAANSE OPSOMMING: Hierdie proefskrif beskryf die bevindings van 'n opname waarin die houdings en gedrag van private grondeienaars ten opsigte van Overberg Kustelike Renosterveld ondersoek word. Altesaam 94% van die oorspronklike renosterveld, 'n hoogs bedreigde, grasagtige struikveld in die laaglande van die Kaapse Planteryk (KPR) , is reeds getransformeer en verander. Persoonlike onderhoude is met 36 boere in die Overbergstreek van die Wes-Kaap gevoer en 'n vraelys is aan hulle gestel. Die "Statistical Package for Social Scientists" (SPSS) is gebruik om die volgende aspekte kwantitatief te beskryf: die bestuur en benutting van Overberg Kustelike Renosterveld; die waarde wat toegeskryf word aan hierdie renosterveld; asook grondeienaars se kennis oor die bewaringsbelang van Overberg Kustelike Renosterveld en hul bereidwilligheid om dit te bewaar. Houdings jeens aansporingsmiddele om bewaring te bevorder, bewareas en die provinsiale bewaringsowerhede is ook ondersoek. Private grondeienaars se bereidwilligheid om betrokke te raak by bewaringswerk is verder ondersoek deur gebruik te maak van die SPSS-kruistabelleringsanalise. In vergelyking met boere wat nie bewaring ondersteun nie, is gevind dat die meer bewaringsgesinde boere jonger is, nie noodwendig 'n beter opvoeding het nie, en groter plase (>500 ha) besit waarop meer brokkies oorblywende Overberg Kustelike Renosterveld (>300 ha) gevind word. Grondeienaars se houding jeens Overberg Kustelike Renosterveld was grootliks negatief. Dit is as gevolg van sekere verwante plante en diere wat deur boere as probleme ervaar word, en omdat dit nie ekonomies lewensvatbaar is om renosterveld te behou nie. Die beskikbaarheidstelling van aansporingsmiddele (waarvan hulpverlening met omheinings en grondbestuur die aantreklikste was), en 'n toename in landbou-voorligting is egter praktiese en positiewe dryfvere wat bewaring kan bevorder. Daar is ook 'n behoefte vir meer opvoeding van grondeienaars oor aspekte soos die skaarsheid van Overberg Kustelike Renosterveld, sodat die verdere transformasie van oorblywende fragmente voorkom kan word. Die tweede gedeelte van die studie was daarop gemik om 'n gebruikersvriendelike indeks te ontwikkel waarmee die bewaringsgesindheid van grondeienaars teenoor die bewaring van renosterveld gemeet kan word. Deur gebruik te maak van Spearman se rangorde-korrelasies en Cronbach se Alpha is die data aan 'n herhalende item-analise onderwerp. Die resultate het 'n indeks met twee dimensies en 'n Cronbach Alpha-koëffisiënt van 0.67 gelewer. Die dimensies of komponente van bewaringsgesindheid jeens renosterveld het onder meer ingesluit: i) 'n grondeienaar se opvatting oor die moontlike voordeel van Overberg Kustelike Renosterveld en ii) sy/haar bereidwilligheid om dit bewaar. Die gemiddelde telling vir bewaringsgesindheid was 0.6 (± 0.03), wat in die algemeen 'n simpatieke houding jeens die bewaring van Overberg Kustelike Renosterveld onder grondeienaars aandui. Hiervolgens blyk dit dat alhoewel heelwat grondeienaars weens die lae landbouwaarde van renosterveld nie 'n hoë waarde op hierdie plantegroeitipe plaas nie, hulle bereidwilligheid om dit te bewaar soms verwagtinge oortref. Dit is dalk 'n moontlike verklaring vir die bogemiddelde gemiddelde bewaringsgesindheidtelling. 'n Variansie-analise het 'n noemenswaardige, positiewe korrelasie met bewaringsgesindheid ten opsigte van die volgende veranderlikes getoon: i) area van Overberg Kustelike Renosterveld; ii) die grondeienaar se moontlike lidmaatskap van 'n bewaringsgroep; iii) die teenwoordigheid van ekotoerisme-bedrywighede op die eiendom; en iv) die tydperk wat die eiendom al in besit van die grondeienaar se familie is. Die voorgestelde gebruik van die indeks is toepaslik omdat die indeks-tellings (wat vir gebruik in 'n Geografiese Inligtingstelsel databasis aangepas kan word) bewaringslui kan help met besluite random die toekenning van hulpbronne. Die aanname is dat individuele grondeienaars wie hoë tellings aangeteken het, met alle waarskynlikheid meer geredelik aan bewaringsinisiatiewe sal wil deelneem.

Thesis (MSc)--University of Stellenbosch, 2005. / ENGLISH ABSTRACT: The primary aim of this study was to assess and compare the ecological integrity of the Lourens, Palmiet and Hout Bay Rivers (South Western Cape, South Africa) by examining the macroinvertebrate community structure at a series of representative sampling sites along the course of each river, using the South African Scoring System - Version 5 (SASS-5) rapid bioassessment method. Secondary aims included an examination of the effects of seasonal variability, biotope availability and site-specific environmental variables on the macroinvertebrate community structure at sampling sites, as well as the preliminary testing of the Integrated Habitat Assessment System (IHAS) for aquatic macroinvertebrates. According to results obtained, the ecological integrity of sampling sites in the Mountain Stream Zone of the three rivers was consistently good. The Hout Bay River in the upper portions of the Orange Kloof Reserve was particularly near-pristine, with this area having been identified in this study as a potential biodiversity ‘hot-spot’ for aquatic macroinvertebrates. Downstream of the Mountain Stream Zone, there was a significant deterioration in the ecological integrity of all three rivers due to a number of probable causes. Results based on recorded SASS Scores and Average Score per Taxon (ASPT) values, using ‘biological bands’ generated from reference sites in the South Western Cape, were generally similar to and supported by the corresponding multivariate analyses undertaken. From the results of the various analyses undertaken in this investigation and some of the problems encountered in interpreting the data, a number of recommendations are made regarding future bioassessment studies based on the SASS within the national River Health Programme (RHP). To test the IHAS, secondary data were obtained from reference sites in the Mpumalanga and Western Cape Provinces of South Africa. Assuming that SASS Scores at reference sites are the highest scores attainable, one would expect to find a positive relationship between SASS Scores and IHAS scores at reference sites. The assumption in this investigation was that this relationship should be linear. Non-parametric correlation analyses were undertaken between SASS-4/5 Scores and IHAS scores, using Kendall’s Rank-correlation Coefficient (τ), with separate analyses undertaken for different geomorphological zones and biotope groups. Correlations between SASS Scores and IHAS scores were generally weak (τ-values mostly < 0.3) and unsatisfactory, with no significant correlations (p < 0.05) for two-thirds of the data sets analysed and a wide degree of scatter generally observed amongst data points in respective scatter plots. The performance of the IHAS varied between geomorphological zones and biotope groups, with the Foothill: Gravel-bed Zone in Mpumalanga showing the best results, particularly when the stones-in-current biotope group was analysed separately. Further testing of the IHAS is required to confirm its relative performance in different bioregions/ecoregions, geomorphological zones and biotope groups, which should be undertaken as a priority research area within the RHP. Unsuccessful attempts to test the IHAS by means of multiple regression analyses were undertaken, suggesting that such techniques should be avoided in further testing of the IHAS. / AFRIKAANSE OPSOMMING: Die hoofdoel van hierdie studie was om die ekologiese toestand van die Lourens-, Palmiet- en Houtbaairiviere (Suidwes Kaap, Suid Afrika) te bepaal en te vergelyk deur die bestudering van die makroinvertebraatgemeenskapstruktuur by verteenwoordigende monsterpunte langs die riviere, met gebruik van die “South African Scoring System” – Weergawe 5 (SASS-5) snelle biologiese bepalingsmetode. Sekondêre doelwitte het die bepaling van die gevolge van seisoenele veranderlikheid, biotoop beskikbaarheid en ligging-bepaalde omgewingsveranderlikes op die makroinvertebraatgemeenskapstruktuur by monsterpunte ingesluit, asook die inleidende toetsing van die “Integrated Habitat Assessment System” (IHAS) vir watermakroinvertebrate. Volgens die resultate verkry, was die ekologiese toestand van monsterpunte in die Bergstroomsone van die drie riviere konsekwent goed. Die Houtbaairivier in die boonste gedeelte van die Oranjekloofreservaat was veral feitlik onversteurd en hierdie streek is in die studie as ‘n potensiaal biodiversiteit “hot-spot” vir watermakroinvertebrate geidentifiseer. Stroomafwaarts van die Bergstroomsone was daar ‘n beduidende verswakking in die ekologiese toestand van al drie riviere, as gevolg van ‘n aantal moontlike oorsake. Resultate gebaseer op bepaalde “SASS Scores” en ‘Gemiddelde Waarde per Takson’ (“Average Score per Taxon” - ASPT) waardes, met gebruik van ‘biologiese bande’ wat van verwysingsmonsterpunte in die Suidwes Kaap afgelei is, was oor die algemeen soortgelyk aan en gestaaf deur die ooreenstemmende multiveranderlikke (“multivariate”) statistiese analises wat gedoen is. Uit die resultate van die verskeie analises wat in hierdie ondersoek gedoen is en sommige van die probleme wat in die dataverklaring gevind is, is ‘n aantal aanbevelings gemaak met betrekking tot toekomende biologiese bepalingstudies vir die nasionale Riviergesondheidsprogram (“River Health Programme” - RHP) wat op die SASS gebaseer is. Om die IHAS te toets is sekondêre data van verwysingsmonsterpunte in die Mpumalanga en Wes Kaap Provinsies van Suid Afrika verkry. As aangeneem word dat die “SASS Scores” by verwysingsmonsterpunte die hoogste moontlike tellings is wat bereik kan word, sou ‘n positiewe verwantskap tussen “SASS Scores” en IHAS tellings by verwysingsmonsterpunte verwag word. Die veronderstelling in hierdie studie was dat dié verwantskap lineêr moet wees. Nie-parametriese korrelasieanalise tussen “SASS-4/5 Scores” en IHAS tellings is gemaak, deur gebruik van Kendall se Rangkorrelasiekoëffisiënt (τ), met afsonderlike analises vir verskillende geomorfologiese sones en biotoopgroepe verrig. Korrelasies tussen “SASS Scores” en IHAS tellings was algemeen swak (τ-waardes < 0.3) en onbevredigend, met geen beduidende korrelasies (“p” < 0.05) vir twee-derdes van die datastelle wat geanaliseer is nie en ‘n wye verspreiding tussen datapunte in die onderskeie “scatter plots” wat waargeneem is. Die funksionering van die IHAS was verskillend tussen geomorfologiese sones en biotoopgroepe. Die beste resultate is vir die Voorheuwel: Gruisbeddingsone in Mpumalanga verkry, veral indien die klippe-in-stroom biotoopgroep afsonderlik geanaliseer is. Verdere toetsing van die IHAS is nodig om die relatiewe funksionering in verskillende biostreke/”ecoregions”, geomorfologiese sones en biotoopgroepe te bevestig en dit behoort voorangs te geniet binne die RHP. Pogings om die IHAS deur middel van veelvoudige regressie analise te toets het misluk, wat aandui dat sulke tegnieke vermy moet word in verder toetsing van die IHAS.

Thesis (MSc)--University of Stellenbosch, 2005. / ENGLISH ABSTRACT:This study concentrates on the Bontebok (Damaliscus pygargus pygargus Pallas 1766) population of the Bontebok National Park. Using a combination of literature research, field observations, satellite imagery, Geographic Information System mapping and dung analysis, their habitat preferences within the Park are ascertained and compared to different behavioural and environmental factors. Considering vegetation composition and height, grass cover, burning history, general topography, water availability, actual diet quality (from faecal analysis) and population distribution, the best predictor of spatial pattern are investigated with GIS Dempster-Shafer models. The historical density distribution of the animals in the Park can be used to predict sustainable stocking rates. As far as possible, the results for the Bontebok are compared to those of the other larger herbivores in Bontebok National Park i.e. Red Hartebeest, Cape Mountain Zebra and Grey Rhebuck. These results should be useful for Wildlife Managers to know how to manage veld for Bontebok and at what stocking rates this can be done sustainably. / AFRIKAANSE OPSOMMING: Hierdie studie fokus op die Bontebok (Damaliscus pygargus pygargus, Pallas 1766) bevolking van die Bontebok Nasionale Park. Deur ‘n kombinasie van literatuurstudie, veldobservasies, satlietbeelde, Geografiese Inligtingstelsel (GIS) kartering en mis-analise te gebruik, word Bontebok habitatvoorkeure in die Park vasgestel en met verskillende gedrags- en omgewingsfaktore vergelyk. Deur te kyk na plantegroei-samestelling en hoogte, grasbedekking, brandgeskiedenis, algemene topografie, water-beskikbaarheid, diëet kwaliteit en die Bontebokke se waargenome ruimtelike verspreiding, word die faktor wat die verspreidingspatroon die beste voorspel, m.b.v. GIS Dempster-Shafer modelle ondersoek. Die historiese digtheidsverspreiding van diere in die Park word gebruik om volhoubare veldladings te voorspel. Sover moontlik word die resultate vir Bontebok vergelyk.met die ander groter herbivore in die Bontebok Nasionale Park, te wete Rooi Hartebees, Kaapse Bergsebra en Vaalribbok. Hierdie resultate behoort vir Natuurlewe-bestuurders tot nut te wees as riglyne vir veldbestuur en volhoubare veldladings van Bontebok.

Thesis (MSc)--University of Stellenbosch, 2006. / ENGLISH ABSTRACT: Understanding the responses of species to climate change is a scientific problem that requires urgent attention, especially under current conditions of global climate change. The large and rapid rates of climate change reported for sub-Antarctic Marion Island makes the island highly suitable for studying the biotic consequences of climate change. Furthermore, the extreme environments on the island result in a close coupling of the biotic (e.g. population dynamics) and abiotic (e.g. climate) factors. Therefore, examining the response of the dominant and keystone plant species on the island, Azorella selago Hook. (Apiaceae), to climate-associated environmental change (e.g. temperature) may provide insight into how A. selago and the associated species communities will be affected by climate change. This study described the variability in microclimate temperatures associated with A. selago across altitudinal gradient and between the eastern and western sides of Marion Island. Microclimate temperatures were also compared to the island’s Meteorological data to determine variation between temperatures experienced by A. selago cushion-plants in the field and those recorded at the island’s Meteorological Station. Temperature variation inside and outside A. selago cushions was also examined. Azorella selago cushions were found to have a buffering effect on temperature, such that species occurring epiphytically on A. selago experience more moderate temperatures than the surrounding environment. However, A. selago were found to experience more extreme temperatures than temperatures recorded at the Meteorological Station. Therefore, A. selago may possibly experience greater environmental warming than recorded by the Meteorological Station. While temperatures decline with altitude, temperature conditions on the western side of the island were more temperate than the eastern side. This presents the first record of temperature conditions on the western side of the island. This study also quantified fine-scale (e.g. within-site) and broad-scale (e.g. island-wide) variability patterns of A. selago (morphology, phenology, and epiphyte load) across Marion Island. Altitudinal gradient and climatic exposure at different sides of the island were used to understand the likely effects of climate associated environmental change on this dominant component of the fellfield habitat. Site-specific processes were found to determine the spatial structure of A. selago characteristics at fine-scales. However, broad-scale observations established strong responses of A. selago characteristics to altitudinal gradients and different sides of the island. Azorella selago morphological features (e.g. plant size and leaf size) were found to be more responsive to differences between the eastern and western sides of the island than to altitudinal gradient. Azorella selago micro-morphological features (e.g. leaf trichomes and stomatal densities) were also found to be more responsive to climatic exposure at different sides of the island than to altitudinal gradient. However, differences in A. selago epiphyte density (e.g. Agrostis magellanica) and phenology resembled microclimate temperatures in that they were more responsive to altitudinal gradient than to side of the island differences. From these results it can therefore be predicted that the A. selago of Marion Island is likely to be morphologically fairly resilient to moderate climatic shifts, although at lower altitudes and on the eastern side of the island, it may be outcompeted by the epiphytic grass, Agrostis magellanica. The results also suggest that the warming climate of Marion Island may result in an early occurrence of phenological processes particularly at lower altitudes and the eastern side. Azorella selago at lower altitudes and on the eastern side of Marion Island are therefore expected to largely show more symptoms of climate change (e.g. warming) on this species. Azorella selago is also predicted to move up altitudinal gradients in response to warming. / AFRIKAANSE OPSOMMING: ’n Begrip van hoe spesies reageer op klimaatsverandering is ’n wetenskaplike vraag wat onmiddellike aandag benodig, veral onder huidige globale klimaatsverandering. Die groot en snelle tempo waarteen klimaatsverandering waargeneem word op sub-Antarktiese Marion Eiland, maak die eiland hoogs geskik om die biotiese gevolge van klimaatsverandering te bestudeer. Verder veroorsaak die uiterste omgewing van die eiland tot ’n nabye koppeling tussen die biotiese (bv. populasie dinamika) en abiotiese (bv. klimaat) faktore. Dus, deur die reaksies van ’n dominante- en sleutel-spesie op die eiland, Azorella selago Hook. (Apiaceae), op klimaat-geassosieerde omgewings verandering (bv. temperatuur) te bestudeer, mag insig verskaf hoe A. selago en geassosieerde spesie gemeenskappe geaffekteer sal word deur klimaatsverandering. Hierdie studie beskryf die wispelturigheid in mikroklimaat temperature geassosieer met A. selago oor ’n hoogte gradiënt asook tussen die oostelike en westelik dele van Marion Eiland. Mikroklimaat temperature was ook vergelyk met die eiland se Meteorologiese data met die doel om die mate van variasie tussen temperature verduur deur A. selago kussing-plante in die natuurlike omgewing met die van die eiland se Meteorologiese stasie te vergelyk. Temperatuur variasie binne en buite A. selago kussing-plante is ook vasgestel. Dit was gevind dat Azorella selago kussing-plante die temperatuur buffer, met die gevolg dat spesies wat epifities op A. selago voorkom, meer gematigde temperature ondervind as die onmiddellike omgewing om die plant. Daar is egter gevind dat A. selago meer uiterste temperature ondervind as temperature gemeet by die Meteorologiese stasie. Dus mag A. selago groter omgewings verwarming ervaar as wat temperature gemeet by die Meteorologiese stasie dui. Terwyl temperatuur afneem met ’n toename in hoogte, was temperatuur aan die westekant van die eiland mere gematig as die oostekant. Dit verskaf die eerste rekord van temperatuur toestande aan die westekant van die eiland. Hierdie studie bepaal ook die fyn-skaal (e.g. binne-terrein) en groot-skaal (e.g. oor die eiland) variasie patrone van A. selago (morfologie, fenologie, en epifiet lading) oor Marion Eiland. Die hoogte gradiënt en klimaat blootstelling aan verskillende kante van die eiland is gebruik om die waarskynlike effekte van klimaats-geassosieerde omgewings verandering op die dominante deel van die felfield habitat te verstaan. Daar is gevind dat terrein spesifieke prosesse die ruimtelike struktuur van A. selago se klein-skaal eienskappe bepaal. Groot-skaalse waarnemings dui egter ’n sterk antwoord van A. selago eienskappe op die hoogte gradiënt en aan verskillende kante van die eiland. Azorella selago morfologiese eienskappe (e.g. plant- en blaar grootte) is gevind om meer te reageer op verskille tussen oostelike en westelike kante van die eiland as op die hoogte gradiënt. Azorella selago mikromorfologiese eienskappe (e.g. blaar trigome en stomata digtheid) is ook gevind om meer te reageer op omgewings blootstelling tussen verskillende kante van die eiland as op die hoogte gradiënt. Verskille in A. selago epifiet digtheid (e.g. Agrostis magellanica) en fenologie het egter mikroklimaat temperature gevolg, in dat beide meer gereageer het op die hoogte gradiënt as eiland-kant verskille. Hierdie resultate voorspel dus dat dit waarskynlik is dat A. selago van Marion Eiland morfologies redelik terugspringend sal wees ten opsigte van matige klimaatsverandering, al mag dit uitgekompeteer word deur die epifitiese gras, Agrostis magellanica by lae hoogtes en aan die oostekant van die eiland. Hierdie resultate dui ook dat verwarming van Marion Eiland se klimaat ’n vervroeging van fenologiese prosesse mag hê, veral by lae hoogtes en aan die oostekant van die eiland. Dus word dit verwag dat Azorella selago by lae hoogtes en aan die oostekant van Marion Eiland om meer simptome van klimaatsverandering (e.g. verwarming) te dui. Dit word ook voorspel dat Azorella selago opwaarts teen die hoogte gradiënt sal beweeg in reaksie tot verwarming.