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  • About
  • The Global ETD Search service is a free service for researchers to find electronic theses and dissertations. This service is provided by the Networked Digital Library of Theses and Dissertations.
    Our metadata is collected from universities around the world. If you manage a university/consortium/country archive and want to be added, details can be found on the NDLTD website.
1

AVAILABLE ENERGY AND SPECIES DIVERSITY: THEORY AND EXPERIMENTS WITH BEES (COMMUNITIES, FLOWERS, FORAGING).

WRIGHT, DAVID HAMILTON. January 1984 (has links)
A general biogeographic theory of insular species diversity, species-energy theory, is produced by replacing area in species-area models with a measure of available energy. Islands with more available energy support larger populations, which have lower extinction rates. Given similar immigration rates, islands with greater available energy are therefore expected to support greater equilibrium numbers of species. Assuming that total population size is proportional to energy supply, and that species-abundance distributions are lognormal and of similar form, the species-energy relationship is approximated by S = kEᶻ. Species-energy theory explains 70-80% of the variation in species number of angiosperms and of birds on such widely varying islands as Greenland and Jamaica. The effects of energy on the structure of a subalpine bee community in Colorado were investigated. As available nector declined, during mornings and over the season, foraging profitability for Bombus appositus (Hymenoptera: Apidae) decreased. This change was manifested by increased foraging trip durations: nector loads did not change. Total colony profits increased as colonies grew over the season, but profit relative to colony size declined, due to reduced profitability of individual foraging trips. These results support the hypothesis of resource limitation in this species. Assemblages of bees foraging on patches of flowers showed effects of energy availability on species composition and dynamics. Bees foraging in enriched patches had lower departure rates than bees in control patches, and, consequently, increased equilibrium numbers of individuals and species present per patch. Both behavioral and mechanical factors influenced departure rates. A species-specific arrival-departure rate model satisfactorily described the foraging assemblages and their response to enrichment. Experiments performed on 2 species of flowers with different corolla tube lengths demonstrated that bee species respond differently to resources of unequal availability, necessitating a species-level approach. Analogies with island systems are discussed. Energy is important to communities in general and bees in particular on a variety of scales. By implication, human resource diversion from natural ecosystems may have profound impacts on global diversity and extinction.
2

Ecological genetics of Poa alpina

MacEwan, A. January 1986 (has links)
No description available.
3

Maintenance of phenotypic variation in the butterfly, Maniola jurtina

Goulson, David January 1991 (has links)
No description available.
4

Ecological genetics of two polymorphic enzymes in three species of Megalorchestia (Amphipoda:Talitridae)

McDonald, John H. 16 December 1983 (has links)
Graduation date: 1984
5

The genetics of cultivar and host specificity of Phytophthora sojae and P. vignae

May, K. Unknown Date (has links)
No description available.
6

Ecological genetics of Trifolium repens

Pusey, J. G. January 1965 (has links)
The theory of Evolution by Natural Selection deals substantially with events now past, and with processes too slow for contemporary study. Experimental studies of evolution are possible, but are liable to the criticism that they deal with artificial, or at least extreme and atypical situations, involving rapid evolutionary change. Such criticism can be avoided to a degree by studying situations now in equilibrium. It is suggested that interesting results may be obtained by studying polymorphisms, and comparing the equilibria reached in populations of an organism in different environments. The chosen approach was to find a case of polymorphism convenient to study, followed by an attempt to interpret the pattern of its variation in terms of features of environments where a morph was common selecting in its favour, and vice versa. The material presented falls into three parts. 1) Material and Methods: Background and Preliminary Studies. The system chosen for study was the white V-shaped leaf markings of Trifolium repens. The species is widespread, abundant, and variable; the character is expressed on vegetative organs, and is therefore always accessible; and the known variation in this character had already been interpreted in terms of the action of a set of several alleles at one locus. The results of crosses indicate general agreement with previous reports. One heterozygous plant apparently largely failed to transmit one allele, V<sup>b</sup>, through the pollen. Segregations indicate that a new phenotype, 'smeared', is determined by a particular V<sup>h</sup> allele. Two more new phenotypes, 'shaded' and 'marginal' are described. The places where these and other rare phenotypes have been found are listed. New reports and the author's other experience are combined with previous reports in a new general account of leaf marking in the species, which also deals with at least three distinct classes of red markings. There appears to be more variation, implying more alleles, than previous accounts allow for, and reportedly distinct types, for example, V<sup>h</sup>, V<sup>l</sup> and V<sup>f</sup>, seem to be linked by a series of intermediates, among which clear dividing lines are hard or impossible to draw. Experimental studies relating to the techniques of population sampling are reported. Scarification by 10 to 20 minutes exposure to concentrated sulphuric acid was chosen to deal with hard seed. The effect of a restricted number of mother plants on the accuracy of a seed sample is discussed; predictions concerning the frequencies of different types in progeny from single heads grown separately were roughly consistent with observations. Possible differences between vegetative and seed samples are discussed. Morph frequencies in a sample of plants bearing inflorescences were not found to be significantly different from those of the whole vegetative sample. The consequences of this species' ability to reproduce vegetatively are discussed, and a brief clone-mapping project is described. The procedures used to obtain and score material, and to obtain information about its background are described. The amounts of material of different types from different sources is summarised in Table 6.2. Problems involved in estimating gene frequencies are discussed, and the derivation of the values used to represent the different morph frequencies is outlined; they are basically phenotypic frequency figures, gene frequency calculations being used only where it was found to be necessary. 2) Observations on Population Samples. Polymorphism in respect of white leaf marks was found to be present in all except 15 out of 624 samples. The commonest group of phenotypes are the 'simple V-marks', referred to as 'L'. The next commonest is the unmarked type, 'O', present in all except 72 out of 624 samples, with an overall frequency among the plants scored of 17 per cent. Study of the sample data reveals a deficiency of the double marked phenotypes expected to be showed by V<sup>by</sup>V<sup>l</sup> plants. This is explained as the result of a degree of dominance of V<sup>by</sup> in such combinations; the effect of such dominance is allowed for in the frequency figures representing the frequency of 'L' marks, and of double-marks containing two members of the 'L-series' of simple-V-producing alleles. The possibility of demonstrating interaction between the frequencies of different morphs is discussed. There are indications of lower frequencies of 'By', 'B', and 'F' marks when the unmarked phenotype is common. The possibilities for the main object of the work, discovering associations between morph frequencies and environmental factors are shown in Table 8.1, giving the sets of data presentable for particular methods of analysis. Data on 148 British samples, scored in the field by the author, were treated by Multiple Regression Analysis (using the KDF 9 Computer of the Oxford Computing Laboratory), to test for association of morph frequencies with geographical location, altitude, and soil pH. Significant increases in the proportion of unmarked plants are shown with greater distance north and higher altitude. Frequency data for all the classes of morphs described is shown in the form of maps, of the British Isles, Western Europe, and the whole of the species' natural range. These maps confirm the northwards increase in frequency of the unmarked form found in Britain. Maps for the rare morphs show various patterns, most of which seem to involve central regions of higher frequency north of the Mediterranean (France - Alps - Greece) with lower frequencies elsewhere. Regression analysis of a set of seed samples from Spain supports the conclusion from British vegetative samples of an increase in unmarked frequency with higher altitude. Examination of the information about the background of other samples suggests higher unmarked frequencies in pastures than on waste ground, meadows being intermediate. British data suggest an association with wet, and particularly with badly-drained sites, with trodden paths, and with dense vegetation. There are patterns of response to water regime elsewhere but (e.g. Polish data) they tend to suggest the opposite association, of high unmarked frequency with dry conditions. 3) Comparison with other species. New observations are reported on some other related species. Available information on marking in other species of Trifolium is reviewed, and it is pointed out that it is scattered, difficult of access (much unpublished), and sometimes ambiguous or contradictory. Summary of this data indicates the presence of white V-markings in nearly 30 species of Trifolium, and its absence in at least as many more. Marks are probably entirely absent in subgenus Chronosemium and perhaps in part of subgenus Trifolium. but seem to occur in most of the other subgenera for which there is information. When marks are present, it appears that they are nearly always variable. Only three species are definitely reported as always marked, and in two of these there is variation between different types of marking. This suggests that the factors producing or preserving polymorphism in T. repens act also in other marked species. The presence of red leaf marks of various kinds is reported in 21 species of Trifolium. Also in the tribe Trifolieae, both red and white V-markings are found in Parochetus communis. Material grown by the author showed great variability within each plant in leaf mark, but no clear differences between plants. Red leaf marks, variable, and in some cases approaching a V-shape, are also found in Medicago. Several lines of evidence indicate relationships between the white markings, and the various red marking systems in T. repens. It is suggested that the white and red V-markings have a common evolutionary origin. Some examples of leaf marking with analogous properties in genera unrelated to T. repens are also briefly reviewed. In discussion some possible challenges to the validity of the results claimed are discussed, and evidence is presented suggesting that the reported genetic clines in unmarked frequency are real. Selective factors affecting unmarked frequency are tentatively suggested to be temperature and water regime. The problem of relating these to markings on leaves is discussed, and also the possibility that the phenomena of leaf marking are by-products of unknown processes, and are of no intrinsic importance. It is suggested that the inter- actions between red and white markings support the hypothesis that leaf markings themselves are of selective importance; and some possibilities as to what form this selective importance might take are mentioned. Possibilities for further work indicated in the course of the studies presented here are discussed. They include studies on important problems of population dynamics, which affect the design of techniques for sampling for leaf markings, but in which also observations on leaf markings could be used as means to ends of wider significance. Ways in which the methods used in the present study could be improved in a repeated study are suggested; however, it is felt that the clines observed provide a starting point for experimental work, and that this might be more rewarding than further descriptive work.
7

Life history variation in Mimulus guttatus (Scrophulariaceae), the importance of ecological pressures in space and time

McCombie, Helen January 1995 (has links)
No description available.
8

The ecological genetics of flower colour variation in Cirsium palustre

Mogford, D. J. January 1972 (has links)
The thistle Cirsium palustre exists over most of lowland Britain as a predominantly purple flowered species. However the populations of seacliffs and mountains exhibit a flower colour polymorphism, occurring as homozygous white morphs, homozygous and heterozygous intermediate morphs, and homozygous and heterozygous purple morphs. The degree of polymorphism of the seacliff populations of the Gower Peninsula is correlated with population size, the larger populations being less polymorphic. This might be interpreted as indicating chance fixation of the white alleles. Such an explanation would gain support from the very small size of the more polymorphic populations and from the drastic fluctuations in population size which have been demonstrated as having occurred in these populations over a period of four consecutive seasons. However other explanations based on selective effects are possible. On seacliffs the frequency of the white morphs is inversely related to exposure, and decreases in population size are accompanied by differential survival of the purple morphs. On mountains the distribution of the polymorphism is markedly correlated with altitude. For the mountains of southern Mid Wales, populations below 1000 ft. are strongly monomorphic purple. Above 1000 ft. the degree of polymorphism increases abruptly, with white frequencies reaching over 80%. Morph frequencies among North Wales populations bear a similar relation to altitude but the increase in polymorphism occurs at about 1200 ft. In each case the general trend is that purple frequency declines with increase in white frequency, and that the frequency of intermediates shows a unimodal distribution with a quite precise peak. For both sets of populations this peak occurs at an altitude about 250 ft. higher than that at which the increase in white frequency occurs. It is possible that the occurrence of the polymorphism on seacliffs and mountains may be related to a limitation of cross pollination consequent upon the exposure of seacliffs and the combined climatic characteristics of mountains, which include increase in exposure, mist and rain and decrease in temperature. Evidence on the levels of outbreeding in these populations was inconclusive but evidence in other species suggested that pollination might indeed be limited in these conditions. An increase in homozygosity consequent upon inbreeding would promote the frequency of the white morphs. Moreover the white morphs were subject to preferential pollination and both this and certain forms of heterogeneity in morph distribution were likely to promote the frequency of inbreeding among white morphs. However in conditions of limited pollination the degree of general outbreeding of the white morphs will be increased by preferential pollination and this may be assumed to be a fitness advantage which may be of particular importance in the maintenance of the polymorphism. In addition it is likely that the presence of white morphs within a population may result in the attraction of higher numbers of pollinators or encourage foraging for longer periods in which case the polymorphism may be said to be adaptive in the sense of Fisher (1930). Other selective effects are also apparent. The occasional presence of highly polymorphic populations in valley bases and the regular occurrence of predominantly purple populations in mountain forests may both to some extent provide evidence for an effect of temperature other than upon pollination. Some evidence suggests that both exposure and moisture may also be of individual importance. Selection was apparent even by the arrangement of morph types within a population subject to no obvious environmental heterogeneity. It is possible that the polymorphism is maintained by a physiological heterozygous advantage and that this may be responsible for the maintenance of white and intermediate morphs in low frequency in the predominantly purple populations of inland lowland regions. The maintenance of the polymorphism imposed a significant selective mortality upon the species. This was indicated by the above instance of selection within a uniform community and also in several instances in which intrapopulational selection occurred between segments of population subject to differing exposure. However the growth in cultivation of seeds set in natural populations revealed that the complexity of the genetic system was sufficient to allow widely different morph frequencies to be maintained in different populations without the necessity of high selection in each generation.
9

The evolutionary ecology of tropane alkaloids /

Shonle, Irene Katherine. January 1999 (has links)
Thesis (Ph. D.)--University of Chicago, Dept. of Ecology and Evolution, June 1999. / Includes bibliographical references. Also available on the Internet.
10

Ecological genetics of floral longevity in Campanula rotundifolia, the Alpine Harebell /

Giblin, David Emmett, January 2001 (has links)
Thesis (Ph. D.)--University of Missouri-Columbia, 2001. / Typescript. Vita. Includes bibliographical references. Also available on the Internet.

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