A major challenge in savanna rangeland studies is estimating woody vegetation cover and densities over large areas where field based census alone is impractical. It is therefore crucial that the management and conservation oriented research in savannas identify data sources that provides quick, timely and economical means to obtain information on vegetation cover. Satellite remote sensing can provide such information. Remote sensing investigations, however, require establishing statistical relationships between field and remotely sensed data. Usually regression is the empirical method applied to field and remotely sensed data for the spatial estimation of woody vegetation variables. Geostatistical techniques, which take spatial autocorrelation of variables into consideration, have rarely been used for this purpose. We investigated the possibility of improving woody biomass predictions in tropical savannas using cokriging. Cokriging was used to evaluate the cross-correlated information between SPOT (Satellites Pour l’Observation de la Terre or Earth-observing Satellites)-derived vegetation variables and field sampled woody vegetation percentage canopy cover and density. The main focus was to estimate woody density and map the distribution of woody cover in an African savanna environment. In order to select the best SPOT-derived vegetation variable that best correlate with field sampled woody variables, several spectral vegetation and texture indices were evaluated. Next, variogram models were developed: one for woody canopy cover and density, one for the best SPOT-derived vegetation variable, and a crossvariogram between woody variables and best SPOT-derived data. These variograms were then used in cokriging to estimate woody density and map its spatial distribution. Results obtained indicate that through cokriging, the estimation accuracy can be improved compared to ordinary kriging and stepwise linear regression. Cokriging therefore provided a method to combine field and remotely sensed data to accurately estimate woody cover variables. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2008.

Thesis (MSc)--Stellenbosch University, 2015. / ENGLISH ABSTRACT: Understanding the effects of climate change is one of the cardinal issues within the natural resource management circles. Increased droughts are part of these changes. Afrotemperate forests, as well as their drier Afromontane counterparts suffer from periodic and seasonal droughts respectively. To better understand the effect of droughts on these forests, three key species namely Olea capensis (Iron wood), Podocarpus latifolius (Common Yellow wood) and Pterocelastrus tricuspidatus (Candle wood), were analysed using dendroecologic techniques. Two sites in the Southern Cape were selected according to a West-to-East moisture gradient, with the drier site being close to George and the medium moist site at the Diepwalle estate in the vicinity of Knysna. Growth ring measurements from each of the species were used to calculate basal area and basal area increment during the lifetime of the trees. Drought years for the sites were then selected based on the Standardised Precipitation and Evapotranspiration Index (SPEI), also indicated by the growth during the drought periods. Differences in growth patterns for all three species were observed. An event analysis was then used to quantify the difference in the resistance (Rt), recovery (Rc), resilience (Rs) and relative resilience (RRs). With values standardised around one (Rt, Rc and Rs) and zero (RRs), it was seen that the Candle wood had the highest (~0.92) resistance and the Yellow wood had the highest (~1.3) recovery after the drought. Iron wood stood apart from the other two species in the sense that it only reacted negatively towards the drought one year after the event in most cases. It was concluded that each of the species were significantly different in their reactions towards drought. This specific difference in drought reaction can give way to the possibility that the species together adapted to relieve the stress of a short drought by splitting the available resources over a longer period. / AFRIKAANSE OPSOMMING: Dit is van uiterse belang vir bestuur doeleindes om die veranderende klimaat oor die wêreld te verstaan, insluitend die droogtes wat daarmee gepaard gaan. Die Afrotemperate woud-tipe, asook sy droeër teenstaander, die Afromontane, lei gereeld aan sporadiese en seisonale droogtes. Om hierdie woud-tipe se reaksie tot droogtes beter te verstaan, was drie boom spesies naamlik Ysterhout (Olea capensis), Kershout (Pterocelastrus tricuspidatus) en gewone Geelhout (Podocarpus latifolius), gekies vir die gebruik in ‘n dendro-ekologiese studie. Twee areas was gekies van ‘n wes-tot-oos droogte gradient, met die droeër blok in die George omgewing en die meer vogtige een naby aan Knysna. Die jaarring metings van elke boom was gebruik om beide die basale oppervlakte en die basale oppervlak groei per jaar aan te teken. ‘n Gestandardiseerde reenval en evapotranspirasie indeks (SPEI) was gebruik om vas te stel jare waarin matige tot sterk droogtes gebeur het. Hierdie gekose jare het aanduiding gegee dat daar wel ‘n verskil waargeneem was in die groei patrone van elke spesie gedurende die droogtes. ‘n Gebeurtenis analise is gebruik om ‘n kwantitatiewe verskil te kon sien in die weerstand (Rt), herstel (Rc), weerstandbiedendheid (Rs) en relatiewe weerstandbiedendheid (RRs). Die was waargeneem dat Kerhout die hoogste weerstand (0.92) toon, terwyl die Geelhout ‘n hoër herstel waarde (1.3) gehad het. Ysterhout het apart van die ander twee spesies gestaan in dìe dat dit eers een jaar na die droogte ‘n reaksie getoon het teenoor die droogte. Dit was dus gevind dat daar spesifieke verskil is tussen al drie van die spesies teen opsigte van stres reaksies was. Hierdie verskil kan dan wel ook moontlik aandui dat hierdie spesies en woud-tipe op so ‘n anier aangepas is dat dit die stress gedurende ‘n kort droogte versprei oor ‘n langer tydperk.

Thesis (MSc (Geography and Environmental Studies))--University of Stellenbosch, 2007. / The Agulhas Plain, an area rich in fynbos, was monitored within six months after the February 2006 fire. The potential of using medium resolution imagery, specifically from the Moderate Resolution Imaging Spectroradiometer (MODIS) in determining the re-growth rates of indigenous and alien vegetation types after fire was explored. Pixels representing dense areas of each vegetation type were selected. There was a significant difference in the pixels selected for each vegetation type. A time series of Normalized Difference Vegetation Index (NDVI) data was derived and fitted to functions, such as Double Logistics and Asymmetric Gaussian as implemented in the TIMESAT software. The results show that alien vegetation grows faster after a fire occurrence than in its absence. Within the specified months of monitoring, it was observed that fynbos grew faster than the alien vegetation. Also, the re-growth rates of vegetation on the coastal soils were higher than those of vegetation on the inland soils. The determination of the re-growth rate was necessary to assist resource managers determine the appropriate time for follow-up of clearing invaded sites after fire.

The aim of this thesis is to examine the role of anthropogenic disturbance in the creation of a socio-ecological landscape. Three key questions were answered: what impact has past anthropogenic disturbance had on present vegetative characteristics; what value did this disturbed landscape have to local people; how did the local peoples' worldviews and eco-cosmologies influence how they perceived, valued and managed their landscape? Research was based in a rural, predominantly amaXhosa village in the Kat River valley, Eastern Cape, South Africa. Four major landscape components characterised the environment, namely dense forests, former grazing lands, abandoned fields and old settlements. A combination of qualitative and quantitative methods were adopted, consisting of a vegetation and soil survey and social science methods including semistructured interviews, focus groups, transect walks and participant observation. The key finding was that anthropogenic disturbance is necessary to enhance the potential of the area to support human habitation. However, anthropogenic disturbance can have positive or negative effects for both local people and the environment. An intermediate level of disturbance is a key factor leading to a resilient socio-ecological system. Various anthropogenic disturbances have had significant affects on vegetation characteristics in terms of species richness, and a change in vegetation composition and species heights. Of the three anthropogenic landscapes examined, former grazing lands were the least intensively disturbed. They were also more species rich and structurally diverse than areas that were under agricultural production or used as a settlement. Dense forests, although the least desirable landscape to local people were, together with sacred pools, sources of ecosystem renewal and played a critical role during times of disturbance. Anthropogenic disturbance has added value to local people's livelihoods; this disturbance has resulted in a patchy landscape that supplies a wide variety of resources to local people. Local people are dependent on their environment for their basic needs; this has encouraged sustainable management practices. The local AmaXhosa still retain many elements of their traditional worldview, which has many characteristics of Animism. The environment is an integral component of their belief system and certain sites and species have significant cultural value and are protected.

Savannahs regulate an agro-ecosystem crucial for the production of domestic livestock, one of the main sources of income worldwide as well as in South African rural communities. Nevertheless, globally these ecosystem functions are threatened by intense human exploitation, inappropriate land use and environmental changes. Leaf area index (LAI) defined as one half the total green leaf area per unit ground surface area, is an inventory of the plant green leaves that defines the actual size of the interface between the vegetation and the atmosphere. Thus, LAI spatial data could serve as an indicator of rangeland productivity. Consequently, the accurate and rapid estimation of LAI is a key requirement for farmers and policy makers to devise sustainable management strategies for rangeland resources. In this study, the main focus was to assess the utility and the accuracy of the PROSAILH radiative transfer model (RTM) to estimate LAI in the South African rangeland on the recently launched Landsat 8 sensor data. The Landsat 8 sensor has been a promising sensor for estimating grassland LAI as compared to its predecessors Landsat 5 to 7 sensors because of its increased radiometric resolution. For this purpose, two PROSAIL inversion methods and semi- empirical methods such as Normalized difference vegetation index (NDVI) were utilized to estimate LAI. The results showed that physically based approaches surpassed empirical approach with highest accuracy yielded by artificial neural network (ANN) inversion approach (RMSE=0.138), in contrast to the Look-Up Table (LUT) approach (RMSE=0.265). In conclusion, the results of this study proved that PROSAIL RTM approach on Landsat 8 data could be utilized to accurately estimate LAI at regional scale which could aid in rapid assessment and monitoring of the rangeland resources. / Environmental Sciences / M. Sc. (Environmental Science)

Organisms inhabit environments that have many dimensions, each of which can vary temporally and spatially. The spatial-temporal variations of environmental stressors and disturbances may have major but different effects on indigenous and invasive species, favouring either of them at different times and places. The invasive mussel Mytilus galloprovincialis invaded the South African coast 30 years ago and, on the south coast of South Africa, it now competes and co-exists with the indigenous Perna perna in the lower eulittoral zone (referred to here as the mussel zone) The invasive and indigenous species dominate the upper and the lower mussel zones respectively, while the two co-exist in the mid-zone. My results show that intertidal mussels experience, and respond to, spatial and temporal fluctuations of several biotic and abiotic stressors. The invasive and the indigenous species adopt different strategies when reacting to environmental factors and their physiological and behavioural responses vary in time and in different habitats as different pressures become of overriding importance. Attachment strength of both species decreased in summer and increased in winter, and was higher on the open coast than in bays for both species, showing a strong positive correlation with wave force in time and space. P. perna had significantly higher attachment strength than M. galloprovincialis but, contrary to previous studies, the difference in gonad index between the two species varied according to the habitat. In bay habitats, M. galloprovincialis had a higher maximum reproductive effort than P. perna, however, on the open coast, there was no significant difference between the two species, suggesting that for the invasive species wave action is a limiting factor not only in terms of the attachment strength but also of energy availability for reproductive tissue development. Major spawning events occurred during periods of low wave action while minor spawning coincided with periods of intense hydrodynamic stress. On the open coast, gonad index was negatively correlated with attachment strength for both species while, in bays, there was no correlation between these two factors for either. The two species also showed different behaviour. In the field, M. galloprovincialis moved significantly more than P. perna over a period of six months. The higher mobility of the invasive species was also confirmed in the laboratory where, in general, M. galloprovincialis formed clumps more readily than P. perna. Taken collectively, these results suggest that channelling more energy into attachment strength limits reproductive tissue development and that, while the indigenous species invests more in byssal production, the invasive species adopts a more dynamic strategy looking for aggregation or a safer arrangement. Higher endolithic infestation and a greater expression of heat shock proteins (Hsps) in mussel populations on the open coast than in bays indicate that this habitat is a more stressful environment not only in terms of wave action. Endolith damaged mussels had significantly lower attachment strengths and condition indices than clean mussels, probably due to the need to channel energy into shell repair. The constant shell repair and expression of Hsps typical of open coast populations are energetically demanding processes. These observations suggest that on the open coast, mussels are subjected to more severe energetic constraints than in bay habitats. Wave and sand stress fluctuated seasonally with the former having a greater effect on mussel mortality on the open coast and the latter a higher impact on bay populations. Overall, mussel mortality rates were higher on the open coast than in bays. My results show that populations on the open coast had fewer private haplotypes and less genetic endemism than those inside bays. Gene flow analysis showed the relatively stable bay habitats act as source populations with greater genetic migration rates out of bays than into them. These differences in genetic structure on scales of las of kilometers show that coastal configuration strongly affects selection, larval dispersal and haplotype diversity. Environmental gradients that are key factors in species distribution over large geographical scales can also be responsible for micro-scale distributions. My results show that M. galloprovincialis colonizes the upper mussel zone where temperature is high, but is less tolerant to this stressor and has to maintain a high expression of Hsps. This suggests that temperature is probably a limiting factor in its invasion towards the sub-tropical east coast. There are inter- and intra-specific differences in responses to the environment which highlight the efforts of M. galloprovincialis and P. perna to optimize resource utilization for survival and reproduction. Determining these differences is crucial to understanding patterns of co-existence between competing indigenous and invasive species.

This thesis presents a history of the landscape of Nqabara, an administrative area in a rural and coastal area of the Eastern Cape Province, South Africa. In the process of investigating landscape history, the inquiry engages with a range of data sources from diverging discursive contexts, including data from ethnographic fieldwork, from the consultation of archival documents and historical reports as well as from comparative historic and ethnographic research, necessitating a critical consideration of the epistemological contexts of data production and the dialogue between researcher and data. Furthermore, in its aim to move beyond historical description towards explanation, the study interrogates the dualist ontological conceptualisations of nature and culture, society and ecology, object and meaning upon which are built three dominant conceptual frameworks concerned with human-environment relationships: social-ecological systems theory, transdisciplinary landscape studies and political ecology. Drawing primarily upon the works of James Gibson, Anthony Giddens and Tim Ingold, an ontological foundation is developed to guide the enquiry and move towards an alternative understanding of the relationship of people’s livelihoods with respect to the landscape in which it is lived, which I call here the praxisembodiment perspective. This ontology takes the situated patterns of action of a situated agent-in-its-environment as its point of departure and proceeds to develop a framework explaining how relations among the patterns of action of different agents-in-their-environment, emerge in structures that simultaneously enable and constrain future action. The foundation is thereby provided for a monist understanding of how landscape and social structure emerge simultaneously from the complex intersection of patterns of actions and interactions of agents in their environment. This framework calls for an understanding of time, space and scale, not as independent variables influencing process and action, but as emergent properties arising from the patterns of actions of situated agents. Finally the alternative ontology is applied to the history of landscape and livelihoods of people of Nqabara. It is concluded that an appropriate understanding and explanation of the critical transformations in the landscape as well as in social institutions, should be sought through analysis of the complex ways in which patterns of action of multiple spatial and temporal rhythms and between multiple agents in an environment, intersect and resonate.

The conservation of leopards in the Eastern Cape Province requires a holistic approach that considers both predator-human interactions as well as the biology of the carnivore. Numerous studies have been conducted on leopards within protected areas; however more information regarding the species is needed outside these areas to facilitate effective management of predators. The spatial ecology of the leopard (Panthera pardus) were studied in the Baviaanskloof and GAENP in the Eastern Cape. The Baviaanskloof is an extensive area of mountainous terrain (approximately 2665km²) which has a mosaic of land uses, and leopards move from conservation areas to farmland where they come into contact and conflict with farmers. This study examined the spatial ecology of leopards living on farmlands adjacent to protected land. The space utilization and activity patterns of six leopards were analysed. These animals were caught and released on farmlands in the Baviaanskloof (n=4) or translocated (n=2) when not possible to release on site. The animals were caught by means of fall-door, walk-in traps and fitted with Vectronic GPS collars that facilitated the collection of high quality GPS fixes from each animal. Data was collected using VHF and UHF telemetry to download data. An understanding of spatial requirements in areas with different land use, and the extent of overlap of space use with other leopards allow, for the first time in the region, the calculation of possible maximum population size. Analysis of range size was carried out using two methods: minimum convex polygon, and Kernel Utilization Distribution. Finally, a key predictor of space use is prey availability. I assessed the prey base using a grid of camera traps. The studied leopards revealed large range utilization with minimal overlap. The activity patterns suggest there is no preference between diurnal and nocturnal activity patterns and the population density was estimated between 0.3 - 1.3 leopards per 100km². Large home range sizes and low population densities suggest that leopards require large areas of suitable habitat, and that conservation efforts need to be extended beyond protected areas to ensure the long-term viability of leopard populations in such areas.

The principal objective of this thesis was to gain an understanding of the movement patterns of spotted grunter Pomadasys commersonnii, an estuarine-dependent fishery species, in the turbid, freshwater dominated Great Fish Estuary. Both manual and automated telemetry methods were used to monitor the movements of spotted grunter during two separate studies conducted in summer and spring 2003 and 2004. Acoustic transmitters were surgically implanted into twenty spotted grunter with lengths between 263 and 387 mm TL in the first study and twenty spotted grunter ranging between 362 and 698 mm TL in the second study. The specific objectives were to gain an understanding of (i) the time spent in the estuarine environment (ii) the space use and home range size, and (iii) the abiotic factors governing the movement patterns of spotted grunter in the estuary. The nursery function of estuarine environments was highlighted in this study as adolescent spotted grunter spent a significantly larger proportion of their time in the estuary than adult fish (p < 0.0001; R² = 0.62). The increased frequency of sea trips, with the onset of sexual maturity, provided testimony of the end of the estuarinedependent phase of their life-cycle. Although considered to be predominantly marine, the adult spotted grunter in the Great Fish Estuary utilised the estuary for considerable periods. Adults are thought to frequent estuaries to forage, seek shelter and to possibly rid themselves of parasites. During this study, the number of sea trips made by tagged fish ranged from 0 to 53, and the duration ranged from 6 hours to 28 days. The tidal phase and time of day had a significant effect (p < 0.05) on the sea trips undertaken by fish. Most tagged spotted grunter left the estuary during the night (84%) on the outgoing tide, and most returned in the evening (77%) during the incoming tide. Sea temperature (p < 0.0001; R² = 0.34), barometric pressure (p = 0.004; R² = 0.19) and wind (p = 0.01) had a significant effect on the number of spotted grunter recorded in the estuary. Spotted grunter were more prone to return to the estuary after high barometric pressure, when low sea temperatures (upwelling events) prevailed. There was a significant positive relationship between home range size and fish length (p = 0.004; R² = 0.20). Small spotted grunter (< 450 mm TL) appeared to be highly resident, with a small home range (mean size = 129 167 m²), that was generally confined to a single core area. Larger individuals (> 450 mm TL) occupied larger home ranges (mean size = 218 435 m²) with numerous core areas. The home ranges of small and large spotted grunter overlapped considerably yielding evidence of two high use areas, situated 1.2 km and 7 km from the mouth of the Great Fish Estuary. Tagged spotted grunter were located in a wide range of salinity, turbidity and temperature, but were found to avoid temperatures below 16 ºC. The daily change in environmental variables (salinity, temperature and turbidity) had a significant effect on the change in fish position in the estuary (p < 0.0001; R² = 0.38). The distribution of tagged spotted grunter, particularly the larger individuals, in the Great Fish Estuary was influenced by the tidal phase (p < 0.05); they moved upriver on the incoming tide and downriver on the outgoing tide. This study provides an understanding of the movement patterns of spotted grunter in the estuary and between the estuarine and marine environments. Consequently, it provides information that will assist in the design of a management plan to promote sustainability of this important fishery species. The techniques used and developed in this study also have direct application for further studies on other important estuarinedependent fishery species.

Studies of the dunefield systems crossing the Cape St. Francis headland in the Eastern Cape have focused on the role that wind plays in sediment transfer in coastal dunefield systems, with limited consideration of the role of water. The aim of this study was to improve understanding of the morphology, processes and patterns within the Oyster Bay HBD system, focussing particularly on surface water and groundwater interactions and the role of surface water in sediment transfer across the dunefield system. An extensive field survey was conducted, to collect related data, complimented by spatial and temporal analysis of the study area using GIS. The key findings from this research were the apparent differences between the western and eastern regions of the dunefield with regard to specific drivers and the respective processes and responses. Wind is the major driver of change up to and across the crest of the dunefield. In the eastern region water (ground water, surface water and the Sand River System) is the primary agent of sediment flux through processes of aggregation and slumping as well as episodic events including debris flows. This study has highlighted a need for further quantitative studies that investigate the movement of sediment through dunefield systems such as this (where water is at or near the land surface). The paradigm that sediment flux is entirely due to wind is almost certainly simplistic, and deeper understanding of these systems is needed / Maiden name: Elkington, Lauren