<p>Continuous measurements of carbon fluxes and meteorological variables were made at a newly initiated flux tower site at an oak-dominant temperate deciduous forest in Southern Ontario, Canada from January to December 2012. Results indicate this forest was a moderate carbon sink in 2012. Annual values of net ecosystem productivity (NEP), gross ecosystem productivity (GEP) and ecosystem respiration (R) were 263 ± 30, 1192 and 922 g C m^-2, respectively. An unusual warm period in March caused a strong increase in R. Erratic peaks of daily air temperature in April also increased R. A drought in July and early August reduced NEP rates when soil moisture values reached the lowest point of the year in late July and early August (minimum 0.023 m³ m^-3). This decrease in NEP was mostly caused by a decrease in GEP, rather than increased R. Water use efficiency at this deciduous forest was 2.86 g C kg^-1 H₂O, indicating conservative water use by the forest. Downwelling photosynthetic active radiation (PAR) was a dominant environmental control on photosynthesis, followed by air temperature and vapour pressure deficit, except in extreme dry periods when soil water stress affected carbon uptake. Extremely cloudy days in the growing season resulted in net carbon release due to low photosynthetic uptake values. Results indicate that large climatic fluctuations in this region may cause high instability in photosynthetic carbon uptake and release from soil carbon pools. This study helps to evaluate and quantify the responses of deciduous forests in the Great Lakes region to future climate change and extreme weather events.</p> / Master of Science (MSc)

Este trabajo está enfocado en la relación entre la productividad y la riqueza de especies de helechos terrestres, y en la disponibilidad de nutrientes a través de un gradiente altitudinal en Ecuador. El trabajo de campo fue realizado durante tres años entre 2009 y 2011, el cual fue llevado a cabo en 8 niveles altitudinales entre 500m y 4000m en la Cordillera Oriental de los Andes ecuatorianos. Diversidad, biomasa, productividad y caracteres funcionales de las hojas de los helechos terrestres fueron estudiados en tres cuadrantes permanentes de 400m2 por cada elevación. Se reportaron un total de 91 especies de helechos terrestres en 32 géneros y 18 familias. A través del gradiente altitudinal, el pico más alto de riqueza de especies se registró en elevaciones medias y la riqueza de especies estuvo débilmente relacionada a medidas de productividad del ecosistema y más fuerte relacionado a la productividad de los hehechos terrestres, lo cual podría estar determinado por un incremento en el número de individuos de helechos y por la disponibilidad de nichos. Sin embargo, en cada nivel altitudinal la relación entre la riqueza de especies de helechos y la productividad de helechos fue negativa, lo cual principalmente se debe a la competencia interespecífica; es decir, en ambas escalas espaciales de este estudio, la diversidad de helechos parece estar limitada por el número de nichos disponibles y la competencia para ocupar estos nichos. Al comparar los caracteres funcionales de las hojas de helechos y árboles a través del gradiente altitudinal se observó similaridades entre ellos. Sin embargo, se encontró una diferencia en el área específica de las hojas entre helechos y árboles, lo cual refleja los diferentes niveles de luz de ambas formas de vida. Respecto los cambios altitudinales en biomasa y productividad de helechos y árboles se encontró una notable diferencia entre estas dos formas de vida, lo cual apoya la hipótesis de que ambos grupos de plantas están creciendo limitados por diferentes factores.

The affect on leaf area index (LAI) and net ecosystem production (NEP) of the mountain pine beetle (Dendroctonus ponderosae) (MPB) outbreak in British Columbia affecting lodgepole pine (Pinus contorta var. latifolia) forests was examined from 1999 to 2008. The process-based carbon (C) cycle model – Boreal Ecosystem Productivity Simulator (BEPS) with remotely sensed LAI inputs was used to produce annual NEP maps, which were validated using field measurements. The annual NEP ranged from 2.43 to -8.03 MtC between 1999 and 2008, with sink to source conversion in 2000. The inter-annual variability for both LAI and NEP displayed initial decreases followed by a steadily increasing trend from 2006 to 2008 with NEP returning to near C neutrality in 2008 (-1.84 MtC). The resistance of LAI and NEP to MPB attack was attributed to ecosystem resilience in the form of secondary overstory growth and increased production of non-attacked host trees.

The affect on leaf area index (LAI) and net ecosystem production (NEP) of the mountain pine beetle (Dendroctonus ponderosae) (MPB) outbreak in British Columbia affecting lodgepole pine (Pinus contorta var. latifolia) forests was examined from 1999 to 2008. The process-based carbon (C) cycle model – Boreal Ecosystem Productivity Simulator (BEPS) with remotely sensed LAI inputs was used to produce annual NEP maps, which were validated using field measurements. The annual NEP ranged from 2.43 to -8.03 MtC between 1999 and 2008, with sink to source conversion in 2000. The inter-annual variability for both LAI and NEP displayed initial decreases followed by a steadily increasing trend from 2006 to 2008 with NEP returning to near C neutrality in 2008 (-1.84 MtC). The resistance of LAI and NEP to MPB attack was attributed to ecosystem resilience in the form of secondary overstory growth and increased production of non-attacked host trees.

In response to global climate change, many important earth-systems-oriented science programs have been established in the past. One such program, the Fluxnet program, studies the response of world forests and other natural ecosystems by measuring biospheric fluxes of carbon dioxide (CO2), water vapour, and energy with eddy-covariance (EC) techniques to assess the role of world ecosystems in offsetting increases in CO2 emissions and related impacts on global climate. The EC methodology has its limitations particularly when weather is inclement and during system stoppages. These limitations create non-trivial problems by creating data gaps in the monitored data stream, diminishing the integrity of the dataset and increasing uncertainty with data interpretation. This Thesis deals with the development of a parsimonious, semi-empirical approach for gap filling of net ecosystem productivity (NEP) data. The approach integrates the effects of environmental controls on diurnal NEP. The approach, because of its limited number of parameters, can be rapidly optimized when appropriate meteorological, site, and NEP target values are provided. The procedure is verified by applying it to several gap-filling case studies, including timeseries collected over balsam fir (Abies Balsamea (L.) Mill.) forests in New Brunswick (NB), Canada and several other forests along a north-south temperaturemoisture gradient from northern Europe to the Middle East. The evaluation showed that the model performed relatively well for most sites; i.e., r2 ranged from 0.68-0.83 and modelling efficiencies, from 0.89-0.97, demonstrating the possibility of applying the model to forests outside NB. Inferior model performance was associated with sites with less than complete input datasets.

This thesis investigates the abiotic control of carbon (C) and water vapour fluxes (FCO₂ and E, respectively) in a New Zealand Pinus radiata D. Don plantation and a nearby clearcut. It concentrates on the limitation of these fluxes imposed by growing season soil water deficit. This results from low precipitation (658 mm a⁻¹) in combination with a limited root zone water storage capacity of the very stony soil (> 30% by volume). The thesis analyses results from seven eddy covariance flux measurement campaigns between November 1994 and March 1996. The study site was located in Balmoral Forest, 100 km north-west of Christchurch (42° 52' S, 172° 45' E), in a (in November 1994) 8-year-old stand. One set of measurements was conducted in an adjacent clearcut. Ecosystem flux measurements were accompanied by separate measurements of ground fluxes and of the associated environmental variables. Flux analysis focussed on the underlying processes of assimilation (Ac), canopy stomatal conductance (Gc) and respiration (Reco), using biophysical models coupled to soil water balance and temperature subroutines. Aiming to link time inegrated net ecosystem C (NEP) to tree growth, sequestration in tree biomass (NPP) was quantified by regular measurements of stem diameter using allometric relationships. Average rates of FCO₂ and E were highest in spring (324 mmol m⁻² d⁻¹ and 207 mol m⁻² d⁻¹, respectively) when the abiotic environment was most favourable for Gc and Ac. During summer, fluxes were impeded by the depletion of available soil water (θ) and the co-occurrence of high air saturation deficit (D) and temperature (T) and were equal or smaller than during winter (FCO₂ = 46 mmol m⁻² d⁻¹ in summer and 115 mmol m⁻² d⁻¹ in winter; E = 57 and 47 mol m⁻² d⁻¹, respectively). With increasingly dry soil, fluxes and their associated ratios became predominantly regulated by D rather than quantum irradiance, and on particularly hot days the ecosystem was a net C source. Interannually, forest C and water fluxes increased strongly with rainfall, and the simultaneously reduced D and T. For two succeeding years, the second having 3 % more rain, modelled NEP was 515 and 716 g C m⁻² a⁻¹, Ac 1690 and 1841 g C m⁻² a⁻¹ and Reco 1175 and 1125 g C m⁻² a⁻¹. NEP / E increased in wetter (and cooler) years (1.3 and 1.5 g kg⁻¹), reflecting a relatively larger gain in NEP. Responding mainly to increased rainfall during commonly dry parts of the year (ie summer), and reflecting the otherwise benign maritime climate of New Zealand, NEP during the winter months could exceed NEP during the middle of the notional tree growing season. Annual Ac, NEP, and NPP were strongly linearly related. This relation did not hold during bi-weekly periods when the processes of intermediate C storage were influential. Separate knowledge of tree growth and C fluxes allowed quantification of autotrophic, and heterotrophic respiration (Rhet≈ 0.4 NEP), as well as fine-root turnover (≈0.2 NEP). The ratio of NEP and stem volume growth was conservative (0.24 t C m⁻³) and allows a direct connection to be made between ecosystem carbon fluxes and forest yield tables. In the absence of living roots, the clearcut flux measurements demonstrated the expected limitation of Rhet by soil temperature (Ts) and θ. However, an additional 'pumping effect' was discovered at the open site whereby turbulence increased CO₂ efflux considerably when the soil surface was wet. Accounting for the combined effects of Ts, θ and turbulence, annual Rhet at the clear-cut site (loss to the atmosphere) was »50 % of NEP (C sequestered from the atmosphere) in the nearby forest. Clearly, there is an important contribution of C fluxes during early stages of ecosystem development to the total C sequestered over the lifetime of a plantation.

Technical complexities and the high cost of satellite images have hindered the adoption of remote sensing technology and tools for nature conservation works in Ethiopia as in many developing countries. The terrestrial and aquatic ecosystems in Abijjata Shala Lakes National Park (ASLNP) and the Important Bird Areas (IBAs) around the park are considered to be one of the most important home ranges for birds. However, little is known about the effect of land use/land cover (LULC) dynamics, due to lack of technical know how and logistical problems. However, it has been shown in this study that sophisticated image management works are not always relevant. Instead a simple method of utilizing the thermal band has been demonstrated. A new approach of long-term dynamics analysis method has also been suggested. A successful classification of images was achieved after such simple enhancement tests. It has been discovered that, there were more active LULC change processes in the area in the first study period (1973 to 1986) than during the second study period (1986-2000). In the first period nearly half of the landscape underwent land cover change processes with more than 26% of the entire landscape experiencing forest or land degradation. In the second period the extent of the change process was limited to only 1/3 of the total area with a smaller amount of degradation processes than before. During the entire study period, agriculture was responsible for the loss of more than 4/5 of the total terrestrial productive ecosystem. More than 37.6% of the total park area has been experiencing this loss for the past 3 decades. Only 1/5 of this area has a chance to revive, the remaining has undergone a permanent degradation. Lake Abijjata lost half of its size during the past 30 years. In the Zeway-Awassa basin 750 km², 2428km² and 3575km² of terrestrial lands and water bodies are within a distance of 10km, 20km and 30km from IBAs respectively. There are ecologically important areas where two or more IBAs overlap. In areas where more than two to five IBAs overlap, up to 85km² of areas have been recently degraded. High livestock density is one of the reasons for degradation. Using a monthly MODIS data from 2000-2005 and a series of interpolation techniques, the productivity of the area as well as the standing biomass were estimated. Moreover, a new method of spatially accurate livestock density assessment was developed in this study. Only 0.3% of the park area is found to be suitable for productive livestock development but nearly all inhabitants think the area is suitable. Feed availability in ASLNP is scarce even during rainy seasons. Especially the open woodlands are subject to overgrazing. Such shortage forces the inhabitants to cut trees for charcoal making to buy animal feed and non-food consumables. While more than 95% of the inhabitants in the park expanded their agriculture lands, only 13.3% of the farmers managed to produce cereals for market. The application of low cost remote sensing and GIS methods provided ample information that enables to conclude that low productivity and household food insecurity are the main driving forces behind land cover changes that are negatively affecting the natural and semi-natural ecosystems in the central and southern Rift Valley of Ethiopia. The restoration of natural ecosystems or conservation of biodiversity can be achieved only if those driving forces are tackled sustainably.

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